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<document id="4F020C4D3EE6F0B70E23E19626946C5B" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA0312FF91FF2FFAACFD491374" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus rolwalingensis Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="62" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="61" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="DFAA9172DD649DD9BC6A90AAF58EC106" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2517AE0F79EE1D23EB2CDD8F38C979F5">
<mods:title id="6BFE6B6AABC76A3841762693D7B9F09C">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
</mods:titleInfo>
<mods:name id="C4CEAF65FE2F6CFD77BAF6D58EA247DB" type="personal">
<mods:role id="BF1488D9DE0E324242368CCB101504FE">
<mods:roleTerm id="A39F6EAFA4CDD2F03CB0FE57F7F6E438">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F5FFBF0EA5052C2202FA6CF01B270D3">Schmidt, Joachim</mods:namePart>
</mods:name>
<mods:typeOfResource id="C852F946B18D0FE8369CF599B0B4FD89">text</mods:typeOfResource>
<mods:relatedItem id="050452CC54497523D3762638BF42F5B3" type="host">
<mods:titleInfo id="A66FEF074BD680239401E4F535412445">
<mods:title id="F3453A7115ED56484C976582FDA479BE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FAE8A1017E089209D411FB352E6C0E90">
<mods:date id="C5A41D6F84C22BA05163D4C5FC869BA5">2009</mods:date>
<mods:detail id="5EEF7C4EF829DFC44ADF17A081F766ED" type="pubDate">
<mods:number id="45B5EF229116BF8679C2394A1212AE44">2009-08-06</mods:number>
</mods:detail>
<mods:detail id="8607120812E024A24FFE1F5BD4B2EEF5" type="volume">
<mods:number id="1EA794A47C0C9DDD31664105C6B404BC">2178</mods:number>
</mods:detail>
<mods:detail id="5C484D65CFCD87359C9CBF3E0B29FF12" type="issue">
<mods:number id="83829C546175E9FDE3852AF8ADF7B80F">1</mods:number>
</mods:detail>
<mods:extent id="79D0A8F9D42BF13A9E0C7B0095EDAC42" unit="page">
<mods:start id="1D14350E3282B6D0B57B857069724F41">1</mods:start>
<mods:end id="A07474169ACC6E46C1C32ACC96661CD6">72</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="949995FBFE46FA497562433182C7F300">
<mods:url id="389779595A6BCA2FFFD0483C2942D53D">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
</mods:location>
<mods:classification id="500F4811A516750FA1A1A88009BC15CB">journal article</mods:classification>
<mods:identifier id="024497535D49E84628C0ABB7637505CF" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
<mods:identifier id="3547022E53AD2773AE6FD1102362B9A0" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="6F6D1C5AC72A8C47BBFA9B2FCECB229A" type="Zenodo-Dep">5312276</mods:identifier>
</mods:mods>
<treatment id="733A87FA0312FF91FF2FFAACFD491374" ID-DOI="http://doi.org/10.5281/zenodo.5320130" ID-Zenodo-Dep="5320130" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374" httpUri="http://treatment.plazi.org/id/733A87FA0312FF91FF2FFAACFD491374" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<subSubSection id="B38965670312FF90FF2FFAACFE351009" box="[151,505,1307,1334]" pageId="60" pageNumber="61" type="nomenclature">
<paragraph id="FB2C36EC0312FF90FF2FFAACFE351009" blockId="60.[151,505,1307,1368]" box="[151,505,1307,1334]" pageId="60" pageNumber="61">
<heading id="A06481800312FF90FF2FFAACFE351009" bold="true" box="[151,505,1307,1334]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
<emphasis id="C9E7EAFE0312FF90FF2FFAACFE351009" bold="true" box="[151,505,1307,1334]" pageId="60" pageNumber="61">
<taxonomicName id="3C934D6F0312FF90FF2FFAACFE7C100A" ID-CoL="57ZZ5" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,432,1308,1334]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="rolwalingensis" status="sp. nov.">
<emphasis id="C9E7EAFE0312FF90FF2FFAACFE7C100A" bold="true" box="[151,432,1308,1334]" italics="true" pageId="60" pageNumber="61">Trechus rolwalingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D457850312FF90FE0FFAABFE351009" box="[439,505,1307,1333]" pageId="60" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38965670312FF90FF2FFA8EFE411064" box="[151,397,1342,1368]" pageId="60" pageNumber="61" type="description">
<paragraph id="FB2C36EC0312FF90FF2FFA8EFE411064" blockId="60.[151,505,1307,1368]" box="[151,397,1342,1368]" pageId="60" pageNumber="61">
(
