treatments-xml/data/01/31/87/0131878A0723FF8CFA49FACB61374DFF.xml

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<document id="7E2B2E12EF86A12A02801C5AB9491C82" ID-CLB-Dataset="62482" ID-DOI="10.5281/zenodo.6603807" ID-GBIF-Dataset="916bf600-5bc0-438d-9c6c-7fff4c5a388f" ID-ISBN="978-84-941892-3-4" ID-Zenodo-Dep="6603807" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654101401716" checkinUser="carolina" docAuthor="Don E. Wilson, Thomas E. Lacher, Jr &amp; Russell A. Mittermeier" docDate="2016" docId="0131878A0723FF8CFA49FACB61374DFF" docLanguage="en" docName="hbmw_6_Geomyidae_0234.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I, Barcelona: Lynx Edicions" docTitle="Heterogeomys cavator" docType="treatment" docVersion="10" lastPageNumber="248" masterDocId="FD08FFF2072CFF83FFF3FFF96B0F4602" masterDocTitle="Geomyidae" masterLastPageNumber="269" masterPageNumber="234" pageNumber="248" updateTime="1699338600100" updateUser="ExternalLinkService">
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<mods:title id="1ACC7C89A25D4BED0507EE7D2D668848">Geomyidae</mods:title>
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<mods:namePart id="9FE6E29CBD898D40263A720917C6364A">Don E. Wilson</mods:namePart>
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<mods:namePart id="854030225E37CD6940A62F98878A0A7C">Thomas E. Lacher, Jr</mods:namePart>
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<mods:namePart id="67AB3999A1C867CB9A2B1602DEF0F732">Russell A. Mittermeier</mods:namePart>
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<mods:publisher id="2DAD39335A3379C222BCCCE1C59B128F">Lynx Edicions</mods:publisher>
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<mods:title id="2DDFDBBE6D3ACA95FFA0818752A37F6E">Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I</mods:title>
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<treatment id="0131878A0723FF8CFA49FACB61374DFF" ID-DOI="http://doi.org/10.5281/zenodo.6603729" ID-GBIF-Taxon="195708588" ID-Zenodo-Dep="6603729" LSID="urn:lsid:plazi:treatment:0131878A0723FF8CFA49FACB61374DFF" httpUri="http://treatment.plazi.org/id/0131878A0723FF8CFA49FACB61374DFF" lastPageNumber="248" pageId="15" pageNumber="248">
<subSubSection id="C18265170723FF8CFA49FACB6EFC4362" box="[1466,1523,1330,1376]" pageId="15" pageNumber="248" type="multiple">
<paragraph id="8927369C0723FF8CFA49FACB6EFC4362" blockId="15.[1464,2497,1330,1419]" box="[1466,1523,1330,1376]" pageId="15" pageNumber="248">
<heading id="D26F81F00723FF8CFA49FACB6EFC4362" box="[1466,1523,1330,1376]" pageId="15" pageNumber="248">
<figureCitation id="11A32A190723FF8CFA49FACB6EFC4362" box="[1466,1523,1330,1376]" captionStart="Plate 13: Geomyidae" captionStartId="14.[104,134,3318,3339]" captionTargetBox="[12,2731,15,3644]" captionTargetPageId="13" captionText="25. Michoacan Pocket Gopher (Zygogeomys trichopus), 26. Giant Pocket Gopher (Orthogeomys grandis), 27. Hispid Pocket Gopher (Heterogeomys hispidus), 28. Big Pocket Gopher (Heterogeomys lanius), 29. Chiriqui Pocket Gopher (Heterogeomys cavator), 30. Darien Pocket Gopher (Heterogeomys dariensis), 31. Cherries Pocket Gopher (Heterogeomys cherrier), 32. Variable Pocket Gopher (Heterogeomys heterodus), 33. Underwoods Pocket Gopher (Heterogeomys underwood), 34. Bullers Pocket Gopher (Pappogeomys bulleri), 35. Yellow-faced Pocket Gopher (Cratogeomys castanops), 36. Goldmans Pocket Gopher (Cratogeomys goldman), 37. Oriental Basin Pocket Gopher (Cratogeomys fulvescens), 38. Merriams Pocket Gopher (Cratogeomys merriami), 39. Cofre de Perote Pocket Gopher (Cratogeomys perotensis), 40. Volcan de Toluca Pocket Gopher (Cratogeomys planiceps), 41. Smoky Pocket Gopher (Cratogeomys fumosus)" figureDoi="http://doi.org/10.5281/zenodo.6603903" httpUri="https://zenodo.org/record/6603903/files/figure.png" pageId="15" pageNumber="248">29.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF9F6FACB6CF24362" box="[1541,2045,1330,1376]" pageId="15" pageNumber="248" type="vernacular_names">
