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<document id="D261D6A773FD9E4E74B7E061AB2B3843" ID-CLB-Dataset="29418" ID-DOI="10.11646/zootaxa.4482.3.2" ID-GBIF-Dataset="fd24ef18-76ac-46f1-aec2-2faf5b0ff325" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440707" ID-ZooBank="3FBE11E2-9F97-4A29-92A9-67780C782E0D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538375301724" checkinUser="plazi" docAuthor="Li, Jheng-Jhang, Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2018" docId="376A1B4DFFB6FF95D1CDFC7C4548F812" docLanguage="en" docName="zootaxa.4482.3.2.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parasesarma gracilipes Li &amp; Rahayu &amp; Ng 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="470" masterDocId="CB536335FFA4FF86D15AFFF74432FFDA" masterDocTitle="Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific" masterLastPageNumber="490" masterPageNumber="451" pageNumber="469" updateTime="1698559507815" updateUser="ExternalLinkService">
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<mods:title id="F10F6F888C1FAB31F438FF31800384EC">Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific</mods:title>
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<mods:namePart id="37E5144A7B65DCD2DD75A4F7DE73AB92">Li, Jheng-Jhang</mods:namePart>
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<mods:namePart id="052FCC9100EC04B6734A68BBAF4099A7">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="5CE96A67F193552C2979DC1246B8AF2F">Ng, Peter K. L.</mods:namePart>
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<mods:title id="BBC56E803362BA157691EEA19659F532">Zootaxa</mods:title>
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<mods:date id="5F97DCF7008293BA1F8BDAF5F3EB7271">2018</mods:date>
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<treatment id="376A1B4DFFB6FF95D1CDFC7C4548F812" ID-DOI="http://doi.org/10.5281/zenodo.5986793" ID-GBIF-Taxon="148421722" ID-Zenodo-Dep="5986793" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812" httpUri="http://treatment.plazi.org/id/376A1B4DFFB6FF95D1CDFC7C4548F812" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<subSubSection id="F7D9F9D0FFB6FF94D1CDFC7C45D3FC1D" pageId="18" pageNumber="469" type="nomenclature">
<paragraph id="BF7CAA5BFFB6FF94D1CDFC7C45C9FC7C" blockId="18.[151,507,907,967]" box="[151,507,907,934]" pageId="18" pageNumber="469">
<heading id="E4341D37FFB6FF94D1CDFC7C45C9FC7C" bold="true" box="[151,507,907,934]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="8DB77649FFB6FF94D1CDFC7C45C9FC7C" bold="true" box="[151,507,907,934]" pageId="18" pageNumber="469">
<taxonomicName id="78C3D1D8FFB6FF94D1CDFC7C4583FC7F" ID-CoL="84G48" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,433,907,933]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="8DB77649FFB6FF94D1CDFC7C4583FC7F" bold="true" box="[151,433,907,933]" italics="true" pageId="18" pageNumber="469">Parasesarma gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB6FF94D0E3FC7B45C9FC7C" box="[441,507,908,934]" pageId="18" pageNumber="469" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="BF7CAA5BFFB6FF94D1CDFC5945D3FC1D" blockId="18.[151,507,907,967]" box="[151,481,942,967]" pageId="18" pageNumber="469">
(
<figureCitation id="27F8B6DEFFB6FF94D1C5FC5944C9FC1D" box="[159,251,942,967]" captionStart="FIGURE 11" captionStartId="19.[151,250,1654,1676]" captionTargetBox="[152,1435,481,1633]" captionTargetId="figure@19.[152,1435,481,1633]" captionTargetPageId="19" captionText="FIGURE 11. Parasesarma gracilipes n. sp.. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="18" pageNumber="469">Figs. 11</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D05DFC584517FC1D" box="[263,293,943,967]" captionStart="FIGURE 12" captionStartId="20.[151,250,1613,1635]" captionTargetBox="[177,1359,226,1572]" captionTargetId="figure@20.[152,1435,193,1592]" captionTargetPageId="20" captionText="FIGURE 12. Parasesarma gracilipes n. sp., holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440733/files/figure.png" pageId="18" pageNumber="469">12</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D06BFC584552FC1D" box="[305,352,943,967]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="18" pageNumber="469">19E</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D037FC5845ADFC1D" box="[365,415,943,967]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="18" pageNumber="469">20D</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D0F1FC5845EBFC1D" box="[427,473,943,967]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="18" pageNumber="469">21E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB6FF94D1CDFC004149FB90" pageId="18" pageNumber="469" type="reference_group">