<figureCitation id="63A82A690312FF90FF27FA8EFECD1064" box="[159,257,1342,1368]" captionStart="FIGURES 2–11" captionStartId="13.[151,269,1818,1842]" captionTargetBox="[262,1324,194,1794]" captionTargetId="figure-115@13.[262,1325,194,1794]" captionTargetPageId="13" captionText="FIGURES 2–11. Pronotum. Fig. 2, Trechus thibetanus Jeannel, 1928, Lectotype. Fig. 3, T. dongulaensis sp. n., paratype, male. Fig. 4, T. namtsoensis sp. n., paratype, male. Fig. 5, T. eutrechoides eutrechoides Deuve, 1992, non-type male specimen from Lamna La. Fig. 6, T. glabratus sp. n., paratype, male. Fig. 7, T. korae sp. n., paratype, male. Fig. 8, T. yak yak ssp. n., paratype, male. Fig. 9, T. yak shogulaensis ssp. n., paratype, male. Fig. 10, T. rolwalingensis rolwalingensis ssp. n., paratype, male. Fig. 11, T. rolwalingensis daldunglana ssp. n., paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5312280" httpUri="https://zenodo.org/record/5312280/files/figure.png" pageId="60" pageNumber="61">Figs. 10</figureCitation>
,
<figureCitation id="63A82A690312FF90FEB6FA8EFE961064" box="[270,346,1342,1368]" captionStart="FIGURES 52–64" captionStartId="30.[151,270,1830,1854]" captionTargetBox="[214,1367,199,1799]" captionTargetId="figure-139@30.[214,1372,194,1799]" captionTargetPageId="30" captionText="FIGURES 52–64. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 52–56, 60–63; Figs. 52, 54, 55, 60, 61, 63 in addition with parameres) and in dorsal view (Figs. 57–59, 64). Fig. 52, Trechus damchungensis Deuve, 1997, holotype. Figs. 53, 57, T. cf. damchungensis Deuve, 1997, paratype from Tangula Pass. Fig. 54, T. mieheorum sp. n., paratype. Figs. 55, 58, T. bastropi sp. n., paratype. Fig. 56, T. hodeberti Deuve, 1997, nontype specimen from Lhachen La (= locus typicus). Figs. 59, 60, T. solhoeyi sp. n., paratype. Fig. 61, T. yeti sp. n., paratype. Fig. 62, T. budhaensis sp. n., paratype. Fig. 63, 64, T. rolwalingensis sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312294" httpUri="https://zenodo.org/record/5312294/files/figure.png" pageId="60" pageNumber="61">63, 64</figureCitation>
,
<figureCitation id="63A82A690312FF90FEDDFA8EFE4F1064" box="[357,387,1342,1368]" captionStart="FIGURES 81–86" captionStartId="32.[151,269,1942,1966]" captionTargetBox="[171,1424,190,1921]" captionTargetId="figure-78@32.[151,1436,146,1930]" captionTargetPageId="32" captionText="FIGURES 81–86. Habitus. Fig. 81, Trechus dongulaensis sp. n., paratype, female. Fig. 82, T. glabratus sp. n., paratype, female. Fig. 83, T. rolwalingensis rolwalingensis ssp. n., holotype. Fig. 84, T. stratiotes stratiotes ssp. n., paratype, male. Fig. 85, T. muguensis sp. n., paratype, female. Fig. 86, T. sculptipennis sp. n., paratype, female." figureDoi="http://doi.org/10.5281/zenodo.5312300" httpUri="https://zenodo.org/record/5312300/files/figure.png" pageId="60" pageNumber="61">83</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B38965670312FF90FF2FFA3BFA5613B2" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="FB2C36EC0312FF90FF2FFA3BFDBE10F0" blockId="60.[151,1436,1419,2026]" pageId="60" pageNumber="61">
<emphasis id="C9E7EAFE0312FF90FF2FFA3BFE9C1099" bold="true" box="[151,336,1419,1445]" pageId="60" pageNumber="61">Type material:</emphasis>
<materialsCitation id="4BFB3CB10312FF90FEE1FA3CFDBE10F0" collectingDate="1999-09-18" collectionCode="SMNS" country="Nepal" elevation="4700" location="Yarlung Ri" pageId="60" pageNumber="61" specimenCount="1" typeStatus="holotype">
<typeStatus id="2428884E0312FF90FEE1FA3CFE0A109A" box="[345,454,1420,1446]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
male, with label data “
<collectingCountry id="8384767C0312FF90FD60FA3CFCF4109A" box="[728,824,1420,1446]" name="Nepal" pageId="60" pageNumber="61">NEPAL</collectingCountry>
Rolwaling Vall.,
<location id="FE4C60370312FF90FBB1FA3CFB47109A" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374:FE4C60370312FF90FBB1FA3CFB47109A" box="[1033,1163,1420,1446]" country="Nepal" name="Yarlung Ri" pageId="60" pageNumber="61">Yarlung Ri</location>
base camp
<quantity id="3C6B9B090312FF90FAAFFA3CFF7C10F0" metricMagnitude="3" metricUnit="m" metricValue="4.7" metricValueMax="4.8" metricValueMin="4.6" pageId="60" pageNumber="61" unit="m" value="4700.0" valueMax="4800.0" valueMin="4600.0">
