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<document id="6FDFB255955FEE4B825BF04E2AAC2E5F" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFAC077554EAFE9E7CA3FD6B" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Paramaya De Haan 1837" docType="key" docVersion="4" lastPageNumber="155" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="155" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A192FFF9402B44824BBA371616A81C95">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:name id="F9CD8E606F0AD83F9F6240A5E8102FCB" type="personal">
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<mods:namePart id="79AD7DBCFCCDB266041E5FB5A85F4015">Ng, Peter K. L.</mods:namePart>
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<mods:name id="03C9F7FF40BE212FE1B94F74F9151B50" type="personal">
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<mods:namePart id="15128F9BEA2A096E88AFDAFDFB009A17">Forges, Bertrand Richer De</mods:namePart>
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<mods:typeOfResource id="B78EA8033735AC40A9FAA06D846E58EB">text</mods:typeOfResource>
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<mods:title id="C6FB489B4AA484119BF39F327481EB5D">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="3131BFAA8945491280CE47996641F85C">
<mods:date id="671E1E2A436AED3C5202CF297AEF6C5B">2015</mods:date>
<mods:detail id="E496B3FCD6BF1BB3D506462DC132CB44" type="pubDate">
<mods:number id="3C6F21B0D520430886EC1C20DD488393">2015-05-29</mods:number>
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<mods:detail id="D941AE0CE997B73C5BE60435FE9CB510" type="volume">
<mods:number id="7CDBF4C93EE472AE0CD3D844B2C0418F">63</mods:number>
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<mods:start id="E4C211BB5317938E4F2BA02994D969B3">110</mods:start>
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<mods:classification id="F9007A85EF7B3B671990C32091A89F5C">journal article</mods:classification>
<mods:identifier id="E8186D3F1AB9B92CF3AFD25007A621BE" type="DOI">10.5281/zenodo.5384590</mods:identifier>
<mods:identifier id="1F0B86720B987EA201F1AE57637B5D66" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03A9654BFFAC077554EAFE9E7CA3FD6B" ID-DOI="http://doi.org/10.5281/zenodo.5466803" ID-Zenodo-Dep="5466803" LSID="urn:lsid:plazi:treatment:03A9654BFFAC077554EAFE9E7CA3FD6B" httpUri="http://treatment.plazi.org/id/03A9654BFFAC077554EAFE9E7CA3FD6B" lastPageNumber="155" pageId="45" pageNumber="155">
<subSubSection id="C31A87D6FFAC077554EAFE9E7C18FE96" box="[909,1402,285,309]" pageId="45" pageNumber="155" type="nomenclature">
<paragraph id="8BBFD45DFFAC077554EAFE9E7C18FE96" blockId="45.[909,1402,285,309]" box="[909,1402,285,309]" pageId="45" pageNumber="155">
<heading id="D0F76331FFAC077554EAFE9E7C18FE96" bold="true" box="[909,1402,285,309]" centered="true" fontSize="10" level="2" pageId="45" pageNumber="155" reason="2">
<emphasis id="B974084FFFAC077554EAFE9E7C18FE96" bold="true" box="[909,1402,285,309]" pageId="45" pageNumber="155">
Key to species of
<taxonomicName id="4C00AFDEFFAC0775533FFE9D7C18FE96" ID-CoL="7PRYY" authority="De Haan, 1837" authorityName="De Haan" authorityYear="1837" box="[1112,1402,285,309]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFAC0775533FFE9D7DABFE96" bold="true" box="[1112,1225,286,309]" italics="true" pageId="45" pageNumber="155">Paramaya</emphasis>
De Haan, 1837
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFAC07755424FEDA7CA3FD6B" pageId="45" pageNumber="155" type="key">
<keyStep id="30F4CCF8FFAC07755424FEDA7CA3FE5D" pageId="45" pageNumber="155">
<paragraph id="8BBFD45DFFAC07755424FEDA7CA3FE66" blockId="45.[835,1475,345,511]" pageId="45" pageNumber="155">
<keyLead id="30F17768FFAC07755424FEDA7CA3FE66" pageId="45" pageNumber="155">
1. Pseudorostral horns relatively short; ambulatory meri in adults relatively stout, short (
<figureCitation id="133BC8D8FFAC0775532AFEF67DAAFE2F" box="[1101,1224,373,396]" captionStart="Fig" captionStartId="47.[136,171,1617,1639]" captionTargetBox="[136,1473,190,1599]" captionTargetId="figure-202@47.[136,1473,190,1599]" captionTargetPageId="47" captionText="Fig. 21. General habitus, Paramaya species. A, P. spinigera (as Maja (Paramaya) spinigera, after De Haan, 1837: pl. 24, fig. 4); B, P. spinigera, dried female (70.1 × 58.8 mm) (KPM NH4195), Japan; C, P. spinigera, male (64.7 × 51.2 mm) (NHM 1961.6.5.117), Japan; D, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; E, P. ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384638" httpUri="https://zenodo.org/record/5384638/files/figure.png" pageId="45" pageNumber="155">Fig. 21A–D</figureCitation>
); G1 gently curved (
<figureCitation id="133BC8D8FFAC077552FCFEF67AD0FE0B" captionStart="Fig" captionStartId="49.[135,170,1387,1409]" captionTargetBox="[260,1351,191,1371]" captionTargetId="figure-8@49.[252,1356,186,1374]" captionTargetPageId="49" captionText="Fig. 23. Left G1s, Paramaya species. A–C, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; D–F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; G–I, P. coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu. Scale bars = 5.0 mm [A, D, G]; 1.0 mm [B, C, E, F, H, I]." figureDoi="http://doi.org/10.5281/zenodo.5384642" httpUri="https://zenodo.org/record/5384642/files/figure.png" pageId="45" pageNumber="155">Fig. 23A–C</figureCitation>
);
<collectingCountry id="F31794CDFFAC077554A2FE107A9EFE0B" box="[965,1020,403,424]" name="Japan" pageId="45" pageNumber="155">Japan</collectingCountry>
and
<collectingCountry id="F31794CDFFAC07755349FE117D14FE0B" box="[1070,1142,402,424]" name="Taiwan" pageId="45" pageNumber="155">Taiwan</collectingCountry>
...................................................... .....................................
