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<document id="4A68289CCA85F9C9CAE9B6EF2087DE5E" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FB5EFE94CBAFBBC3FB79FBD7" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Syntomodrillia Woodring 1928" docType="treatment" docVersion="12" lastPageNumber="317" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="316" updateTime="1698184378282" updateUser="plazi">
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<mods:title id="DDCD886E1BED4B94A25B4D12B0BC311C">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="6589AD1C9ED0C219CACEF56A615A8E25">Fallon, Phillip J.</mods:namePart>
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<mods:date id="2C3427181647C99D77808BAB4D5B2E21">2016</mods:date>
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<mods:number id="7D4113F7BFB010F376E891F15478698A">4090</mods:number>
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<treatment id="039F87C4FB5EFE94CBAFBBC3FB79FBD7" ID-DOI="http://doi.org/10.5281/zenodo.6076633" ID-GBIF-Taxon="117911576" ID-Zenodo-Dep="6076633" LSID="urn:lsid:plazi:treatment:039F87C4FB5EFE94CBAFBBC3FB79FBD7" httpUri="http://treatment.plazi.org/id/039F87C4FB5EFE94CBAFBBC3FB79FBD7" lastPageId="316" lastPageNumber="317" pageId="315" pageNumber="316">
<subSubSection id="C32C6559FB5EFE93CBAFBBC3FC91FA13" pageId="315" pageNumber="316" type="nomenclature">
<paragraph id="8B8936D2FB5EFE93CBAFBBC3FD98FD58" blockId="315.[151,624,727,754]" box="[151,624,727,754]" pageId="315" pageNumber="316">
<heading id="D0C181BEFB5EFE93CBAFBBC3FD98FD58" bold="true" box="[151,624,727,754]" fontSize="11" level="1" pageId="315" pageNumber="316" reason="1">
<emphasis id="B942EAC0FB5EFE93CBAFBBC3FD98FD58" bold="true" box="[151,624,727,754]" pageId="315" pageNumber="316">
Genus
<taxonomicName id="4C364D51FB5EFE93CBD6BBCCFD98FD58" ID-CoL="7PHWK" authority="Woodring, 1928" authorityName="Woodring" authorityYear="1928" box="[238,624,727,754]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93CBD6BBCCFE49FD5B" bold="true" box="[238,417,727,753]" italics="true" pageId="315" pageNumber="316">Syntomodrillia</emphasis>
Woodring, 1928
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936D2FB5EFE93CBAFBA04FAA3FC9E" blockId="315.[151,1355,798,820]" box="[151,1355,798,820]" pageId="315" pageNumber="316">
<typeStatus id="548D8870FB5EFE93CBAFBA04FF21FC9E" box="[151,201,799,820]" pageId="315" pageNumber="316">Type</typeStatus>
species:
<taxonomicName id="4C364D51FB5EFE93CA1EBA05FDB4FC9E" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[294,604,798,820]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93CA1EBA05FE32FC9E" box="[294,474,798,820]" italics="true" pageId="315" pageNumber="316">Drillia lissotropis</emphasis>
(Dall, 1881)
</taxonomicName>
by original designation (Recent, off Bahia Honda, Artemisa Prov.,
<collectingCountry id="F3217642FB5EFE93CE3EBA05FAA8FC9E" box="[1286,1344,798,820]" name="Cuba" pageId="315" pageNumber="316">Cuba</collectingCountry>
).