<paragraph id="BF7CAA5BFFB6FF94D1CDFC0046BBFBF1" blockId="18.[151,1436,1014,1098]" pageId="18" pageNumber="469">
<treatmentCitationGroup id="9FD38D75FFB6FF94D1CDFC0046BBFBF1" pageId="18" pageNumber="469">
<taxonomicName id="78C3D1D8FFB6FF94D1CDFC0046C4FBD6" ID-CoL="7Z29P" authority=", De Man 1902: 534" authorityName="De Man" authorityPageNumber="534" authorityYear="1902" box="[151,758,1014,1037]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="leptosoma" subGenus="Parasesarma">
<emphasis id="8DB77649FFB6FF94D1CDFC0044DCFBD6" box="[151,238,1015,1036]" italics="true" pageId="18" pageNumber="469">Sesarma</emphasis>
(
<emphasis id="8DB77649FFB6FF94D05FFC0F45BEFBD7" box="[261,396,1016,1037]" italics="true" pageId="18" pageNumber="469">Parasesarma</emphasis>
)
<emphasis id="8DB77649FFB6FF94D0F9FC014639FBD6" box="[419,523,1014,1036]" italics="true" pageId="18" pageNumber="469">leptosoma</emphasis>
,
<treatmentCitation id="3E628C4AFFB6FF94D345FC0046C4FBD6" author="" box="[543,758,1014,1036]" page="534" pageId="18" pageNumber="469">
<bibRefCitation id="DB52D7AAFFB6FF94D345FC00468CFBD6" box="[543,702,1014,1036]" pageId="18" pageNumber="469" refId="ref17427" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel-gesammelten Dekapoden und Stomatopoden. In: Kukenthal, W. (Ed), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen Der Senckenbergischen Naturforschenden Gesellschaft, 25 (3), 467 - 929, pls. 19 - 27." type="journal article">De Man 1902</bibRefCitation>
: 534
</treatmentCitation>
</taxonomicName>
.—Nobili 1905: 497.—Rathbun 1910: 309, pl. 4, fig. 1. Not
<taxonomicName id="78C3D1D8FFB6FF94D19DFBE146B7FBF1" ID-CoL="7XX38" authority="(Hilgendorf, 1869)" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[199,645,1045,1067]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFB6FF94D19DFBE1458EFBF1" box="[199,444,1045,1067]" italics="true" pageId="18" pageNumber="469">Parasesarma leptosoma</emphasis>
(
<bibRefCitation id="DB52D7AAFFB6FF94D093FBE2464FFBF1" box="[457,637,1045,1067]" pageId="18" pageNumber="469" refId="ref16852" refString="Hilgendorf, F. (1869) Crustaceen. In: C. V. D. Decken (Ed), Reisen in Ost-Afrika in den Jahren 1859 - 1865. Heidelberg - Leipzig, pp. 67 - 116. http: // dx. doi. org / 10.5962 / bhl. title. 10640" type="book chapter">Hilgendorf, 1869</bibRefCitation>
)
</taxonomicName>
.
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</paragraph>
<paragraph id="BF7CAA5BFFB6FF94D1CDFBC24149FB90" blockId="18.[151,1436,1014,1098]" box="[151,1403,1076,1098]" pageId="18" pageNumber="469">
<taxonomicName id="78C3D1D8FFB6FF94D1CDFBC24686FB90" ID-CoL="7XX38" authority=", Rahayu &amp; Setyadi 2009: 51" authorityName="Rahayu &amp; Setyadi" authorityPageNumber="51" authorityYear="2009" box="[151,692,1076,1098]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFB6FF94D1CDFBC245BEFB90" box="[151,396,1076,1098]" italics="true" pageId="18" pageNumber="469">Parasesarma leptosoma</emphasis>
, Rahayu &amp; Setyadi 2009: 51
</taxonomicName>
, unnumbered figure. Not
<taxonomicName id="78C3D1D8FFB6FF94D2E3FBC24145FB90" ID-CoL="7XX38" authority="(Hilgendorf, 1869)" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[953,1399,1076,1098]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFB6FF94D2E3FBC2409CFB90" box="[953,1198,1076,1098]" italics="true" pageId="18" pageNumber="469">Parasesarma leptosoma</emphasis>
(
<bibRefCitation id="DB52D7AAFFB6FF94D5E1FBC34142FB90" box="[1211,1392,1076,1098]" pageId="18" pageNumber="469" refId="ref16852" refString="Hilgendorf, F. (1869) Crustaceen. In: C. V. D. Decken (Ed), Reisen in Ost-Afrika in den Jahren 1859 - 1865. Heidelberg - Leipzig, pp. 67 - 116. http: // dx. doi. org / 10.5962 / bhl. title. 10640" type="book chapter">Hilgendorf, 1869</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB6FF94D1CDFB814611FA99" pageId="18" pageNumber="469" type="materials_examined">
<paragraph id="BF7CAA5BFFB6FF94D1CDFB814611FA99" blockId="18.[151,1437,1142,2031]" pageId="18" pageNumber="469">
<materialsCitation id="0FABA006FFB6FF94D1CDFB814580FB69" ID-GBIF-Occurrence="1913889793" collectingDate="2005-06-11" collectionCode="MZB" collectorName="D. L. Rahayu" country="Indonesia" county="Material" location="Ajkwa River" municipality="Cru" pageId="18" pageNumber="469" specimenCount="1" stateProvince="Papua" typeStatus="holotype">
<emphasis id="8DB77649FFB6FF94D1CDFB8145B1FB55" bold="true" box="[151,387,1142,1167]" pageId="18" pageNumber="469">
<collectingCounty id="561DD2D7FFB6FF94D1CDFB8144CDFB55" box="[151,255,1142,1167]" pageId="18" pageNumber="469">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="607814F9FFB6FF94D0D6FB804616FB55" box="[396,548,1143,1167]" pageId="18" pageNumber="469" type="holotype">HOLOTYPE</typeStatus>
: male (13.6×