<paragraph id="8927369C0723FF8CF9F6FACB6CF24362" blockId="15.[1464,2497,1330,1419]" box="[1541,2045,1330,1376]" pageId="15" pageNumber="248">
<heading id="D26F81F00723FF8CF9F6FACB6CF24362" box="[1541,2045,1330,1376]" pageId="15" pageNumber="248">
<vernacularName id="079B46B20723FF8CF9F6FACB6CF24362" box="[1541,2045,1330,1376]" pageId="15" pageNumber="248">Chiriqui Pocket Gopher</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF7B0FACB62CF4362" box="[2115,2496,1330,1376]" pageId="15" pageNumber="248" type="nomenclature">
<paragraph id="8927369C0723FF8CF7B0FACB62CF4362" blockId="15.[1464,2497,1330,1419]" box="[2115,2496,1330,1376]" pageId="15" pageNumber="248">
<heading id="D26F81F00723FF8CF7B0FACB62CF4362" box="[2115,2496,1330,1376]" pageId="15" pageNumber="248">
<taxonomicName id="4E984D1F0723FF8CF7B0FACB62CF4362" baseAuthorityName="Bangs" baseAuthorityYear="1902" box="[2115,2496,1330,1376]" class="Mammalia" family="Geomyidae" genus="Heterogeomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="species" species="cavator">
<emphasis id="BBECEA8E0723FF8CF7B0FACB62CF4362" box="[2115,2496,1330,1376]" italics="true" pageId="15" pageNumber="248">Heterogeomys cavator</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA4AFA8862124384" box="[1465,2333,1393,1414]" pageId="15" pageNumber="248" type="vernacular_names">
<paragraph id="8927369C0723FF8CFA4AFA8862124384" blockId="15.[1464,2497,1330,1419]" box="[1465,2333,1393,1414]" pageId="15" pageNumber="248">
<heading id="D26F81F00723FF8CFA4AFA8862124384" box="[1465,2333,1393,1414]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA4AFA886D0B4384" bold="true" box="[1465,1540,1393,1414]" pageId="15" pageNumber="248">French:</emphasis>
<vernacularName id="079B46B20723FF8CF9FEFA886DCE4384" box="[1549,1729,1393,1414]" pageId="15" pageNumber="248">Gaufre du Chiriqui</vernacularName>
/
<emphasis id="BBECEA8E0723FF8CF925FA886C3E4384" bold="true" box="[1750,1841,1393,1414]" pageId="15" pageNumber="248">German:</emphasis>
<vernacularName id="079B46B20723FF8CF8C9FA8863064384" box="[1850,2057,1393,1414]" pageId="15" pageNumber="248">Chiriqui-Taschenratte</vernacularName>
/
<emphasis id="BBECEA8E0723FF8CF7ECFA8863754384" bold="true" box="[2079,2170,1393,1414]" pageId="15" pageNumber="248">Spanish:</emphasis>
<vernacularName id="079B46B20723FF8CF78CFA8862124384" box="[2175,2333,1393,1414]" pageId="15" pageNumber="248">Tuza de Chiriqui</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF7E9FA43636B43FD" pageId="15" pageNumber="248" type="reference_group">
<paragraph id="8927369C0723FF8CF7E9FA43636B43FD" blockId="15.[2072,2669,1466,1888]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CF7E9FA4363BA43D5" bold="true" box="[2074,2229,1466,1495]" pageId="15" pageNumber="248">Taxonomy.</emphasis>
<taxonomicName id="4E984D1F0723FF8CF726FA43635143FD" ID-CoL="27b07b88-43e5-401a-bdd8-a932c3e9c912" authority="Bangs, 1902" authorityName="Bangs" authorityYear="1902" class="Mammalia" family="Geomyidae" genus="Macrogeomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="species" species="cavator">Macrogeomys cavator Bangs, 1902</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF78DFA1F63914027" pageId="15" pageNumber="248" type="materials_examined">
<paragraph id="8927369C0723FF8CF78DFA1F63914027" blockId="15.[2072,2669,1466,1888]" pageId="15" pageNumber="248">
<materialsCitation id="39F03CC10723FF8CF78DFA1F63914027" ID-GBIF-Occurrence="3799834312" pageId="15" pageNumber="248">“Boquete, 4,800 feet” (Chiriqui, Panama).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF7E9F9C962EB413B" pageId="15" pageNumber="248" type="discussion">
<paragraph id="8927369C0723FF8CF7E9F9C962EB413B" blockId="15.[2072,2669,1466,1888]" pageId="15" pageNumber="248">
Placed into subgenus
<taxonomicName id="4E984D1F0723FF8CF697F9C96104404F" authorityName="Merriam" authorityYear="1895" box="[2404,2571,1584,1613]" class="Mammalia" family="Geomyidae" genus="Macrogeomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="genus">Macrogeomys</taxonomicName>
.