</paragraph>
<paragraph id="8B8936D2FB5EFE93CBAFBA7AFC91FA13" blockId="315.[151,1436,865,1826]" pageId="315" pageNumber="316">
Woodring named
<taxonomicName id="4C364D51FB5EFE93CA65BA7AFCE5FCD0" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" box="[349,781,865,890]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93CA65BA7AFD96FCD0" box="[349,638,865,890]" italics="true" pageId="315" pageNumber="316">Syntomodrillia lissotropis</emphasis>
(Dall, 1881)
</taxonomicName>
as the
<typeStatus id="548D8870FB5EFE93C861BA79FC61FCD3" box="[857,905,866,889]" pageId="315" pageNumber="316">type</typeStatus>
of the genus without designating a specimen or lot as genus
<typeStatus id="548D8870FB5EFE93CA1FBA9DFEBFFC37" box="[295,343,902,925]" pageId="315" pageNumber="316">type</typeStatus>
despite Dall having identified six lots from three different localities as this species (Dall, 1881: 58–59; Dall, 1889a: 92) (all listed in the section about
<taxonomicName id="4C364D51FB5EFE93C9C4BAB1FC64FC68" box="[764,908,937,962]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93C9C4BAB1FC64FC68" box="[764,908,937,962]" italics="true" pageId="315" pageNumber="316">S. lissotropis</emphasis>
</taxonomicName>
below). To accommodate the specimens of the new lots, Dall emended the description in 1889 to variably include intercostal spiral grooves, which was not mentioned in the original description. Bartsch (1934: 25–26), discovered that Dall’s lots of
<taxonomicName id="4C364D51FB5EFE93CFDDBAEAFAB1FBA0" box="[1253,1369,1009,1034]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93CFDDBAEAFAB1FBA0" box="[1253,1369,1009,1034]" italics="true" pageId="315" pageNumber="316">lissotropis</emphasis>
</taxonomicName>
were comprised of more than one species (five different ones are described in this work). Because Woodring had not specified a particular specimen as
<typeStatus id="548D8870FB5EFE93C916BD21FDB6FBFB" box="[558,606,1082,1105]" pageId="315" pageNumber="316">type</typeStatus>
of the genus, Bartsch selected a specimen that had distinct intercostal grooves from one of Dall’s
<taxonomicName id="4C364D51FB5EFE93CAE7BD46FDBBFBDC" box="[479,595,1117,1142]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93CAE7BD46FDBBFBDC" box="[479,595,1117,1142]" italics="true" pageId="315" pageNumber="316">lissotropis</emphasis>
</taxonomicName>
lots (USNM 87477), which he believed to be representative of the one Woodring intended as
<typeStatus id="548D8870FB5EFE93CAABBD99FE2BFB33" box="[403,451,1154,1177]" pageId="315" pageNumber="316">type</typeStatus>
of
<taxonomicName id="4C364D51FB5EFE93CAD1BD9AFD67FB30" box="[489,655,1153,1178]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93CAD1BD9AFD67FB30" box="[489,655,1153,1178]" italics="true" pageId="315" pageNumber="316">Syntomodrillia</emphasis>
</taxonomicName>
. He named it
<taxonomicName id="4C364D51FB5EFE93C816BD9AFBA7FB30" box="[814,1103,1153,1178]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="woodringi">
<emphasis id="B942EAC0FB5EFE93C816BD9AFBA7FB30" box="[814,1103,1153,1178]" italics="true" pageId="315" pageNumber="316">Syntomodrillia woodringi</emphasis>
</taxonomicName>
, and substituted it as the
<typeStatus id="548D8870FB5EFE93CE54BD99FA74FB33" box="[1388,1436,1154,1177]" pageId="315" pageNumber="316">type</typeStatus>
of the genus. Bartsch believed it to be a different species because Dall’s original 1881 description did not mention intercostal spirals. Woodring (1970: 388–389) later rebutted Bartsch’s action by stating that the presence of spirals was a variable trait, that whether or not
<taxonomicName id="4C364D51FB5EFE93C96EBDF5FD0FFAAC" box="[598,743,1261,1286]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93C96EBDF5FD0FFAAC" box="[598,743,1261,1286]" italics="true" pageId="315" pageNumber="316">S. lissotropis</emphasis>
</taxonomicName>
possessed intercostal spirals was unimportant, citing as proof a fossil species that exhibited variability in this trait, and put
<taxonomicName id="4C364D51FB5EFE93C87BBC09FC3AFA80" box="[835,978,1297,1322]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="woodringi">
<emphasis id="B942EAC0FB5EFE93C87BBC09FC3AFA80" box="[835,978,1297,1322]" italics="true" pageId="315" pageNumber="316">S. woodringi</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName id="4C364D51FB5EFE93CF94BC09FAD4FA80" box="[1196,1340,1297,1322]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93CF94BC09FAD4FA80" box="[1196,1340,1297,1322]" italics="true" pageId="315" pageNumber="316">S. lissotropis</emphasis>
</taxonomicName>
, leaving the latter as the
<typeStatus id="548D8870FB5EFE93CA7EBC2DFE9EFAE7" box="[326,374,1334,1357]" pageId="315" pageNumber="316">type</typeStatus>
of the genus.
<taxonomicName id="4C364D51FB5EFE93C92ABC2EFCDAFAE4" box="[530,818,1333,1358]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93C92ABC2EFCDAFAE4" box="[530,818,1333,1358]" italics="true" pageId="315" pageNumber="316">Syntomodrillia lissotropis</emphasis>
</taxonomicName>
is supported in this work as the valid
<typeStatus id="548D8870FB5EFE93CFEEBC2DFAEEFAE7" box="[1238,1286,1334,1357]" pageId="315" pageNumber="316">type</typeStatus>
of the genus, as intended by Woodring. Bartsch’s action does not fulfill the requirements of ICZN 2000 Article 70 (the
<typeStatus id="548D8870FB5EFE93CE54BC41FA74FADB" box="[1388,1436,1370,1393]" pageId="315" pageNumber="316">type</typeStatus>
species was not misidentified so substituting another is not allowed).
