<documentid="1B7D5178E88915634DD4AFB86278E3AC"ID-CLB-Dataset="50384"ID-DOI="10.5281/zenodo.173868"ID-GBIF-Dataset="3057cbcd-031a-4a23-b2df-c3f768da9262"ID-ISSN="11755326"ID-Zenodo-Dep="173868"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="guilherme"checkinTime="1459785867585"checkinUser="plazi"docAuthor="González, Guillermo & Vandenberg, Natalia J."docDate="2006"docId="527187C4101FBC740650FB7E8261FDA0"docLanguage="en"docName="zt01311p050.pdf.imd"docOrigin="Zootaxa 1311"docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article"docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF"docStyleName="Zootaxa.2001-2006.journal_article"docStyleVersion="3"docTitle="Cycloneda germainii González & Vandenberg 2006, new combination"docType="treatment"docVersion="16"lastPageNumber="35"masterDocId="AE48FFBC100CBC630758FFE38774FFD6"masterDocTitle="Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries"masterLastPageNumber="50"masterPageNumber="13"pageNumber="32"updateTime="1738591164360"updateUser="guilherme">
<mods:titleid="6360051444AA2507C057976EA37F2F1B">Review of lady beetles in the Cycloneda germainii species complex (Coleoptera; Coccinellidae: Coccinellinae: Coccinellini) with descriptions of new and unusual species from Chile and surrounding countries</mods:title>
<taxonomicNameid="1DD84D51101FBC700650FB7E8502FB61"authority="(Crotch)"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[264,630,1181,1207]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="19"pageNumber="32"phylum="Arthropoda"rank="species"species="germainii"status="comb. nov.">
<figureCitationid="42E32A57101FBC700649FB22860DFB0D"box="[273,377,1217,1243]"captionStart="FIGURE 1"captionStartId="4.[264,367,1784,1808]"captionTargetBox="[344,1263,723,1752]"captionTargetId="figure@4.[279,1332,713,1777]"captionTargetPageId="4"captionText="FIGURE 1. Habitus views of Cycloneda species: A, C. lacrimosa González & Vandenberg, new species, male, paratype (3.0 mm); B, C. disconsolata Vandenberg & González, new species, female, holotype (3.3 mm); C, C. patagonica González & Vandenberg, new species, paratype (3.0 mm); D, C. germainii (Crotch), female, common form (3.5 mm)."httpUri="https://zenodo.org/record/173869/files/figure.png"pageId="19"pageNumber="32">Figs. 1D</figureCitation>
;
<figureCitationid="42E32A57101FBC7006DEFB2286D9FB0D"box="[390,429,1217,1243]"captionStart="FIGURE 3"captionStartId="15.[264,367,1565,1589]"captionTargetBox="[272,1316,323,1530]"captionTargetId="figure@15.[256,1331,310,1551]"captionTargetPageId="15"captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range."httpUri="https://zenodo.org/record/173871/files/figure.png"pageId="19"pageNumber="32">3D</figureCitation>
;
<figureCitationid="42E32A57101FBC7006E1FB228682FB0D"box="[441,502,1217,1243]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="19"pageNumber="32">4I–L</figureCitation>
;
<figureCitationid="42E32A57101FBC70055AFB22855CFB0D"box="[514,552,1217,1243]"captionStart="FIGURE 6"captionStartId="18.[264,368,1741,1765]"captionTargetBox="[266,1322,327,1740]"captionTargetId="figure@18.[264,1323,305,1746]"captionTargetPageId="18"captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González & Vandenberg, new species, holotype; B, C. patagonica González & Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924357"httpUri="https://zenodo.org/record/3924357/files/figure.png"pageId="19"pageNumber="32">6C</figureCitation>
;
<figureCitationid="42E32A57101FBC70056CFB22852EFB0D"box="[564,602,1217,1243]"captionStart="FIGURE 8"captionStartId="25.[264,368,1723,1747]"captionTargetBox="[294,1301,321,1698]"captionTargetId="figure@25.[284,1309,311,1730]"captionTargetPageId="25"captionText="FIGURE 8. Female genitalia of Cycloneda species (oriented with anterior end pointing left): A, C. lacrimosa González & Vandenberg, new species, allotype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species; D, Cycloneda germainii (Crotch); E, C. eryngii (Mulsant); F, C. sicardi (Brèthes), specimen from Tacuil, Salta; G, C. boliviana (Mulsant), specimen from Molinos, Salta."figureDoi="http://doi.org/10.5281/zenodo.3924361"httpUri="https://zenodo.org/record/3924361/files/figure.png"pageId="19"pageNumber="32">8D</figureCitation>
;
<figureCitationid="42E32A57101FBC70053FFB2285FFFB0D"box="[615,651,1217,1243]"captionStart="FIGURE 9"captionStartId="26.[264,368,885,909]"captionTargetBox="[276,1316,321,884]"captionTargetId="figure@26.[264,1323,305,886]"captionTargetPageId="26"captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924340"httpUri="https://zenodo.org/record/3924340/files/figure.png"pageId="19"pageNumber="32">9E</figureCitation>