<quantity id="783B07BEFFB6FF94D3E1FB814711FB55" box="[699,803,1142,1167]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="18" pageNumber="469" unit="mm" value="11.9">11.9 mm</quantity>
) (
<collectionCode id="D9D2329EFFB6FF94D266FB804748FB55" box="[828,890,1143,1167]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="18" pageNumber="469">MZB</collectionCode>
<collectingMunicipality id="5F183021FFB6FF94D2DFFB80479DFB55" box="[901,943,1143,1167]" pageId="18" pageNumber="469">Cru</collectingMunicipality>
4803),
<collectingCountry id="C7D4EACBFFB6FF94D550FB814049FB55" box="[1034,1147,1142,1167]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
:
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:
<location id="BA1CFC80FFB6FF94D585FB814141FB55" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812:BA1CFC80FFB6FF94D585FB814141FB55" box="[1247,1395,1142,1167]" country="Indonesia" county="Material" municipality="Cru" name="Ajkwa River" pageId="18" pageNumber="469" stateProvince="Papua">Ajkwa River</location>
,
<collectingDate id="DB397573FFB6FF94D4DBFB80453DFB69" pageId="18" pageNumber="469" value="2005-06-11">
<date id="CB7D8C9BFFB6FF94D4DBFB80453DFB69" pageId="18" pageNumber="469" value="2005-06-11">11 June 2005</date>
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,
<collectorName id="1236CF8DFFB6FF94D046FB6C4580FB69" box="[284,434,1178,1203]" pageId="18" pageNumber="469">D.L. Rahayu</collectorName>
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.
<materialsCitation id="0FABA006FFB6FF94D0E5FB6C4583FB0D" ID-GBIF-Occurrence="1913889791" collectingDate="1999-08-03" collectionCode="ZRC" collectorName="G. Setyadi." country="Indonesia" location="Ajkwa River" pageId="18" pageNumber="469" specimenCode="ZRC 2000.1881" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="607814F9FFB6FF94D0E5FB6C466CFB69" box="[447,606,1179,1203]" pageId="18" pageNumber="469" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="A9C561D2FFB6FF94D330FB6C46FCFB69" box="[618,718,1178,1203]" pageId="18" pageNumber="469" type="female">1 female</specimenCount>
(12.5×
<quantity id="783B07BEFFB6FF94D27BFB6D47BBFB69" box="[801,905,1178,1203]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="18" pageNumber="469" unit="mm" value="10.7">10.7 mm</quantity>
) (
<specimenCode id="EF650220FFB6FF94D2F8FB6C406DFB69" box="[930,1119,1179,1203]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="469">ZRC 2000.1881</specimenCode>
),
<collectingCountry id="C7D4EACBFFB6FF94D52EFB6D40D7FB69" box="[1140,1253,1178,1203]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
:
<location id="BA1CFC80FFB6FF94D5A8FB6D41B7FB69" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812:BA1CFC80FFB6FF94D5A8FB6D41B7FB69" box="[1266,1413,1178,1203]" country="Indonesia" name="Ajkwa River" pageId="18" pageNumber="469">Ajkwa River</location>
<collectingDate id="DB397573FFB6FF94D4D4FB6C4518FB0D" pageId="18" pageNumber="469" value="1999-08-03">
<date id="CB7D8C9BFFB6FF94D4D4FB6C4518FB0D" pageId="18" pageNumber="469" value="1999-08-03">3 August 1999</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB6FF94D06FFB484583FB0D" box="[309,433,1214,1239]" pageId="18" pageNumber="469">G. Setyadi.</collectorName>
</materialsCitation>
—
<materialsCitation id="0FABA006FFB6FF94D08AFB4945F6FB21" ID-GBIF-Occurrence="1913889805" collectingDate="1999-08-12" collectionCode="ZRC" collectorName="G. Setyadi." country="Indonesia" location="Minajerwi" pageId="18" pageNumber="469" specimenCode="ZRC 2000.1882" specimenCount="1" specimenCount-male="1" stateProvince="Papua" typeStatus="paratype">
<specimenCount id="A9C561D2FFB6FF94D08AFB494624FB0D" box="[464,534,1214,1239]" pageId="18" pageNumber="469" type="male">1 male</specimenCount>
(12.6×
<quantity id="783B07BEFFB6FF94D33DFB4946FFFB0D" box="[615,717,1214,1239]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="18" pageNumber="469" unit="mm" value="10.9">10.9 mm</quantity>
) (
<specimenCode id="EF650220FFB6FF94D3B9FB4847ACFB0D" box="[739,926,1215,1239]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="469">ZRC 2000.1882</specimenCode>
),
<collectingCountry id="C7D4EACBFFB6FF94D2EBFB494010FB0D" box="[945,1058,1214,1239]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
:
<location id="BA1CFC80FFB6FF94D577FB494097FB0D" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812:BA1CFC80FFB6FF94D577FB494097FB0D" box="[1069,1189,1214,1239]" country="Indonesia" name="Minajerwi" pageId="18" pageNumber="469" stateProvince="Papua">Minajerwi</location>
,
<collectingRegion id="7D0764B9FFB6FF94D5EAFB4840C5FB0C" box="[1200,1271,1215,1238]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="469">Papua</collectingRegion>
, mangrove,
<collectingDate id="DB397573FFB6FF94D4DAFB484502FB21" pageId="18" pageNumber="469" value="1999-08-12">