<taxonomicName id="4C364D51FB5EFE93C89CBC66FB4BFA3C" box="[932,1187,1405,1430]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="woodringi">
<emphasis id="B942EAC0FB5EFE93C89CBC66FB4BFA3C" box="[932,1187,1405,1430]" italics="true" pageId="315" pageNumber="316">Splendrillia woodringi</emphasis>
</taxonomicName>
is shown in this work to be a different but valid species, not a synonym of
<taxonomicName id="4C364D51FB5EFE93C9DABCB9FC9AFA10" box="[738,882,1441,1466]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="species" species="lissotropis">
<emphasis id="B942EAC0FB5EFE93C9DABCB9FC9AFA10" box="[738,882,1441,1466]" italics="true" pageId="315" pageNumber="316">S. lissotropis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB5EFE93CBFFBCDEFF14F970" pageId="315" pageNumber="316" type="diagnosis">
<paragraph id="8B8936D2FB5EFE93CBFFBCDEFF14F970" blockId="315.[151,1436,865,1826]" pageId="315" pageNumber="316">
<emphasis id="B942EAC0FB5EFE93CBFFBCDEFEA9FA74" bold="true" box="[199,321,1477,1502]" pageId="315" pageNumber="316">Diagnosis.</emphasis>
According to Woodring (1928: 160), very small slender shell, with numerous ribs, a varix some distance from the outer lip, a short, relatively wide aperture, and short anterior canal with a shallow siphonal notch; fasciole not swollen. Nucleus slender with about 2 smooth whorls. Sculpture of broad axial ribs extending from suture-to-suture, reduced on sulcus. Spiral sculpture of microscopic threads or grooves between ribs on the later whorls and of fine threads on the anterior fasciole. Intercostal spirals may be absent in some species, or variably present (Woodring, 1970: 388–389). Anal sinus broad, with a round apex that adjoins the suture. Outer lip with a shallow stromboid notch. Inner lip narrow, edge detached; a weak parietal callus located posteriorly at junction of outer lip.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB5EFE93CBFFBFFDFCAAF87C" pageId="315" pageNumber="316" type="key">
<paragraph id="8B8936D2FB5EFE93CBFFBFFDFC32F88B" blockId="315.[151,1436,865,1826]" pageId="315" pageNumber="316">
<emphasis id="B942EAC0FB5EFE93CBFFBFFDFE28F954" bold="true" box="[199,448,1765,1790]" pageId="315" pageNumber="316">Key characteristics.</emphasis>
The presence of all the following characteristics is diagnostic of species of
<taxonomicName id="4C364D51FB5EFE93CBAFBE12FED6F888" ID-CoL="7PHWK" box="[151,318,1801,1826]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93CBAFBE12FED6F888" box="[151,318,1801,1826]" italics="true" pageId="315" pageNumber="316">Syntomodrillia</emphasis>
</taxonomicName>
and separates the genus from other TWA
<taxonomicName id="4C364D51FB5EFE93C82DBE12FC68F888" ID-CoL="7NHDF" box="[789,896,1801,1826]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
genera:
</paragraph>
<paragraph id="8B8936D2FB5EFE93CBAFBE49FCAAF87C" blockId="315.[151,1436,1873,2006]" pageId="315" pageNumber="316">
1. Shell small, slender, glossy, with a moderately long anterior canal. (Although characterized as short by Woodring, the canal is usually longer than in some
<taxonomicName id="4C364D51FB5EFE93C87BBE6DFC55F827" ID-CoL="7P2D7" box="[835,957,1910,1933]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93C87BBE6DFC55F827" box="[835,957,1910,1933]" italics="true" pageId="315" pageNumber="316">Fenimorea</emphasis>
</taxonomicName>
or
<taxonomicName id="4C364D51FB5EFE93C8D7BE6EFB9BF824" ID-CoL="7PHCB" box="[1007,1139,1909,1934]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93C8D7BE6EFB9BF824" box="[1007,1139,1909,1934]" italics="true" pageId="315" pageNumber="316">Splendrillia</emphasis>
</taxonomicName>
, for instance, which are described as short in this work. Relatively speaking, the canal of
<taxonomicName id="4C364D51FB5EFE93C8A7BE82FBAEF818" ID-CoL="7PHWK" box="[927,1094,1945,1970]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93C8A7BE82FBAEF818" box="[927,1094,1945,1970]" italics="true" pageId="315" pageNumber="316">Syntomodrillia</emphasis>
</taxonomicName>
is moderately long but shorter than that of
<taxonomicName id="4C364D51FB5EFE93CA75BEA6FE3BF87C" ID-CoL="7P77Q" box="[333,467,1981,2006]" class="Gastropoda" family="Drilliidae" genus="Leptadrillia" kingdom="Animalia" order="Neogastropoda" pageId="315" pageNumber="316" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB5EFE93CA75BEA6FE3BF87C" box="[333,467,1981,2006]" italics="true" pageId="315" pageNumber="316">Leptadrillia</emphasis>
</taxonomicName>
, which is characterized as long.)