;
<figureCitationid="42E32A57101FBC7005CFFB2285C1FB0D"box="[663,693,1217,1243]"captionStart="FIGURE 10"captionStartId="27.[264,368,1868,1892]"captionTargetBox="[287,1302,680,1870]"captionTargetId="figure@27.[287,1312,671,1870]"captionTargetPageId="27"captionText="FIGURE 10. Distributions of Cycloneda species."figureDoi="http://doi.org/10.5281/zenodo.3924342"httpUri="https://zenodo.org/record/3924342/files/figure.png"pageId="19"pageNumber="32">10</figureCitation>
<taxonomicNameid="1DD84D51101FBC700650FAF285DAFAF1"authority="Crotch 1874: 106"authorityName="González & Vandenberg"authorityPageNumber="106"authorityYear="2006"baseAuthorityName="Crotch"box="[264,686,1296,1320]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="19"pageNumber="32"phylum="Arthropoda"rank="species"species="germainii">
<bibRefCitationid="BE494B23101FBC7006A9FAF385DAFAF1"author="Crotch"box="[497,686,1296,1319]"firstAuthor="Crotch"pageId="19"pageNumber="32"refId="ref18126"refString="Crotch, G. R. (1874) A revision of the coleopterous family Coccinellidae. University Press, London, 311 pp."type="book"year="1874">Crotch 1874: 106</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700377FAD383BDFA91"author="Bosq"box="[1071,1225,1328,1351]"firstAuthor="Bosq"pageId="19"pageNumber="32"pagination="5 - 29"refId="ref18015"refString="Bosq, J. M. (1952) Enumeracion de predatores observados en la Republica Argentina (Coleopteros y Hemipteros). Ministerio de Agricultura y Ganaderia, Serie A, No 54, 5 - 29."type="journal article"year="1952">Bosq 1952:25</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700660FAB3868AFAB1"author="Brethes, J."box="[312,510,1360,1383]"firstAuthor="Brethes"journalOrPublisher="Anales del Museo Nacional de Historia Natural"pageId="19"pageNumber="32"pagination="145 - 175"part="33"refId="ref18054"refString="Brethes, J. (1925) Sur une collection de Coccinellidae (et un Phalacridae) du British Museum. Anales del Museo Nacional de Historia Natural, 33, 145 - 175."title="Sur une collection de Coccinellidae (et un Phalacridae) du British Museum"type="journal article"year="1925">Bréthes 1925: 152</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700551FAB38471FAB1"author="Korschefsky"box="[521,773,1360,1383]"firstAuthor="Korschefsky"pageId="19"pageNumber="32"refId="ref18255"refString="Korschefsky, R. (1932) Coleopterorum Catalogus, Pars 120, Coccinellidae II. W. Junk, Berlin, 659 pp."type="book"year="1932">Korschefsky 1932: 510</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700448FAB38378FAB1"author="Blackwelder"box="[784,1036,1360,1383]"firstAuthor="Blackwelder"pageId="19"pageNumber="32"pagination="343"refId="ref17970"refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Bulletin of the United States National Museum, 185, 343 - 550."type="journal article"year="1945">Blackwelder 1945: 454</bibRefCitation>
<bibRefCitationid="BE494B23101FBC7006A3FA9385E2FA51"author="Bosq"box="[507,662,1392,1415]"firstAuthor="Bosq"pageId="19"pageNumber="32"pagination="5 - 29"refId="ref18015"refString="Bosq, J. M. (1952) Enumeracion de predatores observados en la Republica Argentina (Coleopteros y Hemipteros). Ministerio de Agricultura y Ganaderia, Serie A, No 54, 5 - 29."type="journal article"year="1952">Bosq 1952: 13</bibRefCitation>
<taxonomicNameid="1DD84D51101FBC700650FA7285B1FA71"authority=": Gordon 1987: 12"authorityName="González & Vandenberg"authorityPageNumber="12"authorityYear="2006"baseAuthorityName="Crotch"box="[264,709,1424,1448]"class="Insecta"family="Coccinellidae"genus="Coccinellina"higherTaxonomySource="GBIF"kingdom="Animalia"order="Coleoptera"pageId="19"pageNumber="32"phylum="Arthropoda"rank="species"species="germainii">
<bibRefCitationid="BE494B23101FBC700555FA7385B1FA71"author="Gordon"box="[525,709,1424,1447]"firstAuthor="Gordon"pageId="19"pageNumber="32"pagination="1 - 46"refId="ref18222"refString="Gordon, R. D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46."type="journal article"year="1987">Gordon 1987: 12</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitationid="BE494B23101FBC700589FA7384FFFA71"author="Vandenberg"box="[721,907,1424,1447]"firstAuthor="Vandenberg"lastPageId="19"lastPageNumber="50"pageId="19"pageNumber="49"pagination="221 - 236"refId="ref18479"refString="Vandenberg, N. J. (2002) The new world genus Cycloneda Crotch (Coleoptera: Coccinellidae: Coccinellini): Historical review, new diagnosis, new generic and specific synonyms, and an improved key to North American species. Proceedings of the Entomological Society of Washington, 104 (1), 221 - 236."type="journal article"year="2002">Vandenberg 2002</bibRefCitation>