<taxonomicName id="4C00AFDEFFAC07755329FE337CA3FE66" ID-CoL="4D2M7" authority="(De Haan, 1837)" baseAuthorityName="De Haan" baseAuthorityYear="1837" box="[1102,1473,431,453]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="155" phylum="Arthropoda" rank="species" species="spinigera">
<emphasis id="B974084FFFAC07755329FE337C77FE66" box="[1102,1301,432,453]" italics="true" pageId="45" pageNumber="155">Paramaya spinigera</emphasis>
(De Haan, 1837)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFAC07755424FE4F7CA3FE5D" blockId="45.[835,1475,345,511]" pageId="45" pageNumber="155">
<keyLead id="30F17768FFAC07755424FE4F7CA3FE5D" pageId="45" pageNumber="155">
– Pseudorostral horns very long; ambulatory meri in adults relatively slender, long (
<figureCitation id="133BC8D8FFAC0775533FFE6A7DB6FE5D" box="[1112,1236,489,511]" captionStart="Fig" captionStartId="47.[136,171,1617,1639]" captionTargetBox="[136,1473,190,1599]" captionTargetId="figure-202@47.[136,1473,190,1599]" captionTargetPageId="47" captionText="Fig. 21. General habitus, Paramaya species. A, P. spinigera (as Maja (Paramaya) spinigera, after De Haan, 1837: pl. 24, fig. 4); B, P. spinigera, dried female (70.1 × 58.8 mm) (KPM NH4195), Japan; C, P. spinigera, male (64.7 × 51.2 mm) (NHM 1961.6.5.117), Japan; D, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; E, P. ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384638" httpUri="https://zenodo.org/record/5384638/files/figure.png" pageId="45" pageNumber="155">Figs. 21E, F</figureCitation>
,
<figureCitation id="133BC8D8FFAC07755387FE6A7C6FFE5C" box="[1248,1293,489,511]" captionStart="Fig" captionStartId="48.[113,148,1157,1179]" captionTargetBox="[113,1443,191,1130]" captionTargetId="figure-15@48.[113,1451,191,1144]" captionTargetPageId="48" captionText="Fig. 22. General habitus. A, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; B, “Maja spinigera” (ca. 70 mm carapace length), Sri Lanka, not preserved (photograph: R. Pethiyagoda); C, D, “Maja spinigera” (after Alcock &amp; Anderson, 1898: pl. 34, fig. 3)." figureDoi="http://doi.org/10.5281/zenodo.5384640" httpUri="https://zenodo.org/record/5384640/files/figure.png" pageId="45" pageNumber="155">22A</figureCitation>
); G1 otherwise ..2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F4CCF8FFAC07755424FDA17CA3FD6B" pageId="45" pageNumber="155">
<paragraph id="8BBFD45DFFAC07755424FDA17CA3FDD2" blockId="45.[835,1474,546,712]" pageId="45" pageNumber="155">
<keyLead id="30F17768FFAC07755424FDA17CA3FDD2" pageId="45" pageNumber="155">
2. Surface of thoracic sternum relatively smooth, with very low rounded granules (
<figureCitation id="133BC8D8FFAC07755347FDBC7D18FDF6" box="[1056,1146,575,597]" captionStart="Fig" captionStartId="86.[113,148,1750,1772]" captionTargetBox="[113,1452,388,1735]" captionTargetId="figure-138@86.[113,1452,388,1735]" captionTargetPageId="86" captionText="Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea." figureDoi="http://doi.org/10.5281/zenodo.5384702" httpUri="https://zenodo.org/record/5384702/files/figure.png" pageId="45" pageNumber="155">Fig. 50C</figureCitation>
); G1 gently curved (
<figureCitation id="133BC8D8FFAC07755220FDBC7CD5FDF6" box="[1351,1463,575,597]" captionStart="Fig" captionStartId="49.[135,170,1387,1409]" captionTargetBox="[260,1351,191,1371]" captionTargetId="figure-8@49.[252,1356,186,1374]" captionTargetPageId="49" captionText="Fig. 23. Left G1s, Paramaya species. A–C, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; D–F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; G–I, P. coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu. Scale bars = 5.0 mm [A, D, G]; 1.0 mm [B, C, E, F, H, I]." figureDoi="http://doi.org/10.5281/zenodo.5384642" httpUri="https://zenodo.org/record/5384642/files/figure.png" pageId="45" pageNumber="155">Fig. 23G–I</figureCitation>
);
<collectingCountry id="F31794CDFFAC0775540CFDDF7AD9FDD2" box="[875,955,604,625]" name="Vanuatu" pageId="45" pageNumber="155">Vanuatu</collectingCountry>
...........................................