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB59FE94CBAFB983FEBCFC62" pageId="316" pageNumber="317" type="description">
<paragraph id="8B8936D2FB59FE94CBAFB983FD98FF7F" blockId="316.[151,1436,151,357]" pageId="316" pageNumber="317">2. Axial ribs that extend from suture-to-suture on spire whorls, to fasciole on last, slightly reduced in height, and recurved on shoulders of later whorls;</paragraph>
<paragraph id="8B8936D2FB59FE94CBAFB9FBFE15FEEA" blockId="316.[151,1436,151,357]" pageId="316" pageNumber="317">3. Spiral sculpture of fine threads or grooves, usually below the periphery of later whorls (absent on shoulder), stronger on shell base, and of ridges on the anterior fasciole. Spiral sculpture may be very faint or variably absent in some species; and</paragraph>
<paragraph id="8B8936D2FB59FE94CBAFB857FB57FECF" blockId="316.[151,1436,151,357]" box="[151,1215,332,357]" pageId="316" pageNumber="317">4. A varix (usually not hump-like) approximately ¼- to ⅓-turn from the edge of the outer lip.</paragraph>
<paragraph id="8B8936D2FB59FE94CBFFB88EFB18FE5F" blockId="316.[151,1437,404,1149]" pageId="316" pageNumber="317">
Most species in the genus are similar-looking and difficult to distinguish, differing only slightly in shell and whorl profile, size, rib form, and a few other other characteristics, attributes that are not always evident in imperfect specimens. Locality is important because ranges of most
<taxonomicName id="4C364D51FB59FE94C8B1B8C7FBC7FE5F" box="[905,1071,476,501]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C8B1B8C7FBC7FE5F" box="[905,1071,476,501]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
are very limited.
</paragraph>
<paragraph id="8B8936D2FB59FE94CBFFBB1BFB6EFD2F" blockId="316.[151,1437,404,1149]" pageId="316" pageNumber="317">
<emphasis id="B942EAC0FB59FE94CBFFBB1BFE21FDBD" bold="true" box="[199,457,512,535]" pageId="316" pageNumber="317">Nomenclatural notes.</emphasis>
There has been a gradual suppression of the use of
<taxonomicName id="4C364D51FB59FE94CF16B8E4FB3CFDB2" box="[1070,1236,511,536]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CF16B8E4FB3CFDB2" box="[1070,1236,511,536]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
as a genus since Powell (1966) reduced it to a subgenus of
<taxonomicName id="4C364D51FB59FE94C954BB3FFD18FD97" box="[620,752,548,573]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C954BB3FFD18FD97" box="[620,752,548,573]" italics="true" pageId="316" pageNumber="317">Splendrillia</emphasis>
</taxonomicName>
. The subgenus has been dropped altogether by Turgeon
<emphasis id="B942EAC0FB59FE94CE5CBB3EFA75FD97" box="[1380,1437,548,573]" italics="true" pageId="316" pageNumber="317">et al.</emphasis>
(1998). However, species in
<taxonomicName id="4C364D51FB59FE94CAC9BB5CFD70FDCA" box="[497,664,583,608]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CAC9BB5CFD70FDCA" box="[497,664,583,608]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
are sufficiently different morphologically from
<taxonomicName id="4C364D51FB59FE94CFE0BB5CFAB4FDCA" box="[1240,1372,583,608]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CFE0BB5CFAB4FDCA" box="[1240,1372,583,608]" italics="true" pageId="316" pageNumber="317">Splendrillia</emphasis>
</taxonomicName>
(and
<taxonomicName id="4C364D51FB59FE94CBAFBB77FEF5FD2F" box="[151,285,620,645]" class="Gastropoda" family="Drilliidae" genus="Leptadrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CBAFBB77FEF5FD2F" box="[151,285,620,645]" italics="true" pageId="316" pageNumber="317">Leptadrillia</emphasis>
</taxonomicName>
) to merit separation at the genus level (see discussion of similar genera next).