<taxonomicName id="4E984D1F0723FF8CF5D1F9C963724076" pageId="15" pageNumber="248">Heterogeomys</taxonomicName>
considered a subgenus of
<taxonomicName id="4E984D1F0723FF8CF604F9AA6346409E" pageId="15" pageNumber="248">Orthogeomys</taxonomicName>
by J. L. Patton in 2005 but was resurrected to genus status by T. A. Spradling and colleagues in 2016. Molecular studies suggest that H. cavatoris sister to
<taxonomicName id="4E984D1F0723FF8CF61BF9086336413B" authorityName="Goldman" authorityYear="1912" class="Mammalia" family="Geomyidae" genus="Heterogeomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="species" species="dariensis">H. dariensis</taxonomicName>
. Three subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CF7EBF8C663DB41D6" pageId="15" pageNumber="248" type="distribution">
<caption id="DDE766140723FF8CF7EBF8C663DB41D6" ID-DOI="http://doi.org/10.5281/zenodo.6603871" ID-Zenodo-Dep="6603871" httpUri="https://zenodo.org/record/6603871/files/figure.png" inLine="true" pageId="15" pageNumber="248" targetBox="[1459,2049,1466,1879]" targetPageId="15">
<paragraph id="8927369C0723FF8CF7EBF8C662A74162" blockId="15.[2072,2669,1466,1888]" box="[2072,2472,1855,1888]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CF7EBF8C662A74162" bold="true" box="[2072,2472,1855,1888]" pageId="15" pageNumber="248">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="8927369C0723FF8CFA45F89C61464184" blockId="15.[1456,2665,1893,3069]" box="[1462,2633,1893,1926]" pageId="15" pageNumber="248">
<taxonomicName id="4E984D1F0723FF8CFA45F89C61464184" authority="Bangs, 1902" authorityName="Bangs" authorityYear="1902" box="[1462,2633,1893,1926]" class="Mammalia" family="Geomyidae" genus="Heterogeomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="subSpecies" species="cavator" subSpecies="cavator">H.c.cavatorBangs,1902—CordilleradeTalamancaofECostaRicaandWPanama.</taxonomicName>
</paragraph>
<paragraph id="8927369C0723FF8CFA45F86C614B41AC" blockId="15.[1456,2665,1893,3069]" box="[1462,2628,1941,1966]" pageId="15" pageNumber="248">
<taxonomicName id="4E984D1F0723FF8CFA45F86C614B41AC" authority="Goodwin, 1943" authorityName="Goodwin" authorityYear="1943" box="[1462,2628,1941,1966]" class="Mammalia" family="Geomyidae" genus="Heterogeomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="subSpecies" species="cavator" subSpecies="nigrescens">H.c.nigrescensGoodwin,1943—WendofCordilleradeTalamancainCCostaRica.</taxonomicName>
</paragraph>
<paragraph id="8927369C0723FF8CFA45F84A63DB41D6" blockId="15.[1456,2665,1893,3069]" box="[1462,2260,1971,2004]" pageId="15" pageNumber="248">
<taxonomicName id="4E984D1F0723FF8CFA45F84A6DF541D6" authority="Bangs, 1902" authorityName="Bangs" authorityYear="1902" box="[1462,1786,1971,2004]" class="Mammalia" family="Geomyidae" genus="Heterogeomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="subSpecies" species="cavator" subSpecies="pansa">H. c. pansa Bangs, 1902</taxonomicName>
— SE Costa Rica into SW Panama.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C18265170723FF8CFA46F82562034EEA" pageId="15" pageNumber="248" type="description">
<paragraph id="8927369C0723FF8CFA46F82562034EEA" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA46F8256DA141FF" bold="true" box="[1461,1710,2012,2045]" pageId="15" pageNumber="248">Descriptive notes.</emphasis>
Head-body 220-270 mm, tail 110-130 mm; weight 500-910 g. Dorsal pelage of the Chiriqui Pocket Gopher is dark brown, almost black, and ventral pelage 1s gray-brown. Pelage is dense and coarse in individuals from high elevations and sparse in individuals from low elevations. The Chiriqui Pocket Gopher has a fusiform body shape typical of all pocket gophers and possesses fur-lined cheek pouches that open external to the mouth. Anterior surface of each upper incisor has a single medial groove. Chromosomal complement has 2n = 44 and FN = 78.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA47F7096C834F5C" pageId="15" pageNumber="248" type="biology_ecology">