<date id="CB7D8C9BFFB6FF94D4DAFB484502FB21" pageId="18" pageNumber="469" value="1999-08-12">12 August 1999</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB6FF94D018FB1445F6FB21" box="[322,452,1250,1275]" pageId="18" pageNumber="469">G. Setyadi.</collectorName>
</materialsCitation>
—
<materialsCitation id="0FABA006FFB6FF94D0B9FB15466AFAC5" ID-GBIF-Occurrence="1913889800" collectingDate="2000-04-04" collectionCode="ZRC" collectorName="D. L. Rahayu." country="Indonesia" location="Kamora" pageId="18" pageNumber="469" specimenCode="ZRC 2000.1883" specimenCount="1" specimenCount-male="1" stateProvince="Papua" typeStatus="paratype">
<specimenCount id="A9C561D2FFB6FF94D0B9FB15461DFB21" box="[483,559,1250,1275]" pageId="18" pageNumber="469" type="male">1 male</specimenCount>
(14.2×12.0 mm) (
<specimenCode id="EF650220FFB6FF94D255FB1447E3FB21" box="[783,977,1251,1275]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="469">ZRC 2000.1883</specimenCode>
),
<collectingCountry id="C7D4EACBFFB6FF94D2B0FB154069FB21" box="[1002,1115,1250,1275]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
:
<collectingRegion id="7D0764B9FFB6FF94D536FB144081FB20" box="[1132,1203,1251,1274]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="469">Papua</collectingRegion>
,
<location id="BA1CFC80FFB6FF94D59FFB14412DFB20" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812:BA1CFC80FFB6FF94D59FFB14412DFB20" box="[1221,1311,1251,1274]" country="Indonesia" name="Kamora" pageId="18" pageNumber="469" stateProvince="Papua">Kamora</location>
mangrove
<collectingCountry id="C7D4EACBFFB6FF94D1CDFAF14535FAC5" box="[151,263,1286,1311]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
,
<collectingDate id="DB397573FFB6FF94D04FFAF0459FFAC5" box="[277,429,1286,1311]" pageId="18" pageNumber="469" value="2000-04-04">
<date id="CB7D8C9BFFB6FF94D04FFAF0459FFAC5" box="[277,429,1286,1311]" pageId="18" pageNumber="469" value="2000-04-04">4 April 2000</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB6FF94D0E1FAF0466AFAC5" box="[443,600,1286,1311]" pageId="18" pageNumber="469">D.L. Rahayu.</collectorName>
</materialsCitation>
—
<materialsCitation id="0FABA006FFB6FF94D32CFAF14611FA99" ID-GBIF-Occurrence="1913889788" collectingDate="2005-06-11" collectionCode="ZRC" collectorName="D. L. Rahayu." country="Indonesia" location="Ajkwa River" pageId="18" pageNumber="469" specimenCode="ZRC 2018.0793" specimenCount="1" specimenCount-female="1" stateProvince="Papua" typeStatus="paratype">
<specimenCount id="A9C561D2FFB6FF94D32CFAF146E6FAC5" box="[630,724,1286,1311]" pageId="18" pageNumber="469" type="female">1 female</specimenCount>
(13.4×
<quantity id="783B07BEFFB6FF94D272FAF047A2FAC5" box="[808,912,1287,1311]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="18" pageNumber="469" unit="mm" value="11.7">11.7 mm</quantity>
) (
<specimenCode id="EF650220FFB6FF94D2F3FAF04055FAC5" box="[937,1127,1287,1311]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="469">ZRC 2018.0793</specimenCode>
),
<collectingCountry id="C7D4EACBFFB6FF94D527FAF140DCFAC5" box="[1149,1262,1286,1311]" name="Indonesia" pageId="18" pageNumber="469">Indonesia</collectingCountry>
:
<collectingRegion id="7D0764B9FFB6FF94D5A1FAF04176FAC4" box="[1275,1348,1287,1310]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="469">Papua</collectingRegion>
:
<location id="BA1CFC80FFB6FF94D408FAF144E5FA99" LSID="urn:lsid:plazi:treatment:376A1B4DFFB6FF95D1CDFC7C4548F812:BA1CFC80FFB6FF94D408FAF144E5FA99" country="Indonesia" name="Ajkwa River" pageId="18" pageNumber="469" stateProvince="Papua">Ajkwa River</location>
,
<collectingDate id="DB397573FFB6FF94D1B8FADC454BFA99" box="[226,377,1322,1347]" pageId="18" pageNumber="469" value="2005-06-11">
<date id="CB7D8C9BFFB6FF94D1B8FADC454BFA99" box="[226,377,1322,1347]" pageId="18" pageNumber="469" value="2005-06-11">11 June 2005</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB6FF94D0DEFADC4611FA99" box="[388,547,1322,1347]" pageId="18" pageNumber="469">D. L. Rahayu.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB6FF94D19DFAB940BCF859" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="BF7CAA5BFFB6FF94D19DFAB945CEF8E1" blockId="18.[151,1437,1142,2031]" pageId="18" pageNumber="469">
<emphasis id="8DB77649FFB6FF94D19DFAB94573FABD" bold="true" box="[199,321,1358,1383]" pageId="18" pageNumber="469">Diagnosis.</emphasis>
Carapace (