<bibRefCitationid="BE494B23101FBC7005DEFA538456FA11"author="Bosq"box="[646,802,1456,1479]"firstAuthor="Bosq"pageId="19"pageNumber="32"pagination="5 - 29"refId="ref18015"refString="Bosq, J. M. (1952) Enumeracion de predatores observados en la Republica Argentina (Coleopteros y Hemipteros). Ministerio de Agricultura y Ganaderia, Serie A, No 54, 5 - 29."type="journal article"year="1952">Bosq 1952: 25</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700683FA338502FA31"author="Bosq"box="[475,630,1488,1511]"firstAuthor="Bosq"pageId="19"pageNumber="32"pagination="5 - 29"refId="ref18015"refString="Bosq, J. M. (1952) Enumeracion de predatores observados en la Republica Argentina (Coleopteros y Hemipteros). Ministerio de Agricultura y Ganaderia, Serie A, No 54, 5 - 29."type="journal article"year="1952">Bosq 1952: 25</bibRefCitation>
<bibRefCitationid="BE494B23101FBC700503FA138423F9D1"author="Korschefsky"box="[603,855,1520,1543]"firstAuthor="Korschefsky"pageId="19"pageNumber="32"refId="ref18255"refString="Korschefsky, R. (1932) Coleopterorum Catalogus, Pars 120, Coccinellidae II. W. Junk, Berlin, 659 pp."type="book"year="1932">Korschefsky 1932: 510</bibRefCitation>
<bibRefCitationid="BE494B23101FBC70043AFA138314F9D1"author="Blackwelder"box="[866,1120,1520,1543]"firstAuthor="Blackwelder"pageId="19"pageNumber="32"pagination="343"refId="ref17970"refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Bulletin of the United States National Museum, 185, 343 - 550."type="journal article"year="1945">Blackwelder 1945: 454</bibRefCitation>
<bibRefCitationid="BE494B23101FBC7006BAF9F385FFF9F1"author="Brethes, J."box="[482,651,1552,1575]"firstAuthor="Brethes"journalOrPublisher="Anales del Museo Nacional de Historia Natural"pageId="19"pageNumber="32"pagination="145 - 175"part="33"refId="ref18054"refString="Brethes, J. (1925) Sur une collection de Coccinellidae (et un Phalacridae) du British Museum. Anales del Museo Nacional de Historia Natural, 33, 145 - 175."title="Sur une collection de Coccinellidae (et un Phalacridae) du British Museum"type="journal article"year="1925">Brèthes 1925: 6</bibRefCitation>
<bibRefCitationid="BE494B23101FBC70068AF9D385E3F991"author="Brethes, J."box="[466,663,1584,1607]"firstAuthor="Brethes"journalOrPublisher="Anales del Museo Nacional de Historia Natural"pageId="19"pageNumber="32"pagination="145 - 175"part="33"refId="ref18054"refString="Brethes, J. (1925) Sur une collection de Coccinellidae (et un Phalacridae) du British Museum. Anales del Museo Nacional de Historia Natural, 33, 145 - 175."title="Sur une collection de Coccinellidae (et un Phalacridae) du British Museum"type="journal article"year="1925">Brèthes 1925: 152</bibRefCitation>
Distinguished from other members of the genus by the combination of antenna composed of 10 antennomeres (
<figureCitationid="42E32A57101FBC7005D3F94285ABF96D"box="[651,735,1697,1723]"captionStart="FIGURE 9"captionStartId="26.[264,368,885,909]"captionTargetBox="[276,1316,321,884]"captionTargetId="figure@26.[264,1323,305,886]"captionTargetPageId="26"captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924340"httpUri="https://zenodo.org/record/3924340/files/figure.png"pageId="19"pageNumber="32">Fig.9E</figureCitation>
), pronotum with an even creamcolored anterior and lateral border, and elytron with a pale subapical mark positioned relatively close to the apex (
<figureCitationid="42E32A57101FBC70060BF91286A3F8DD"box="[339,471,1777,1803]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="19"pageNumber="32">Figs. 4I–L</figureCitation>
<figureCitationid="42E32A57101FBC70031EF8A283DEF88D"box="[1094,1194,1857,1883]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="19"pageNumber="32">Figs. 4E</figureCitation>
vs. 4I, 4H vs. 4L), but the latter species has 11 antennomeres, the lateral margin of the elytron more arcuate in dorsal view and, in profile (
<figureCitationid="42E32A571018BC7705B0FEA78433FE88"box="[744,839,324,350]"captionStart="FIGURE 3"captionStartId="15.[264,367,1565,1589]"captionTargetBox="[272,1316,323,1530]"captionTargetId="figure@15.[256,1331,310,1551]"captionTargetPageId="15"captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range."httpUri="https://zenodo.org/record/173871/files/figure.png"pageId="20"pageNumber="33">Fig. 3E</figureCitation>