<elevation id="70BED1DF0312FF90FAAFFA3CFF7C10F0" metricMagnitude="3" metricUnit="m" metricValue="4.7" metricValueMax="4.8" metricValueMin="4.6" pageId="60" pageNumber="61" unit="m" value="4700.0" valueMax="4800.0" valueMin="4600.0">4600–4800 m</elevation>
</quantity>
, 16.–
<date id="8F2D102C0312FF90FF53FA02FEA910F0" box="[235,357,1458,1484]" pageId="60" pageNumber="61" value="1999-09-18">
<collectingDate id="9F69E9C40312FF90FF53FA02FEA910F0" box="[235,357,1458,1484]" pageId="60" pageNumber="61" value="1999-09-18">18.9.1999</collectingDate>
</date>
lg. Schmidt” (
<collectionCode id="9D82AE290312FF90FDA8FA02FDAA10F0" box="[528,614,1458,1484]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="60" pageNumber="61">SMNS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FB2C36EC0312FF90FF7DFA69FA5613B2" blockId="60.[151,1436,1419,2026]" pageId="60" pageNumber="61">
<materialsCitation id="4BFB3CB10312FF90FF7DFA69FEAA1326" collectingDate="1999-09-18" collectionCode="BMNH, CKAB, CSCHM, CWR, MNHN, SMNS" country="Nepal" elevation="4700" location="Yarlung Ri" pageId="60" pageNumber="61" specimenCount="29" specimenCount-female="11" specimenCount-male="18" typeStatus="paratype">
<typeStatus id="2428884E0312FF90FF7DFA69FE8C10CF" box="[197,320,1497,1523]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ED95FD650312FF90FEF5FA69FE7210CF" box="[333,446,1497,1523]" pageId="60" pageNumber="61" type="male">18 males</specimenCount>
,
<specimenCount id="ED95FD650312FF90FE74FA69FD9810CF" box="[460,596,1497,1523]" pageId="60" pageNumber="61" type="female">11 females</specimenCount>
, with same label data as holotype (
<collectionCode id="9D82AE290312FF90FBBDFA69FBA810CF" box="[1029,1124,1497,1523]" pageId="60" pageNumber="61">BMNH</collectionCode>
,
<collectionCode id="9D82AE290312FF90FBCAFA69FB0410CF" box="[1138,1224,1497,1523]" pageId="60" pageNumber="61">CKAB</collectionCode>
,
<collectionCode id="9D82AE290312FF90FB6EFA69FA8E10CF" box="[1238,1346,1497,1523]" pageId="60" pageNumber="61">CSCHM</collectionCode>
,
<collectionCode id="9D82AE290312FF90FAE8FA69FA5B10CF" box="[1360,1431,1497,1523]" pageId="60" pageNumber="61">CWR</collectionCode>
,
<collectionCode id="9D82AE290312FF90FF2FF9B0FF3B1326" box="[151,247,1536,1562]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="60" pageNumber="61">MNHN</collectionCode>
,
<collectionCode id="9D82AE290312FF90FEBDF9B0FE911326" box="[261,349,1536,1562]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="60" pageNumber="61">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10312FF90FECCF9B0FEB3137C" collectingDate="1999-09-16" collectionCode="CSCHM" collectorName="J. Schmidt" country="Nepal" elevation="4050" location="Rolwaling Valley" municipality="Na" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="ED95FD650312FF90FECCF9B0FE071326" box="[372,459,1536,1562]" pageId="60" pageNumber="61" type="male">1 male</specimenCount>
,
<collectingCountry id="8384767C0312FF90FE61F9B0FDEA1326" box="[473,550,1536,1562]" name="Nepal" pageId="60" pageNumber="61">Nepal</collectingCountry>
,
<location id="FE4C60370312FF90FD8CF9B0FCC21326" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374:FE4C60370312FF90FD8CF9B0FCC21326" box="[564,782,1536,1562]" country="Nepal" municipality="Na" name="Rolwaling Valley" pageId="60" pageNumber="61">Rolwaling Valley</location>
,
<collectingMunicipality id="1B48AC960312FF90FCA4F9B0FCF31326" box="[796,831,1536,1562]" pageId="60" pageNumber="61">Na</collectingMunicipality>
vill. [village]
<quantity id="3C6B9B090312FF90FC4FF9B0FB681326" box="[1015,1188,1536,1562]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="60" pageNumber="61" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">
<elevation id="70BED1DF0312FF90FC4FF9B0FB681326" box="[1015,1188,1536,1562]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="60" pageNumber="61" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100 m</elevation>
</quantity>
,
<date id="8F2D102C0312FF90FB0BF9B0FA8E1326" box="[1203,1346,1536,1562]" pageId="60" pageNumber="61" value="1999-09-16">
<collectingDate id="9F69E9C40312FF90FB0BF9B0FA8E1326" box="[1203,1346,1536,1562]" pageId="60" pageNumber="61" value="1999-09-16">16.IX.1999</collectingDate>
</date>
, leg.