<figureCitation id="27F8B6DEFFB6FF94D0E5FAB9467DFABC" box="[447,591,1358,1383]" captionStart="FIGURE 11" captionStartId="19.[151,250,1654,1676]" captionTargetBox="[152,1435,481,1633]" captionTargetId="figure@19.[152,1435,481,1633]" captionTargetPageId="19" captionText="FIGURE 11. Parasesarma gracilipes n. sp.. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="18" pageNumber="469">Figs. 11A, E</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D301FAB846BEFABD" box="[603,652,1359,1383]" captionStart="FIGURE 12" captionStartId="20.[151,250,1613,1635]" captionTargetBox="[177,1359,226,1572]" captionTargetId="figure@20.[152,1435,193,1592]" captionTargetPageId="20" captionText="FIGURE 12. Parasesarma gracilipes n. sp., holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440733/files/figure.png" pageId="18" pageNumber="469">12A</figureCitation>
) squarish in general outline, 1.1 times broader than long; regions well defined, separated by shallow grooves; surface with scattered tufts of short setae; postfrontal region distinct, separated into 4 lobes by deep grooves; front deflexed downwards (
<figureCitation id="27F8B6DEFFB6FF94D2F4FA614077FA75" box="[942,1093,1430,1455]" captionStart="FIGURE 11" captionStartId="19.[151,250,1654,1676]" captionTargetBox="[152,1435,481,1633]" captionTargetId="figure@19.[152,1435,481,1633]" captionTargetPageId="19" captionText="FIGURE 11. Parasesarma gracilipes n. sp.. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="18" pageNumber="469">Figs. 11C, D</figureCitation>
) margin intensely concave in dorsal view; lateral margin straight, subparallel along most of length before curving to join almost straight posterior carapace margin; cornea extending or just reaching tip of external orbital tooth (
<figureCitation id="27F8B6DEFFB6FF94D56EFA2940FBFA2C" box="[1076,1225,1502,1527]" captionStart="FIGURE 11" captionStartId="19.[151,250,1654,1676]" captionTargetBox="[152,1435,481,1633]" captionTargetId="figure@19.[152,1435,481,1633]" captionTargetPageId="19" captionText="FIGURE 11. Parasesarma gracilipes n. sp.. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="18" pageNumber="469">Figs. 11A, E</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D58DFA284139FA2D" box="[1239,1291,1503,1527]" captionStart="FIGURE 12" captionStartId="20.[151,250,1613,1635]" captionTargetBox="[177,1359,226,1572]" captionTargetId="figure@20.[152,1435,193,1592]" captionTargetPageId="20" captionText="FIGURE 12. Parasesarma gracilipes n. sp., holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440733/files/figure.png" pageId="18" pageNumber="469">12A</figureCitation>
). Ischium of third maxilliped with shallow median sulcus, merus with distinct submedian ridge; exopod slender, tip reaching half-length of outer margin of merus, flagellum long. Male cheliped palm with 2 transverse pectinate crests (11 and 9 corneous teeth, respectively) on upper surface; upper surface of dactylus with 7 or 8 symmetrical, obliquely elongate dactylar tubercles, proximal 3 tubercles steep, sharp, the others tubercles large, distalmost tubercle indistinct (
<figureCitation id="27F8B6DEFFB6FF94D04AF9654595F970" box="[272,423,1682,1707]" captionStart="FIGURE 12" captionStartId="20.[151,250,1613,1635]" captionTargetBox="[177,1359,226,1572]" captionTargetId="figure@20.[152,1435,193,1592]" captionTargetPageId="20" captionText="FIGURE 12. Parasesarma gracilipes n. sp., holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440733/files/figure.png" pageId="18" pageNumber="469">Figs. 12C, D</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D0EFF96445D4F971" box="[437,486,1683,1707]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="18" pageNumber="469">19E</figureCitation>
). Ambulatory legs relatively slender for this species-complex, P3 and P4 about 1.6 times carapace width; P3 and P4 coxae without dense setae; P3 merus 3.1 times as long as broad; P3 propodus 5.9 times as long as broad; P3 dactylus 0.3 times length of propodus (
<figureCitation id="27F8B6DEFFB6FF94D2C0F92D4034F929" box="[922,1030,1754,1779]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="18" pageNumber="469">Fig. 20D</figureCitation>
). G1 relatively stout (
<figureCitation id="27F8B6DEFFB6FF94D45EF92D41AAF929" box="[1284,1432,1754,1779]" captionStart="FIGURE 12" captionStartId="20.[151,250,1613,1635]" captionTargetBox="[177,1359,226,1572]" captionTargetId="figure@20.[152,1435,193,1592]" captionTargetPageId="20" captionText="FIGURE 12. Parasesarma gracilipes n. sp., holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440733/files/figure.png" pageId="18" pageNumber="469">Figs. 12E–H</figureCitation>
,
<figureCitation id="27F8B6DEFFB6FF94D1CDF90844FAF8CD" box="[151,200,1791,1815]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="18" pageNumber="469">20E</figureCitation>
); apical process corneous, elongated, slightly bent at angle of 45°, stout, ending in rounded tip. G2 longer than quarter length of G1 (
<figureCitation id="27F8B6DEFFB6FF94D0D0F8D545C2F8E1" box="[394,496,1826,1851]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="18" pageNumber="469">Fig. 21E</figureCitation>
).