), with the elytron more wedge shaped, broader in basal 1/4 than in apical 1/4. These two species also can be distinguished by subtle differences in the shape of the basal lobe of the male genitalia (
<figureCitationid="42E32A571018BC77033BFE7783CBFE78"box="[1123,1215,404,430]"captionStart="FIGURE 6"captionStartId="18.[264,368,1741,1765]"captionTargetBox="[266,1322,327,1740]"captionTargetId="figure@18.[264,1323,305,1746]"captionTargetPageId="18"captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González & Vandenberg, new species, holotype; B, C. patagonica González & Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924357"httpUri="https://zenodo.org/record/3924357/files/figure.png"pageId="20"pageNumber="33">Fig. 6C</figureCitation>
<taxonomicNameid="1DD84D511018BC770518FE5F844EFE03"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[576,826,444,469]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="33"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D511018BC770248FE5F86F8FE2B"authorityName="Gonzalez & Vandenberg"authorityYear="2006"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="20"pageNumber="33"phylum="Arthropoda"rank="species"species="patagonica">
. Form ovoid, moderately convex, feebly explanate, lateral margin weakly arcuate, appearing somewhat flattened in basal 2/3 beyond arc of humeral angle, apically tapered to blunt point; elytral, pronotal margins narrowly reflexed. Punctation on dorsal surfaces very fine, regular, with each puncture separated by 2.5–4.0X its diameter; surface between punctures shiny, faintly reticulate on head, pronotum, with only faint trace of reticulation visible on elytron.
Dorsal color pattern as follows: Head creamcolored except for irregular blackish band at base beginning at upper 1/4 of eye, pair of small separated blackish spots nearly touching clypeal margin, with vertical row of brown stipples between each spot and base of head; eye dark with hint of silvery reflections; labrum dark brown. Pronotum predominantly black; anterior, lateral margins with even creamcolored band of little more than ½ diameter of eye. Scutellum black. Elytron with ground color ferrugineous yellow orange with deeply incised irregular black figure on disk configured as follows (
<figureCitationid="42E32A571018BC77039FFC0F8269FBD0"box="[1223,1309,1004,1030]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="20"pageNumber="33">Fig. 4J</figureCitation>
): base with broadly arcuate emargination; outer border with anteromedial wedgeshaped emargination, apical one fourth with semicircular emargination; inner border with small semielliptical emargination near midline; with semidetached dark brown spot near anterolateral edge of black figure, situated just below humeral bulge; additional narrow diffuse brown mark beginning at suture, following lateral margin less than 1/4 its distance; union of dark, light areas irregular, suffused, dark reddish brown; with 4 rounded creamcolored marks occupying medial half of basal emargination, entirely filling 3 posterior emarginations; union of dark, light areas irregular, narrowly ferrugineous, suffused. Each puncture of dorsal surfaces with pinpoint of light brown at center, visible only in areas with light to medium background coloration; double to triple staggered row of punctures at inner margin of sutural band, sparsely scattered row along lateral margin beginning just outside of humeral bulge with more pronounced pigmentation. Anterior, lateral margins of pronotum narrowly transparent to light amber, all margins of elytron dark amber.
</paragraph>
<paragraphid="DA6736D21018BC770660F9FF833FF928"blockId="20.[264,1324,284,1870]"pageId="20"pageNumber="33">Ground color of ventral surfaces dark brown; elytral epipleuron, pronotal hypomeron except for triangular area at inside of base light yellow orange to creamcolored; mesepimeron creamcolored; mouthparts, antenna light amber brown; apex of maxillary palps, last two antennomeres darker brown; leg dark brown, with coxa, tarsus, protibia, apex of meso, metatibiae light reddish brown. Ventral surfaces including appendages clothed in sparse decumbent silvery white pubescence; hairs fine, long.</paragraph>
Eyes finely facetted, separated by 2¼X eye diameter; inner orbits nearly parallel in lower half, diverging at upper level. Antenna of 10 antennomeres (