<collectorName id="5666533A0312FF90FA30F9B0FF35137C" pageId="60" pageNumber="61">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290312FF90FEB1F996FEBB137C" box="[265,375,1574,1600]" pageId="60" pageNumber="61">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10312FF90FE34F996FE12135B" collectingDate="2000-05-23" collectionCode="CSCHM" collectorName="J. Schmidt" country="Nepal" elevation="4550" location="Na" municipality="Yarlung Ri" pageId="60" pageNumber="61" specimenCount="29" specimenCount-female="16" specimenCount-male="13" typeStatus="paratype">
<specimenCount id="ED95FD650312FF90FE34F996FE35137C" box="[396,505,1574,1600]" pageId="60" pageNumber="61" type="male">13 males</specimenCount>
,
<specimenCount id="ED95FD650312FF90FDBEF996FD45137C" box="[518,649,1574,1600]" pageId="60" pageNumber="61" type="female">16 females</specimenCount>
, ditto, but:
<location id="FE4C60370312FF90FCAAF996FCF8137C" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374:FE4C60370312FF90FCAAF996FCF8137C" box="[786,820,1574,1600]" municipality="Yarlung Ri" name="Na" pageId="60" pageNumber="61">Na</location>
to
<collectingMunicipality id="1B48AC960312FF90FCE3F996FC11137C" box="[859,989,1574,1600]" pageId="60" pageNumber="61">Yarlung Ri</collectingMunicipality>
base camp
<quantity id="3C6B9B090312FF90FBD1F996FAC3137C" box="[1129,1295,1574,1600]" metricMagnitude="3" metricUnit="m" metricValue="4.55" metricValueMax="4.9" metricValueMin="4.2" pageId="60" pageNumber="61" unit="m" value="4550.0" valueMax="4900.0" valueMin="4200.0">
<elevation id="70BED1DF0312FF90FBD1F996FAC3137C" box="[1129,1295,1574,1600]" metricMagnitude="3" metricUnit="m" metricValue="4.55" metricValueMax="4.9" metricValueMin="4.2" pageId="60" pageNumber="61" unit="m" value="4550.0" valueMax="4900.0" valueMin="4200.0">4200–4900 m</elevation>
</quantity>
,
<date id="8F2D102C0312FF90FAA4F996FA5B137C" box="[1308,1431,1574,1600]" pageId="60" pageNumber="61" value="2000-05-23">
<collectingDate id="9F69E9C40312FF90FAA4F996FA5B137C" box="[1308,1431,1574,1600]" pageId="60" pageNumber="61" value="2000-05-23">23.V.2000</collectingDate>
</date>
, leg.
<collectorName id="5666533A0312FF90FF76F9FDFE9D135B" box="[206,337,1613,1639]" pageId="60" pageNumber="61">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290312FF90FEDBF9FDFE19135B" box="[355,469,1613,1639]" pageId="60" pageNumber="61">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10312FF90FE54F9FDFEB113B2" collectingDate="2000-05-21" collectionCode="CSCHM" collectorName="J. Schmidt" country="Nepal" elevation="4300" location="Na" municipality="Tso Rolpa lake" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="ED95FD650312FF90FE54F9FDFD97135B" box="[492,603,1613,1639]" pageId="60" pageNumber="61" type="female">1 female</specimenCount>
, ditto, but:
<location id="FE4C60370312FF90FD54F9FDFCC3135B" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374:FE4C60370312FF90FD54F9FDFCC3135B" box="[748,783,1613,1639]" municipality="Tso Rolpa lake" name="Na" pageId="60" pageNumber="61">Na</location>
to
<collectingMunicipality id="1B48AC960312FF90FC82F9FDFC3B135B" box="[826,1015,1613,1639]" pageId="60" pageNumber="61">Tso Rolpa lake</collectingMunicipality>
,
<quantity id="3C6B9B090312FF90FBBEF9FDFB7F135B" box="[1030,1203,1613,1639]" metricMagnitude="3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="60" pageNumber="61" unit="m" value="4300.0" valueMax="4400.0" valueMin="4200.0">
<elevation id="70BED1DF0312FF90FBBEF9FDFB7F135B" box="[1030,1203,1613,1639]" metricMagnitude="3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="60" pageNumber="61" unit="m" value="4300.0" valueMax="4400.0" valueMin="4200.0">4200–4400 m</elevation>
</quantity>
,
<date id="8F2D102C0312FF90FB7AF9FDFA8F135B" box="[1218,1347,1613,1639]" pageId="60" pageNumber="61" value="2000-05-21">
<collectingDate id="9F69E9C40312FF90FB7AF9FDFA8F135B" box="[1218,1347,1613,1639]" pageId="60" pageNumber="61" value="2000-05-21">21.V.2000</collectingDate>
</date>
, leg.