</paragraph>
<paragraph id="BF7CAA5BFFB6FF94D19DF8B140BCF859" blockId="18.[151,1437,1142,2031]" pageId="18" pageNumber="469">
<emphasis id="8DB77649FFB6FF94D19DF8B14542F885" bold="true" box="[199,368,1862,1887]" pageId="18" pageNumber="469">Colour in life.</emphasis>
General colour dark brown. Carapace mottled light brown and dark brown, palm of cheliped yellow or whitish yellow, fingers reddish orange or light orange (Rahayu &amp; Setyadi 2003).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB6FF95D19DF8794115FE77" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469" type="discussion">
<paragraph id="BF7CAA5BFFB6FF95D19DF87944C3FF0E" blockId="18.[151,1437,1142,2031]" lastBlockId="19.[151,1436,151,429]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<emphasis id="8DB77649FFB6FF94D19DF8794509F87D" bold="true" box="[199,315,1934,1959]" pageId="18" pageNumber="469">Remarks.</emphasis>
<taxonomicName id="78C3D1D8FFB6FF94D018F878467CF87D" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[322,590,1934,1959]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="8DB77649FFB6FF94D018F878467CF87D" box="[322,590,1934,1959]" italics="true" pageId="18" pageNumber="469">Parasesarma gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB6FF94D30FF87946A1F87D" box="[597,659,1934,1959]" pageId="18" pageNumber="469" rank="species">
<emphasis id="8DB77649FFB6FF94D30FF87946A1F87D" bold="true" box="[597,659,1934,1959]" pageId="18" pageNumber="469">n. sp.</emphasis>
</taxonomicNameLabel>
can easily be separated from other species in the species-complex by its relatively slender and longer ambulatory legs which are the longest among the members (
<figureCitation id="27F8B6DEFFB6FF94D5C7F8454134F811" box="[1181,1286,1970,1995]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="18" pageNumber="469">Fig. 20D</figureCitation>
). The ratio of merus length/width of the P3 is 3.1 (other species as 2.40–2.67) while the P3 propodus is 5.9 times as long as broad (other species as 4.1–5.0) (
<figureCitation id="27F8B6DEFFB7FF95D091FF604604FF6A" box="[459,566,151,176]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="19" pageNumber="470">Fig. 20D</figureCitation>
). Other features that distinguish this species from its congeners are given in
<tableCitation id="F2419FE0FFB7FF95D1CDFF4B44D9FF0E" box="[151,235,188,213]" captionStart="TABLE 1" captionStartId="7.[147,224,586,606]" captionText="TABLE 1. Comparison of major differences between members of the Parasesarma leptosoma species-complex." pageId="19" pageNumber="470">Table 1</tableCitation>
.