<figureCitationid="42E32A571018BC770367F8D783EAF898"box="[1087,1182,1844,1870]"captionStart="FIGURE 9"captionStartId="26.[264,368,885,909]"captionTargetBox="[276,1316,321,884]"captionTargetId="figure@26.[264,1323,305,886]"captionTargetPageId="26"captionText="FIGURE 9. Left antenna of Cycloneda species, dorsal view: AC, C. lacrimosa Gonzlez & Vandenberg, new species (showing variation from 9 to 11 antennomeres); D, C. patagonica Gonzlez & Vandenberg, new species; E, Cycloneda germainii (Crotch); F, C. eryngii (Mulsant); G, C. sicardi (Brèthes); H, C. boliviana (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924340"httpUri="https://zenodo.org/record/3924340/files/figure.png"pageId="20"pageNumber="33">Fig. 9E</figureCitation>
), combined length slightly greater than distance between eyes; third antennomere elongate, about 1½X second, slightly shorter than third and fourth combined. Pronotum convex, lateral margin weakly reflexed at transparent border; pronotal outline with basal margin subsinuate, lateral margin strongly evenly arcuate, anterior margin subtrapezoidally emarginate, median part linear; anterior angles subtriangulate, projecting anteroventrally. Elytron somewhat elongate (
<figureCitationid="42E32A571019BC7606A3FE07850EFE28"box="[507,634,484,510]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="21"pageNumber="34">Figs. 4I–L</figureCitation>
), in dorsal view with humeral angle rounded, very weakly arcuate beyond to about apical 1/3, broadest near middle or just beyond, tapered to rounded apex; outer margin weakly explanate; in lateral view (
<figureCitationid="42E32A571019BC7604B6FDD7833EFD98"box="[1006,1098,564,590]"captionStart="FIGURE 3"captionStartId="15.[264,367,1565,1589]"captionTargetBox="[272,1316,323,1530]"captionTargetId="figure@15.[256,1331,310,1551]"captionTargetPageId="15"captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range."httpUri="https://zenodo.org/record/173871/files/figure.png"pageId="21"pageNumber="34">Fig. 3D</figureCitation>
) unevenly arcuate, dorsally somewhat flattened in anterior 2/3, more abruptly declivitous in posterior 1/3, of equal breadth at anterior, posterior 1/4; epipleuron approximately horizontal, very weakly ascending in anterior 1/3, descending near apex, weakly concave in medial half. Prosternum Tshaped, with median third of transverse basal piece convex, gradually becoming explanate in each lateral third; intercoxal process convex with fine superficial median sulcus along most of length. Mesosternum trapezoidal; anterior border approximately linear, with raised margin. Metasternum broad, with postmesocoxal line reaching lateral margin; shallowly transversely rugulose; discrimen present, distinct except for ends. Abdomen shortened semioval, broadest in anterior half of second ventrite; posterior margin of ventrites 1–4 linear, of 5 arcuately emarginate, 6 rounded at sides, flattened to very weakly arcuately emarginate at middle; postmetacoxal line of first abdominal ventrite curved posterolaterad, closely paralleling posterior margin for much of length, not attaining lateral margin. Tarsal claw with rectangular basal tooth.
<figureCitationid="42E32A571019BC76053CFB8785B5FBA8"box="[612,705,1124,1150]"captionStart="FIGURE 6"captionStartId="18.[264,368,1741,1765]"captionTargetBox="[266,1322,327,1740]"captionTargetId="figure@18.[264,1323,305,1746]"captionTargetPageId="18"captionText="FIGURE 6. Male genitalia of Cycloneda species (above, ventral and right lateral views of phallobase; below, right lateral view of sipho (=aedeagus)): A, C. lacrimosa González & Vandenberg, new species, holotype; B, C. patagonica González & Vandenberg, new species, holotype; C, C. germainii (Crotch); D, C. eryngii (Mulsant)."figureDoi="http://doi.org/10.5281/zenodo.3924357"httpUri="https://zenodo.org/record/3924357/files/figure.png"pageId="21"pageNumber="34">Fig. 6C</figureCitation>
): basal lobe elongate, roughly parallelsided in basal half, subapically slightly swollen with greatest width at apical one fourth; apex tapered, slightly attenuate; parameres slender, reaching three quarters distance to apex of basal lobe.
Similar to male except larger on average; head black with irregular creamcolored patch on lower half adjacent to eye, including eye canthus. Abdomen with posterior margin of ventrite 5 linear, exposed portion of ventrite 6 very short, apically rounded. Female genitalia as in figure 8D.