<collectorName id="5666533A0312FF90FA30F9FDFF3413B2" pageId="60" pageNumber="61">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290312FF90FEB0F9C4FEB913B2" box="[264,373,1652,1678]" pageId="60" pageNumber="61">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10312FF90FE31F9C4FA5913B2" box="[393,1429,1652,1678]" collectingDate="2000-05-21" collectionCode="CSCHM" collectorName="J. Schmidt" country="Nepal" elevation="4400" location="Tsho Rolpa" municipality="Tso Rolpa lake" pageId="60" pageNumber="61" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="ED95FD650312FF90FE31F9C4FE2A13B2" box="[393,486,1652,1678]" pageId="60" pageNumber="61" type="male">2 males</specimenCount>
,
<specimenCount id="ED95FD650312FF90FE4AF9C4FD9713B2" box="[498,603,1652,1678]" pageId="60" pageNumber="61" type="female">1 female</specimenCount>
, ditto, but:
<location id="FE4C60370312FF90FD58F9C4FCA513B2" LSID="urn:lsid:plazi:treatment:733A87FA0312FF91FF2FFAACFD491374:FE4C60370312FF90FD58F9C4FCA513B2" box="[736,873,1652,1678]" name="Tsho Rolpa" pageId="60" pageNumber="61">Tsho Rolpa</location>
,
<quantity id="3C6B9B090312FF90FCCCF9C4FC0313B2" box="[884,975,1652,1678]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="m" value="4400.0">
<elevation id="70BED1DF0312FF90FCCCF9C4FC0313B2" box="[884,975,1652,1678]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="m" value="4400.0">4400 m</elevation>
</quantity>
,
<date id="8F2D102C0312FF90FC64F9C4FB9A13B2" box="[988,1110,1652,1678]" pageId="60" pageNumber="61" value="2000-05-21">
<collectingDate id="9F69E9C40312FF90FC64F9C4FB9A13B2" box="[988,1110,1652,1678]" pageId="60" pageNumber="61" value="2000-05-21">21.V.2000</collectingDate>
</date>
, leg.
<collectorName id="5666533A0312FF90FB2DF9C4FADC13B2" box="[1173,1296,1652,1678]" pageId="60" pageNumber="61">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290312FF90FA98F9C4FA4113B2" box="[1312,1421,1652,1678]" pageId="60" pageNumber="61">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670312FF91FF7DF92AFB59167F" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="description">
<paragraph id="FB2C36EC0312FF90FF7DF92AFD571389" blockId="60.[151,1436,1419,2026]" box="[197,667,1690,1717]" pageId="60" pageNumber="61">
<emphasis id="C9E7EAFE0312FF90FF7DF92AFE931388" bold="true" box="[197,351,1690,1716]" pageId="60" pageNumber="61">Description:</emphasis>
Body length:
<quantity id="3C6B9B090312FF90FDBEF92BFD5B1389" box="[518,663,1691,1717]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="60" pageNumber="61" unit="mm" value="3.45" valueMax="3.7" valueMin="3.2">3.2–3.7 mm</quantity>
.