<paragraph id="8927369C0723FF8CFA47F7096C834F5C" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA47F7096D2C4F13" bold="true" box="[1460,1571,2288,2321]" pageId="15" pageNumber="248">Habitat.</emphasis>
Well-drained soils in a variety of habitats ranging from mixed conifer—hardwood forests at high elevations to open pasturelands near sea level. Elevational range is from nearsea level to ¢.3200 m.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA40F69F632A4C49" pageId="15" pageNumber="248" type="food_feeding">
<paragraph id="8927369C0723FF8CFA40F69F632A4C49" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA40F69F6DB44F85" bold="true" box="[1459,1723,2406,2439]" pageId="15" pageNumber="248">Food and Feeding.</emphasis>
There is no specific information available for this species, but the Chiriqui Pocket Gopher probably feeds on roots, tubers, stems, and leaves of most plants available within the vicinity ofits burrow system. As in all other pocket gophers, the burrow system is a series of shallow feeding tunnels radiating spoke-like from a deeper, central network that contains one or more nest chambers and several smaller chambers for storage of food orfecal pellets.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA41F5AB63F84C71" box="[1458,2295,2642,2675]" pageId="15" pageNumber="248" type="breeding">
<paragraph id="8927369C0723FF8CFA41F5AB63F84C71" blockId="15.[1456,2665,1893,3069]" box="[1458,2295,2642,2675]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA41F5AB6D374C71" bold="true" box="[1458,1592,2642,2675]" pageId="15" pageNumber="248">Breeding.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA43F58362FC4CEB" pageId="15" pageNumber="248" type="activity">
<paragraph id="8927369C0723FF8CFA43F58362FC4CEB" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA43F5836D904C99" bold="true" box="[1456,1695,2682,2715]" pageId="15" pageNumber="248">Activity patterns.</emphasis>
There is no specific information available for this species, but the Chiriqui Pocket Gopheris probably active at any hour of the day, with periods of peak activity around dawn and dusk. It does not hibernate and is active year-round.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA42F50962C74D85" pageId="15" pageNumber="248" type="biology_ecology">
<paragraph id="8927369C0723FF8CFA42F50962C74D85" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA42F509638E4D13" bold="true" box="[1457,2177,2800,2833]" pageId="15" pageNumber="248">Movements, Home range and Social organization.</emphasis>
There is no specific information available for this species, but the Chiriqui Pocket Gopheris likely solitary and aggressively territorial. Individuals probably leave their burrow systems only rarely, meaning that their home range is defined bysize and extent of their burrow system.
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA42F4686D574DD1" pageId="15" pageNumber="248" type="conservation">
<paragraph id="8927369C0723FF8CFA42F4686D574DD1" blockId="15.[1456,2665,1893,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA42F4686C044DAC" bold="true" box="[1457,1803,2961,2990]" pageId="15" pageNumber="248">Status and Conservation.</emphasis>
Classified as Least Concern on The [IUCN Red List (as
<taxonomicName id="4E984D1F0723FF8CF603F4686D434DD1" baseAuthorityName="Bangs" baseAuthorityYear="1902" class="Mammalia" family="Geomyidae" genus="Orthogeomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="248" phylum="Chordata" rank="species" species="cavator">Orthogeomys cavator</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C18265170723FF8CFA42F41D61374DFF" box="[1457,2616,3044,3069]" pageId="15" pageNumber="248" type="bibRefCitation_list">
<paragraph id="8927369C0723FF8CFA42F41D61374DFF" blockId="15.[1456,2665,1893,3069]" box="[1457,2616,3044,3069]" pageId="15" pageNumber="248">
<emphasis id="BBECEA8E0723FF8CFA42F41D6D454DFF" bold="true" box="[1457,1610,3044,3069]" pageId="15" pageNumber="248">Bibliography.</emphasis>
Hafner (1991), Patton (2005b), Reid (1997), Samudio &amp; Pino (2008b), Spradling et al. (2016).
</paragraph>
</subSubSection>
</treatment>
</document>