<figureCitationid="42E32A571019BC760329FA478385FA68"box="[1137,1265,1444,1470]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="21"pageNumber="34">Figs. 4I–L</figureCitation>
<taxonomicNameid="1DD84D511019BC7605F9FA2F844CFA33"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[673,824,1484,1509]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="34"phylum="Arthropoda"rank="species"species="germainii">
<bibRefCitationid="BE494B231019BC7604A3FA2F83EEFA30"author="Gordon"box="[1019,1178,1484,1510]"firstAuthor="Gordon"pageId="21"pageNumber="49"pagination="1 - 46"refId="ref18222"refString="Gordon, R. D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46."type="journal article"year="1987">Gordon 1987</bibRefCitation>
“type [blue label]/type chili, germ.” (
<collectionCodeid="BCC9AE171019BC760577FA1785F0F9D8"box="[559,644,1524,1550]"country="Chile"httpUri="http://grbio.org/cool/jgvc-yfsb"name="Universidad de Concepcion, Museo de Zoologia"pageId="21"pageNumber="34">UCCC</collectionCode>
<taxonomicNameid="1DD84D511019BC760321FA178278F9DB"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[1145,1292,1524,1549]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="34"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D511019BC7605CEF9A78478F988"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[662,780,1604,1630]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="46"phylum="Arthropoda"rank="species"species="germainii">Germainii</taxonomicName>
<taxonomicNameid="1DD84D511019BC760554F98F842BF950"authority="Crotch"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[524,863,1644,1670]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="34"phylum="Arthropoda"rank="species"species="germainii">Coccinella germainii Crotch</taxonomicName>
<taxonomicNameid="1DD84D511019BC760420F95F849BF900"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[888,1007,1724,1750]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="46"phylum="Arthropoda"rank="species"species="germainii">Germainii</taxonomicName>
<taxonomicNameid="1DD84D511019BC760650F8D7860BF898"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[264,383,1844,1870]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="21"pageNumber="46"phylum="Arthropoda"rank="species"species="germainii">Germainii</taxonomicName>
<bibRefCitationid="BE494B23101ABC7506BDFDD7850DFD98"author="Bosq"box="[485,633,564,590]"firstAuthor="Bosq"pageId="22"pageNumber="49"pagination="5 - 29"refId="ref18015"refString="Bosq, J. M. (1952) Enumeracion de predatores observados en la Republica Argentina (Coleopteros y Hemipteros). Ministerio de Agricultura y Ganaderia, Serie A, No 54, 5 - 29."type="journal article"year="1952">Bosq (1952)</bibRefCitation>
<figureCitationid="42E32A57101ABC75059CFD678452FD48"box="[708,806,644,670]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="22"pageNumber="35">Fig. 4K</figureCitation>
) appears to be nothing more than a less common, dark morph of
<taxonomicNameid="1DD84D51101ABC75056FFD4F85BAFD13"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[567,718,684,709]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
which turns up occasionally in the more southern populations.
<bibRefCitationid="BE494B23101ABC7506F1FD37852EFD38"author="Brethes, J."box="[425,602,724,750]"firstAuthor="Brethes"journalOrPublisher="Anales del Museo Nacional de Historia Natural"pageId="22"pageNumber="49"pagination="145 - 175"part="33"refId="ref18054"refString="Brethes, J. (1925) Sur une collection de Coccinellidae (et un Phalacridae) du British Museum. Anales del Museo Nacional de Historia Natural, 33, 145 - 175."title="Sur une collection de Coccinellidae (et un Phalacridae) du British Museum"type="journal article"year="1925">Brèthes (1925)</bibRefCitation>
had two specimens of the dark morph from “Valle del Lago Blanco” Patagonia, one of which he designated as the
<bibRefCitationid="BE494B23101ABC750440FC978377FC58"author="Korschefsky"box="[792,1027,884,910]"firstAuthor="Korschefsky"pageId="22"pageNumber="49"refId="ref18255"refString="Korschefsky, R. (1932) Coleopterorum Catalogus, Pars 120, Coccinellidae II. W. Junk, Berlin, 659 pp."type="book"year="1932">Korschefsky (1932)</bibRefCitation>
and
<bibRefCitationid="BE494B23101ABC750318FC978259FC58"author="Blackwelder"box="[1088,1325,884,910]"firstAuthor="Blackwelder"pageId="22"pageNumber="49"pagination="343"refId="ref17970"refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Bulletin of the United States National Museum, 185, 343 - 550."type="journal article"year="1945">Blackwelder (1945)</bibRefCitation>
, below). Based on penned annotations in a copy of
<bibRefCitationid="BE494B23101ABC750650FC0F86C6FBD0"author="Crotch"box="[264,434,1004,1030]"firstAuthor="Crotch"pageId="22"pageNumber="49"refId="ref18126"refString="Crotch, G. R. (1874) A revision of the coleopterous family Coccinellidae. University Press, London, 311 pp."type="book"year="1874">Crotch (1874)</bibRefCitation>