</paragraph>
<paragraph id="FB2C36EC0312FF90FF7DF971FE42123E" blockId="60.[151,1436,1419,2026]" pageId="60" pageNumber="61">Colour: Dorsal surface dark brown, shiny; palpi, scapus, pedicellus, anterior half of antennal segment III and legs light brown.</paragraph>
<paragraph id="FB2C36EC0312FF90FF7DF8BEFD3212A1" blockId="60.[151,1436,1419,2026]" pageId="60" pageNumber="61">Microsculpture: Head with moderately engraved isodiametric meshes on supraorbital area, neck and in frontal furrows (x40–50). Pronotum with more faintly engraved slightly transverse meshes on disc (x80) but more deeply impressed almost isodiametric meshes in basal depressions. Elytral disc with faintly engraved narrow and more strongly transverse meshes (x100).</paragraph>
<paragraph id="FB2C36EC0312FF91FF7DF819FE9615E4" blockId="60.[151,1436,1419,2026]" lastBlockId="61.[151,1437,152,1608]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">Head: Broad, with eyes small and slightly protruding; temples 2/3–3/4 of length of eyes and strongly wrinkled to the neck. Antennae short, 1.5–2 antennomeres extend beyond the pronotal base. Antennomere III is as long as or slightly longer than antennomere II, antennomere IV is slightly shorter (5/6–6/7) as antennomere III.</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFF55FC3B14FC" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">Pronotum: Large and transverse, subcordate, strongly contracted towards base; proportions: WP/LP = 1.31–1.39, WP/WPB = 1.36–1.42, WP/WH = 1.31–1.38, WE/WP = 1.53–1.59. Surface strongly convex. Sides convexly rounded in anterior 3/4 and curtly concave anterad of hind angles; the latter small but rectangular, seldom slightly obtuse. Marginal gutter moderately narrow, slightly widened anterad of laterobasal depressions. Base rectilinear or weakly convex, slightly curved anteriorly at outer fifth; basal depressions with fine longitudinal wrinkles both sides of pronotal middle.</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFE7DFE8617F3" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">Elytra: Broad oval, broadest little behind mid-length, with proportion WE/LE = 1.36–1.41. Surface strongly convex, not flattened on disc. Shoulders rounded, indistinct. Striae impunctate, first and second striae fully deep impressed, third striae somewhat shallower and reduced at base and extreme apex, fourth stria indistinct and striae V–VII completely reduced. Second and third interval strongly convex. Preapical seta is located in the third interval often close to the second stria and, in most specimens, at the beginning of the posterior elytral seventh, but in some specimens located slightly anteriorly (up to the beginning of the posterior fifth).</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFD6CFB59167F" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">
Male genitalia: Aedeagal median lobe moderately short (LE/LA = 2.87–3.00), with ventral side almost straight in middle, and with terminal lamella slightly bent downwards; the latter shortly bill-like in lateral and in dorsal view. Copulatory piece of internal sac relatively complex as shown in
<figureCitation id="63A82A690313FF91FB82FC99FB5D167F" box="[1082,1169,809,835]" captionStart="FIGURES 52–64" captionStartId="30.[151,270,1830,1854]" captionTargetBox="[214,1367,199,1799]" captionTargetId="figure-139@30.[214,1372,194,1799]" captionTargetPageId="30" captionText="FIGURES 52–64. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 52–56, 60–63; Figs. 52, 54, 55, 60, 61, 63 in addition with parameres) and in dorsal view (Figs. 57–59, 64). Fig. 52, Trechus damchungensis Deuve, 1997, holotype. Figs. 53, 57, T. cf. damchungensis Deuve, 1997, paratype from Tangula Pass. Fig. 54, T. mieheorum sp. n., paratype. Figs. 55, 58, T. bastropi sp. n., paratype. Fig. 56, T. hodeberti Deuve, 1997, nontype specimen from Lhachen La (= locus typicus). Figs. 59, 60, T. solhoeyi sp. n., paratype. Fig. 61, T. yeti sp. n., paratype. Fig. 62, T. budhaensis sp. n., paratype. Fig. 63, 64, T. rolwalingensis sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312294" httpUri="https://zenodo.org/record/5312294/files/figure.png" pageId="61" pageNumber="62">Fig. 63</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670313FF91FF7DFCFFFBD2102F" pageId="61" pageNumber="62" type="etymology">
<paragraph id="FB2C36EC0313FF91FF7DFCFFFA8E1656" blockId="61.[151,1437,152,1608]" box="[197,1346,847,874]" pageId="61" pageNumber="62">
<emphasis id="C9E7EAFE0313FF91FF7DFCFFFE9A1655" bold="true" box="[197,342,847,873]" pageId="61" pageNumber="62">Etymology:</emphasis>
The specific name is derived from the
<typeStatus id="2428884E0313FF91FC98FCE0FC9E1656" box="[800,850,848,874]" pageId="61" pageNumber="62">type</typeStatus>
locality, the Rolwaling Valley (adjective).