</paragraph>
<paragraph id="BF7CAA5BFFB7FF95D19DFF174115FE77" blockId="19.[151,1436,151,429]" pageId="19" pageNumber="470">
<bibRefCitation id="DB52D7AAFFB7FF95D19DFF174548FF22" box="[199,378,224,248]" pageId="19" pageNumber="470" refId="ref17427" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel-gesammelten Dekapoden und Stomatopoden. In: Kukenthal, W. (Ed), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen Der Senckenbergischen Naturforschenden Gesellschaft, 25 (3), 467 - 929, pls. 19 - 27." type="journal article">De Man (1902)</bibRefCitation>
reported
<taxonomicName id="78C3D1D8FFB7FF95D0B6FF1646CDFF22" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[492,767,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFB7FF95D0B6FF1646CDFF22" box="[492,767,223,248]" italics="true" pageId="19" pageNumber="470">Parasesarma leptosoma</emphasis>
</taxonomicName>
from Halmahera,
<collectingCountry id="C7D4EACBFFB7FF95D28CFF284071FF22" box="[982,1091,223,248]" name="Indonesia" pageId="19" pageNumber="470">Indonesia</collectingCountry>
[as
<taxonomicName id="78C3D1D8FFB7FF95D523FF1741BFFF22" authority="(Parasesarma)" authorityName="Say" authorityYear="1817" baseAuthorityName="Parasesarma" box="[1145,1421,224,248]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="genus">
<emphasis id="8DB77649FFB7FF95D523FF1740E8FF2D" box="[1145,1242,224,247]" italics="true" pageId="19" pageNumber="470">Sesarma</emphasis>
(
<emphasis id="8DB77649FFB7FF95D5B7FF1641B0FF22" box="[1261,1410,225,248]" italics="true" pageId="19" pageNumber="470">Parasesarma</emphasis>
)
</taxonomicName>
], and noticed that the specimens had more slender legs than those crabs from Africa he examined. Similarly, Rathbun (1910) recorded
<emphasis id="8DB77649FFB7FF95D09FFEDF45FFFEE5" box="[453,461,296,319]" italics="true" pageId="19" pageNumber="470">“</emphasis>
<taxonomicName id="78C3D1D8FFB7FF95D096FEDF47C2FE9A" authority="Hilgendorf" authorityName="Hilgendorf" authorityYear="1869" box="[460,1008,295,320]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="leptosoma" subGenus="Parasesarma">
<emphasis id="8DB77649FFB7FF95D096FEDF4606FEE5" box="[460,564,296,319]" italics="true" pageId="19" pageNumber="470">Sesarma</emphasis>
(
<emphasis id="8DB77649FFB7FF95D310FEDE46EDFE9A" box="[586,735,297,320]" italics="true" pageId="19" pageNumber="470">Parasesarma</emphasis>
)
<emphasis id="8DB77649FFB7FF95D3AEFED04755FE9A" box="[756,871,295,320]" italics="true" pageId="19" pageNumber="470">leptosoma</emphasis>
Hilgendorf
</taxonomicName>
, var.
<emphasis id="8DB77649FFB7FF95D573FEDF400AFEE5" box="[1065,1080,296,319]" italics="true" pageId="19" pageNumber="470">”</emphasis>
from Jobi Island, Dutch New
<collectingCountry id="C7D4EACBFFB7FF95D1CDFEBB44DAFEBF" box="[151,232,332,357]" name="Guinea-Bissau" pageId="19" pageNumber="470">Guinea</collectingCountry>
(now
<collectingRegion id="7D0764B9FFB7FF95D06DFEBA454CFEBE" box="[311,382,333,356]" country="Indonesia" name="Papua Barat" pageId="19" pageNumber="470">Papua</collectingRegion>
,
<collectingCountry id="C7D4EACBFFB7FF95D0D7FEBB45CCFEBF" box="[397,510,332,357]" name="Indonesia" pageId="19" pageNumber="470">Indonesia</collectingCountry>
), and commented that the specimens had more slender legs than what
<bibRefCitation id="DB52D7AAFFB7FF95D467FEBA44E9FE52" pageId="19" pageNumber="470" refId="ref17312" refString="Man, J. G. De (1889) Uber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452." type="journal article">De Man (1889</bibRefCitation>
; not
<bibRefCitation id="DB52D7AAFFB7FF95D04BFE874579FE52" box="[273,331,368,392]" pageId="19" pageNumber="470" refId="ref17427" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel-gesammelten Dekapoden und Stomatopoden. In: Kukenthal, W. (Ed), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen Der Senckenbergischen Naturforschenden Gesellschaft, 25 (3), 467 - 929, pls. 19 - 27." type="journal article">1902</bibRefCitation>
) mentioned in his description and figure. On this basis of these notes, and partly on the geography, we refer those previous records of
<taxonomicName id="78C3D1D8FFB7FF95D346FE624698FE77" authorityName="Hilgendorf" authorityYear="1869" box="[540,682,404,429]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFB7FF95D346FE624698FE77" box="[540,682,404,429]" italics="true" pageId="19" pageNumber="470">P. leptosoma</emphasis>
</taxonomicName>
from
<collectingCountry id="C7D4EACBFFB7FF95D3B4FE634769FE77" box="[750,859,404,429]" name="Indonesia" pageId="19" pageNumber="470">Indonesia</collectingCountry>
to
<taxonomicName id="78C3D1D8FFB7FF95D225FE624039FE77" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[895,1035,404,429]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="8DB77649FFB7FF95D225FE624039FE77" box="[895,1035,404,429]" italics="true" pageId="19" pageNumber="470">P. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB7FF95D548FE634063FE77" box="[1042,1105,404,429]" pageId="19" pageNumber="470" rank="species">
<emphasis id="8DB77649FFB7FF95D548FE634063FE77" bold="true" box="[1042,1105,404,429]" pageId="19" pageNumber="470">n. sp.</emphasis>
</taxonomicNameLabel>
for the time being.