from the Smithsonian Entomology Library, Weise apparently concurred with the placement of
<taxonomicNameid="1DD84D51101ABC7504D2FBF78355FBFB"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[906,1057,1044,1069]"class="Insecta"family="Coccinellidae"genus="Coccinella"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D51101ABC750421FBDF830CFB83"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[889,1144,1084,1109]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D51101ABC750479FB8784CDFBAB"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[801,953,1124,1149]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D51101ABC7506A8FB5785F0FB1B"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[496,644,1204,1229]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<taxonomicNameid="1DD84D51101ABC7505CFFB3F8459FB23"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[663,813,1244,1269]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<figureCitationid="42E32A57101ABC7503DAFB3F83A8FB20"box="[1154,1244,1244,1270]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="22"pageNumber="35">Fig. 4L</figureCitation>
) has a nearly identical elytral color pattern to examples of
<figureCitationid="42E32A57101ABC750649FACF8605FA90"box="[273,369,1324,1350]"captionStart="FIGURE 4"captionStartId="16.[264,367,1625,1649]"captionTargetBox="[395,1202,315,1596]"captionTargetId="figure@16.[316,1279,307,1620]"captionTargetPageId="16"captionText="FIGURE 4. Elytral color patterns of Cycloneda species, left elytron: A–H, C. eryngii (Mulsant) (A–C, with reduced or broken maculae: A, from Rio Bueno, Valdivia; B, from San Pedro de Atacama; C, from Osorno; D–F, common aberrations; G, with confluent markings from Santiago; H, tricolored aberration from Baños de Cauquenes, Rancagua); I–L, C. germainii (Crotch) (I, with reduced markings from San Martin de los Andes, Neuquen; J, common aberration; K, melanic form, type of C. duplaris Berg; L, form with disconnected maculae, from Baños de Cauquenes, Rancagua); M–N, C. lacrimosa González & Vandenberg, new species, common aberrations; O–P, C. disconsolata Vandenberg & González, new species (O, paratype; P, holotype)."httpUri="https://zenodo.org/record/173872/files/figure.png"pageId="22"pageNumber="35">Fig. 4H</figureCitation>
<taxonomicNameid="1DD84D51101ABC75049AFAB78323FABB"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[962,1111,1364,1389]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<figureCitationid="42E32A57101ABC75057DFA9F850DFA40"box="[549,633,1404,1430]"captionStart="FIGURE 3"captionStartId="15.[264,367,1565,1589]"captionTargetBox="[272,1316,323,1530]"captionTargetId="figure@15.[256,1331,310,1551]"captionTargetPageId="15"captionText="FIGURE 3. Morphology of Cycloneda species. Lateral views of bodies (elytral length standardized to facilitate comparison), A–G: A, C. lacrimosa González & Vandenberg, new species, paratype; B, C. disconsolata Vandenberg & González, new species, holotype; C, C. patagonica González & Vandenberg, new species, paratype; D, Cycloneda germainii (Crotch), representative specimen; E, C. eryngii (Mulsant), representative specimen; F, C. sicardi (Brèthes), specimen from Salta; G, C. boliviana (Mulsant), holotype. Pronotal dorsofrontal views, H–L: H, C. lacrimosa González & Vandenberg, new species, paratype; I, C. boliviana (Mulsant), holotype; J, C. sicardi (Brèthes), specimen from El Rincón, Catamarca; K, C. sicardi (Brèthes), specimen from Tacuil, Salta; L, C. sicardi (Brèthes), common aberration in much of range."httpUri="https://zenodo.org/record/173871/files/figure.png"pageId="22"pageNumber="35">Fig.3D</figureCitation>
) and the elytron narrower, less arcuate basally in dorsal view. The possibility remains that this specimen represents yet another undescribed species closely allied to
<taxonomicNameid="1DD84D51101ABC75057DFA2F85CDFA33"authorityName="González & Vandenberg"authorityYear="2006"baseAuthorityName="Crotch"box="[549,697,1484,1509]"class="Insecta"family="Coccinellidae"genus="Cycloneda"kingdom="Animalia"order="Coleoptera"pageId="22"pageNumber="35"phylum="Arthropoda"rank="species"species="germainii">
<figureCitationid="42E32A57101ABC750368FA1783EEF9D8"box="[1072,1178,1524,1550]"captionStart="FIGURE 10"captionStartId="27.[264,368,1868,1892]"captionTargetBox="[287,1302,680,1870]"captionTargetId="figure@27.[287,1312,671,1870]"captionTargetPageId="27"captionText="FIGURE 10. Distributions of Cycloneda species."figureDoi="http://doi.org/10.5281/zenodo.3924342"httpUri="https://zenodo.org/record/3924342/files/figure.png"pageId="22"pageNumber="35">Fig. 10</figureCitation>
):
</emphasis>
<materialsCitationid="6AB03C8F101ABC75039AFA178387F9E0"ID-GBIF-Occurrence="2651531303"collectingDate="1963-01-06"collectingDateMax="1963-01-31"collectingDateMin="1963-01-06"collectorName="L. E. Pena"country="Chile"elevation="1500"location="Las Cabras"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Coquimbo">