</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFCC6FB1D16E2" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">
<emphasis id="C9E7EAFE0313FF91FF7DFCC6FEB416AC" bold="true" box="[197,376,886,912]" pageId="61" pageNumber="62">Identification:</emphasis>
Within the
<taxonomicName id="3C934D6F0313FF91FDBAFCC7FD9216AC" authorityName="Clairville" authorityYear="1806" box="[514,606,887,912]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C9E7EAFE0313FF91FDBAFCC7FD9216AC" box="[514,606,887,912]" italics="true" pageId="61" pageNumber="62">Trechus</emphasis>
</taxonomicName>
fauna of
<collectingRegion id="3957F80E0313FF91FD74FCC6FCC516AC" box="[716,777,886,912]" country="China" name="Xizang" pageId="61" pageNumber="62">Tibet</collectingRegion>
and
<taxonomicName id="3C934D6F0313FF91FCFAFCC6FC7816AC" box="[834,948,886,912]" class="Cestoda" family="Anoplocephalidae" genus="Himalaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="61" pageNumber="72" phylum="Platyhelminthes" rank="genus">Himalaya</taxonomicName>
this new species is easily to recognize by its unique male genitalia characters, especially by the external shape of the aedeagal median lobe, by the strongly sclerotized median lobe ostium, and the extraordinary form of the copulatory piece.
</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFC5AFBD2102F" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">
<emphasis id="C9E7EAFE0313FF91FF7DFC5AFEB41138" bold="true" box="[197,376,1002,1028]" pageId="61" pageNumber="62">Relationships:</emphasis>
Due to the large and more strongly transverse pronotum with deep laterobasal foveae, the broader oval elytra with more strongly reduced lateral striae, and due to the general shape of the copulatory piece with broader basal portion and thorn-like or needle-like distal portion,
<taxonomicName id="3C934D6F0313FF91FBA4FB88FB28116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1052,1252,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="rolwalingensis" status="sp. nov.">
<emphasis id="C9E7EAFE0313FF91FBA4FB88FB28116D" box="[1052,1252,1080,1105]" italics="true" pageId="61" pageNumber="62">T. rolwalingensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0313FF91FB54FB87FAE2116D" bold="true" box="[1260,1326,1079,1105]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="D2D457850313FF91FB54FB87FAE2116D" box="[1260,1326,1079,1105]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
seems to be member of the species diverse Central Himalayan group of
<taxonomicName id="3C934D6F0313FF91FCC8FBEFFB901144" authority="Ueno, 1972" authorityName="Ueno" authorityYear="1972" box="[880,1116,1118,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tosioi">
<emphasis id="C9E7EAFE0313FF91FCC8FBEFFC011144" box="[880,973,1119,1144]" italics="true" pageId="61" pageNumber="62">T. tosioi</emphasis>
Uéno, 1972
</taxonomicName>
(which, after a preliminary study, includes
<taxonomicName id="3C934D6F0313FF91FED9FB35FD7311A3" authority="Morvan, 1972" authorityName="Morvan" authorityYear="1972" box="[353,703,1157,1183]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="breuningi">
<emphasis id="C9E7EAFE0313FF91FED9FB35FE3011A2" box="[353,508,1157,1182]" italics="true" pageId="61" pageNumber="62">T. breuningi</emphasis>
Morvan, 1972
</taxonomicName>
,
<taxonomicName id="3C934D6F0313FF91FD75FB35FBD511A3" authority="Schmidt, 1998" authorityName="Schmidt" authorityYear="1998" box="[717,1049,1157,1183]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="gorkhai">
<emphasis id="C9E7EAFE0313FF91FD75FB35FC9C11A2" box="[717,848,1157,1182]" italics="true" pageId="61" pageNumber="62">T. gorkhai</emphasis>
Schmidt, 1998
</taxonomicName>
,
<taxonomicName id="3C934D6F0313FF91FB91FB35FABA11A3" authority="Schmidt, 1998" authorityName="Schmidt" authorityYear="1998" box="[1065,1398,1157,1183]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="gurungi">
<emphasis id="C9E7EAFE0313FF91FB91FB35FB6211A2" box="[1065,1198,1157,1182]" italics="true" pageId="61" pageNumber="62">T. gurungi</emphasis>
Schmidt, 1998
</taxonomicName>
,
<taxonomicName id="3C934D6F0313FF91FA3DFB35FE1311FA" authority="Schmidt, 1994" authorityName="Schmidt" authorityYear="1994" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="lamjunensis">
<emphasis id="C9E7EAFE0313FF91FA3DFB35FEEA11F9" italics="true" pageId="61" pageNumber="62">T. lamjunensis</emphasis>
<bibRefCitation id="9F024B1D0313FF91FE96FB1CFE1311FA" author="Schmidt, J." box="[302,479,1196,1222]" pageId="61" pageNumber="62" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="3C934D6F0313FF91FE55FB1CFCA511FA" authority="Schmidt, 1998" authorityName="Schmidt" authorityYear="1998" box="[493,873,1196,1222]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="namunlaensis">
<emphasis id="C9E7EAFE0313FF91FE55FB1CFD6311F9" box="[493,687,1196,1221]" italics="true" pageId="61" pageNumber="62">T. namunlaensis</emphasis>
Schmidt, 1998
</taxonomicName>
, and
<taxonomicName id="3C934D6F0313FF91FC12FB1CFB2A11FA" authority="Schmidt, 1998" authorityName="Schmidt" authorityYear="1998" box="[938,1254,1196,1222]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tamangi">
<emphasis id="C9E7EAFE0313FF91FC12FB1CFBE611F9" box="[938,1066,1196,1221]" italics="true" pageId="61" pageNumber="62">T. tamangi</emphasis>
Schmidt, 1998
</taxonomicName>
). However, the male genitalia characters of the new species are so striking that a more detailed character study of the Himalayan
<taxonomicName id="3C934D6F0313FF91FEA7FB49FEB7102E" authorityName="Clairville" authorityYear="1806" box="[287,379,1273,1298]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C9E7EAFE0313FF91FEA7FB49FEB7102E" box="[287,379,1273,1298]" italics="true" pageId="61" pageNumber="62">Trechus</emphasis>
</taxonomicName>
fauna is needed before further conclusions can be drawn.