</paragraph>
<paragraph id="8B8936D2FB59FE94CBFFBB94FC8FFD67" blockId="316.[151,1437,404,1149]" pageId="316" pageNumber="317">
The identification of species of
<taxonomicName id="4C364D51FB59FE94C90BBB94FD32FD02" box="[563,730,655,680]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C90BBB94FD32FD02" box="[563,730,655,680]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
has been greatly confused on account of the poor quality and description of most
<typeStatus id="548D8870FB59FE94CA4EBBAEFE59FD66" box="[374,433,693,716]" pageId="316" pageNumber="317">types</typeStatus>
and similarity of species to each other.
</paragraph>
<paragraph id="8B8936D2FB59FE94CBFFBBCCFEBCFC62" blockId="316.[151,1437,404,1149]" pageId="316" pageNumber="317">
<emphasis id="B942EAC0FB59FE94CBFFBBCCFE97FD45" bold="true" box="[199,383,727,752]" pageId="316" pageNumber="317">Similar genera.</emphasis>
<taxonomicName id="4C364D51FB59FE94CABDBBCCFD20FD5A" authority="Woodring, 1928" authorityName="Woodring" authorityYear="1928" box="[389,712,727,752]" class="Gastropoda" family="Drilliidae" genus="Leptadrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CABDBBCCFDE3FD5A" box="[389,523,727,752]" italics="true" pageId="316" pageNumber="317">Leptadrillia</emphasis>
Woodring, 1928
</taxonomicName>
is most similar to but differs from
<taxonomicName id="4C364D51FB59FE94CF69BBCCFB1FFD5A" box="[1105,1271,727,752]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94CF69BBCCFB1FFD5A" box="[1105,1271,727,752]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
in that it has a long anterior canal and lacks intercostal spiral grooves.
<taxonomicName id="4C364D51FB59FE94C822BBE7FBD6FCBE" authority="Hedley, 1922" authorityName="Hedley" authorityYear="1922" box="[794,1086,764,789]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C822BBE7FC77FCBF" box="[794,927,764,789]" italics="true" pageId="316" pageNumber="317">Splendrillia</emphasis>
Hedley, 1922
</taxonomicName>
differs in the following ways: axial ribs end at the sulcus, not at the suture; ribs are shorter, usually more oblique, evanescing just below the whorl periphery and rarely reaching the anterior fasciole; anterior canal is shorter; shell surface has a microsculpture of spiral lines; and the varix is hump-like.
<taxonomicName id="4C364D51FB59FE94C96DBA7CFC06FC2A" authority="Bartsch &amp; Rehder, 1939" authorityName="Bartsch &amp; Rehder" authorityYear="1939" box="[597,1006,871,896]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C96DBA7CFD3EFC2A" box="[597,726,871,896]" italics="true" pageId="316" pageNumber="317">Lissodrillia</emphasis>
Bartsch &amp; Rehder, 1939
</taxonomicName>
also has a small glossy shell with ribs that extend from suture-to-suture in some species, but lacks spiral sculpture altogether, most noticeably on the anterior fasciole.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FB59FE94CBFFBACFFB79FBD7" pageId="316" pageNumber="317" type="distribution">
<paragraph id="8B8936D2FB59FE94CBFFBACFFB79FBD7" blockId="316.[151,1437,404,1149]" pageId="316" pageNumber="317">
<emphasis id="B942EAC0FB59FE94CBFFBACFFE88FC47" bold="true" box="[199,352,980,1005]" pageId="316" pageNumber="317">Distribution.</emphasis>
Members of the genus
<taxonomicName id="4C364D51FB59FE94C9B7BACFFCDEFC47" box="[655,822,980,1005]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="316" pageNumber="317" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FB59FE94C9B7BACFFCDEFC47" box="[655,822,980,1005]" italics="true" pageId="316" pageNumber="317">Syntomodrillia</emphasis>
</taxonomicName>
are largely offshore, soft bottom dwellers, widely distributed throughout the region, though species are individually limited in their range. The genus has apparently undergone a significant radiation based on the number of species discovered to-date, and it is likely that many more are yet to be discovered, especially in areas that have not been thoroughly explored. Because of their habitat preference, and small size, they are easily overlooked and so are uncommon in collections.
</paragraph>
</subSubSection>
</treatment>
</document>
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