</paragraph>
</subSubSection>
<caption id="EBBCFAD3FFB7FF95D1CDF98141A5F970" httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="19" pageNumber="470" startId="19.[151,250,1654,1676]" targetBox="[152,1435,481,1633]" targetPageId="19">
<paragraph id="BF7CAA5BFFB7FF95D1CDF98141A5F970" blockId="19.[151,1436,1654,1706]" pageId="19" pageNumber="470">
<emphasis id="8DB77649FFB7FF95D1CDF981452BF951" bold="true" box="[151,281,1654,1676]" pageId="19" pageNumber="470">FIGURE 11</emphasis>
.
<taxonomicName id="78C3D1D8FFB7FF95D07DF9804628F956" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[295,538,1654,1676]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="8DB77649FFB7FF95D07DF9804628F956" box="[295,538,1654,1676]" italics="true" pageId="19" pageNumber="470">Parasesarma gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB7FF95D379F981466FF951" box="[547,605,1654,1675]" pageId="19" pageNumber="470" rank="species">
<emphasis id="8DB77649FFB7FF95D379F981466FF951" bold="true" box="[547,605,1654,1675]" pageId="19" pageNumber="470">n. sp.</emphasis>
</taxonomicNameLabel>
. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views.
</paragraph>
</caption>
<subSubSection id="F7D9F9D0FFB7FF95D19DF92045C3F8CE" pageId="19" pageNumber="470" type="etymology">
<paragraph id="BF7CAA5BFFB7FF95D19DF92045C3F8CE" blockId="19.[151,1437,1751,1992]" pageId="19" pageNumber="470">
<emphasis id="8DB77649FFB7FF95D19DF920457CF92A" bold="true" box="[199,334,1751,1776]" pageId="19" pageNumber="470">Etymology.</emphasis>
The name is derived from the Latin &quot;
<emphasis id="8DB77649FFB7FF95D3A4F9204761F92A" box="[766,851,1751,1776]" italics="true" pageId="19" pageNumber="470">gracilis</emphasis>
&quot; (slender) and &quot;
<emphasis id="8DB77649FFB7FF95D54AF92F4007F935" box="[1040,1077,1752,1775]" italics="true" pageId="19" pageNumber="470">pes</emphasis>
&quot; (legs), referring to the slender ambulatory legs of the species.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB7FF95D19DF8E84653F886" pageId="19" pageNumber="470" type="distribution">
<paragraph id="BF7CAA5BFFB7FF95D19DF8E84653F886" blockId="19.[151,1437,1751,1992]" pageId="19" pageNumber="470">
<emphasis id="8DB77649FFB7FF95D19DF8E84552F8E2" bold="true" box="[199,352,1823,1848]" pageId="19" pageNumber="470">Distribution.</emphasis>
<collectingCountry id="C7D4EACBFFB7FF95D034F8E845EDF8E2" box="[366,479,1823,1848]" name="Indonesia" pageId="19" pageNumber="470">Indonesia</collectingCountry>
: Jobi Island (Rathbun 1910), Halmahera (
<bibRefCitation id="DB52D7AAFFB7FF95D286F8D740B8F8E2" box="[988,1162,1824,1848]" pageId="19" pageNumber="470" refId="ref17427" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel-gesammelten Dekapoden und Stomatopoden. In: Kukenthal, W. (Ed), Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen Der Senckenbergischen Naturforschenden Gesellschaft, 25 (3), 467 - 929, pls. 19 - 27." type="journal article">De Man 1902</bibRefCitation>
),
<collectingRegion id="7D0764B9FFB7FF95D5FEF8D740DBF8ED" box="[1188,1257,1824,1847]" country="Indonesia" name="Papua Barat" pageId="19" pageNumber="470">Papua</collectingRegion>
(Nobili, 1905; Rathbun 1910; Rahayu &amp; Setyadi 2009).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB7FF95D19DF8904548F812" pageId="19" pageNumber="470" type="biology_ecology">
<paragraph id="BF7CAA5BFFB7FF95D19DF8904548F812" blockId="19.[151,1437,1751,1992]" pageId="19" pageNumber="470">
<emphasis id="8DB77649FFB7FF95D19DF8904519F85A" bold="true" box="[199,299,1895,1920]" pageId="19" pageNumber="470">Ecology.</emphasis>
<taxonomicName id="78C3D1D8FFB7FF95D06EF89F4670F85A" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[308,578,1895,1920]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="8DB77649FFB7FF95D06EF89F4670F85A" box="[308,578,1895,1920]" italics="true" pageId="19" pageNumber="470">Parasesarma gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB7FF95D311F89046B9F85A" box="[587,651,1895,1920]" pageId="19" pageNumber="470" rank="species">
<emphasis id="8DB77649FFB7FF95D311F89046B9F85A" bold="true" box="[587,651,1895,1920]" pageId="19" pageNumber="470">n. sp.</emphasis>
</taxonomicNameLabel>
is active on muddy substrate in mangrove forest (Rahayu &amp; Setyadi 2009), occurring in the intertidal, area some distance from the river bank. They move fast between mangrove roots even during the day.
</paragraph>
</subSubSection>
</treatment>
</document>
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