</paragraph>
</subSubSection>
<subSubSection id="B38965670313FF91FF7DFAAFFD491374" pageId="61" pageNumber="62" type="distribution">
<paragraph id="FB2C36EC0313FF91FF7DFAAFFC9B1092" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">
<emphasis id="C9E7EAFE0313FF91FF7DFAAFFD0D1005" bold="true" box="[197,705,1311,1337]" pageId="61" pageNumber="62">Distribution and geographical variation:</emphasis>
<figureCitation id="63A82A690313FF91FD72FA90FCED1006" box="[714,801,1312,1338]" captionStart="FIGURE 98" captionStartId="65.[151,255,1735,1759]" captionTargetBox="[444,1143,426,1711]" captionTargetId="figure-254@65.[444,1143,426,1711]" captionTargetPageId="65" captionText="FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu &amp; Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n." figureDoi="http://doi.org/10.5281/zenodo.5312314" httpUri="https://zenodo.org/record/5312314/files/figure.png" pageId="61" pageNumber="62">Fig. 98</figureCitation>
. High
<taxonomicName id="3C934D6F0313FF91FCD7FA90FC2D1006" box="[879,993,1312,1338]" class="Cestoda" family="Anoplocephalidae" genus="Himalaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="61" pageNumber="72" phylum="Platyhelminthes" rank="genus">Himalaya</taxonomicName>
of Central
<collectingCountry id="8384767C0313FF91FBDFFA90FB7D1006" box="[1127,1201,1312,1338]" name="Nepal" pageId="61" pageNumber="62">Nepal</collectingCountry>
: The nominotypical form is distributed in the upper Rolwaling Valley between Solu Khumbu Massif in the South and Rolwaling Himal in the North. A form somewhat differing in external morphology was found in the western Rolwaling Valley and will be described as a separate subspecies below.
</paragraph>
<paragraph id="FB2C36EC0313FF91FF7DFA0AFD491374" blockId="61.[151,1437,152,1608]" pageId="61" pageNumber="62">
<emphasis id="C9E7EAFE0313FF91FF7DFA0AFEE310E8" bold="true" box="[197,303,1466,1492]" pageId="61" pageNumber="62">Habitat:</emphasis>
Vertical distribution approximately
<quantity id="3C6B9B090313FF91FD67FA0BFC4A10E9" box="[735,902,1467,1493]" metricMagnitude="3" metricUnit="m" metricValue="4.1" metricValueMax="4.6" metricValueMin="3.6" pageId="61" pageNumber="62" unit="m" value="4100.0" valueMax="4600.0" valueMin="3600.0">3600–4600 m</quantity>
, from the cloud forests of the higher parts of the high montane zone (“Obere Nebelwaldstufe” sensu
<bibRefCitation id="9F024B1D0313FF91FC94FA51FC0F10C7" author="Miehe, G." box="[812,963,1505,1531]" pageId="61" pageNumber="62" pagination="181 - 230" refId="ref49280" refString="Miehe, G. (1991) Der Himalaya, eine multizonale Gebirgsregion. In: Walter, H. &amp; Breckle, S. - W. (Eds.), Okologie der Erde, Band 4. Spezielle Okologie der Gemassigten und Arktischen Zonen ausserhalb Euro-Nordasiens. Gustav Fischer Verlag, Stuttgart, pp. 181 - 230." type="book chapter" year="1991">Miehe, 1991</bibRefCitation>
) up to the meadows of the lower alpine zone. In the cloud forests the specimens were sifted from leaf litter, and in the subalpine and alpine zones they were found under stones on humid slopes.
</paragraph>
</subSubSection>
</treatment>
</document>
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