<taxonomicName id="4C00AFDEFFAC077553A4FDDE7CE1FDD1" ID-CoL="4D2M5" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1219,1411,605,626]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="155" phylum="Arthropoda" rank="species" species="coccinea" status="sp. nov.">
<emphasis id="B974084FFFAC077553A4FDDE7CE1FDD1" box="[1219,1411,605,626]" italics="true" pageId="45" pageNumber="155">Paramaya coccinea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFAC077552ECFDDF7CA3FDD2" box="[1419,1473,604,625]" pageId="45" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFAC07755424FDFA7CA3FD6B" blockId="45.[835,1474,546,712]" pageId="45" pageNumber="155">
<keyLead id="30F17768FFAC07755424FDFA7CA3FD6B" pageId="45" pageNumber="155">
– Surface of thoracic sternum distinctly granulated (
<figureCitation id="133BC8D8FFAC0775523BFDFA7CD4FD2C" box="[1372,1462,633,655]" captionStart="Fig" captionStartId="86.[113,148,1750,1772]" captionTargetBox="[113,1452,388,1735]" captionTargetId="figure-138@86.[113,1452,388,1735]" captionTargetPageId="86" captionText="Fig. 50. Male anterior thoracic sternums and abdomens. A, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; B, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; C, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu; D, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; E, Holthuija miersii, male (22.3 × 16.5 mm) (CBM ZC4001), Singapore; F, Holthuija suluensis, male (11.5 × 8.2 mm) (USNM 48507), Philippines; G, Holthuija pauli n. sp., holotype male (37.3 × 28.0 mm) (NMCR), Philippines; H, Holthuija cognata n. sp., holotype male (29.5 × 23.7 mm) (CBM-ZC3662), Japan; I, Holthuija poorei n. sp., holotype male (24.3 × 18.7 mm) (NMV J63749), Timor Sea." figureDoi="http://doi.org/10.5281/zenodo.5384702" httpUri="https://zenodo.org/record/5384702/files/figure.png" pageId="45" pageNumber="155">Fig. 50B</figureCitation>
); G1 more strongly curved especially distally (
<figureCitation id="133BC8D8FFAC0775525AFD167CD5FD0F" box="[1341,1463,661,684]" captionStart="Fig" captionStartId="49.[135,170,1387,1409]" captionTargetBox="[260,1351,191,1371]" captionTargetId="figure-8@49.[252,1356,186,1374]" captionTargetPageId="49" captionText="Fig. 23. Left G1s, Paramaya species. A–C, P. spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; D–F, P. ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; G–I, P. coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu. Scale bars = 5.0 mm [A, D, G]; 1.0 mm [B, C, E, F, H, I]." figureDoi="http://doi.org/10.5281/zenodo.5384642" httpUri="https://zenodo.org/record/5384642/files/figure.png" pageId="45" pageNumber="155">Fig. 23D–F</figureCitation>
);
<collectingCountry id="F31794CDFFAC0775540EFD317AB7FD6B" box="[873,981,690,712]" name="Philippines" pageId="45" pageNumber="155">Philippines</collectingCountry>
.............................................
<taxonomicName id="4C00AFDEFFAC0775538EFD307CE6FD64" ID-CoL="4D2M6" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1257,1412,690,712]" class="Malacostraca" family="Majidae" genus="Paramaya" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="155" phylum="Arthropoda" rank="species" species="ouch" status="sp. nov.">
<emphasis id="B974084FFFAC0775538EFD307CE6FD64" box="[1257,1412,690,712]" italics="true" pageId="45" pageNumber="155">Paramaya ouch</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFAC077552ECFD307CA3FD6B" box="[1419,1473,691,712]" pageId="45" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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