<materialsCitationid="6AB03C8F101ABC75047BF9A78552F950"ID-GBIF-Occurrence="2651531324"collectorName="Banos & de Cauquenes & Rancagua"country="Chile"location="LIBERTADOR"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Libertador General Bernardo O'Higgins">
<collectingRegionid="181CF830101ABC75047BF9A784A9F988"box="[803,989,1604,1630]"country="Chile"name="Libertador General Bernardo O'Higgins"pageId="22"pageNumber="35">LIBERTADOR</collectingRegion>
<materialsCitationid="6AB03C8F101ABC75056DF98F84ECF978"ID-GBIF-Occurrence="2651531395"collectingDate="1976-07-15"collectingDateMax="1976-07-16"collectingDateMin="1976-07-15"collectorName="E. Chillan & H. F. Howden"country="Chile"elevation="1650"location="Shangrila"municipality="Nuble Prov"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Biobio">
<materialsCitationid="6AB03C8F101ABC7504F1F9778533F900"ID-GBIF-Occurrence="2651531405"collectingDate="1968-03-02"collectorName="Flint & Pena"country="Chile"location="Las Trancas"municipality="Nuble"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Biobio">
<materialsCitationid="6AB03C8F101ABC75036BF95F859FF928"ID-GBIF-Occurrence="2651531358"country="Chile"elevation="1900"location="Los Barros"municipality="Laguna del Laja"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Biobio">
<collectingMunicipalityid="3A03ACA8101ABC75036BF95F8387F900"box="[1075,1267,1724,1750]"pageId="22"pageNumber="35">Laguna del Laja</collectingMunicipality>
,
<locationid="DF076009101ABC7503A6F95F862DF928"LSID="urn:lsid:plazi:treatment:527187C4101FBC740650FB7E8261FDA0:DF076009101ABC7503A6F95F862DF928"country="Chile"municipality="Laguna del Laja"name="Los Barros"pageId="22"pageNumber="35"stateProvince="Biobio">Los Barros</location>
<materialsCitationid="6AB03C8F101ABC7506CCF8EF86F1F898"ID-GBIF-Occurrence="2651531412"collectingDate="1959-01"collectorName="L. E. Pena"country="Chile"county="Tolhuaca"location="Malleco"municipality="Curacautin"pageId="22"pageNumber="35"specimenCount="3"stateProvince="Araucania">
<materialsCitationid="6AB03C8F101ABC7506CCF8D784D8F898"ID-GBIF-Occurrence="2651531382"box="[404,940,1844,1870]"collectorName="Punta Arenas"country="Chile"location="MAGALLANES"pageId="22"pageNumber="35"specimenCount="1"stateProvince="Magallanes y de la Antartica Chilena">
<collectingRegionid="181CF830101ABC7506CCF8D78517F898"box="[404,611,1844,1870]"country="Chile"name="Magallanes y de la Antartica Chilena"pageId="22"pageNumber="35">MAGALLANES</collectingRegion>
<materialsCitationid="6AB03C8F101BBC7406C9FEA78413FE50"ID-GBIF-Occurrence="2651531424"collectingDate="1949-02-04"collectingDateMax="1978-02-27"collectingDateMin="1949-02-04"collectorName="Monros & Km. S. Pilolil & C. M. & O. S. Flint & Jr."country="Argentina"county="Confluencia"location="Limay"municipality="Traful"pageId="23"pageNumber="36"specimenCount="1"stateProvince="Neuquen">
<materialsCitationid="6AB03C8F101BBC7404D4FDEF8584FD98"ID-GBIF-Occurrence="2651531422"collectingDate="1980-02-21"collectorName="Fidalgo Willink & Claps Dominguez"country="Argentina"location="Rio San Julian"pageId="23"pageNumber="36"specimenCount="1"stateProvince="SANTA CRUZ">
<locationid="DF076009101BBC7404D4FDEF834EFDF0"LSID="urn:lsid:plazi:treatment:527187C4101FBC740650FB7E8261FDA0:DF076009101BBC7404D4FDEF834EFDF0"box="[908,1082,524,550]"country="Argentina"name="Rio San Julian"pageId="23"pageNumber="36"stateProvince="SANTA CRUZ">Rio San Julián</location>
<materialsCitationid="6AB03C8F101BBC740650FDBF8261FDA0"ID-GBIF-Occurrence="2651531384"box="[264,1301,604,630]"collectionCode="AMNH, BMNH, GGPC, MNHN, UCCC, USNM"country="Argentina"location="Valle del Lago Blanco"pageId="23"pageNumber="36"specimenCount="2"stateProvince="SANTA CRUZ">
<locationid="DF076009101BBC740650FDBF8564FDA0"LSID="urn:lsid:plazi:treatment:527187C4101FBC740650FB7E8261FDA0:DF076009101BBC740650FDBF8564FDA0"box="[264,528,604,630]"country="Argentina"name="Valle del Lago Blanco"pageId="23"pageNumber="36"stateProvince="SANTA CRUZ">Valle del Lago Blanco</location>
<collectionCodeid="BCC9AE17101BBC74059AFDBF8455FDA0"LSID="urn:lsid:biocol.org:col:34925"box="[706,801,604,630]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="23"pageNumber="36">AMNH</collectionCode>
<collectionCodeid="BCC9AE17101BBC7404B7FDBF833FFDA0"LSID="urn:lsid:biocol.org:col:34988"box="[1007,1099,604,630]"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="23"pageNumber="36">MNHN</collectionCode>
,
<collectionCodeid="BCC9AE17101BBC74030FFDBF83DEFDA0"box="[1111,1194,604,630]"country="Chile"httpUri="http://grbio.org/cool/jgvc-yfsb"name="Universidad de Concepcion, Museo de Zoologia"pageId="23"pageNumber="36">UCCC</collectionCode>
,
<collectionCodeid="BCC9AE17101BBC7403EEFDBF8279FDA0"LSID="urn:lsid:biocol.org:col:34871"box="[1206,1293,604,630]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871"name="Smithsonian Institution, National Museum of Natural History"pageId="23"pageNumber="36"type="Museum">USNM</collectionCode>
