182 lines
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182 lines
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<document id="8D1F6C1005EDB5E0537DDCF846DEC097" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA4BFF86CBAFB983FB63FA3A" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calliclava McLean 1971" docType="treatment" docVersion="12" lastPageNumber="47" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="47" updateTime="1698184378282" updateUser="plazi">
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<mods:titleInfo id="CB1BFDA6531D2971AEB019369B9B86C7">
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<mods:title id="E0BECB62A323BD8BA4ED0AA51D58890D">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="E95F12F36AB32F314FAC59CAD5626716">Fallon, Phillip J.</mods:namePart>
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<mods:typeOfResource id="A83061353CE9C527F25EB42A5C45991F">text</mods:typeOfResource>
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<mods:title id="91FA224A7AE533A26588402EC741B525">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="350A25220EEDE41E678947784266F480">
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<mods:date id="FFB92F02302BEEE242F2259DEE37CFA2">2016</mods:date>
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<mods:detail id="242B1C70F2582BF78E86CA46B67A2F26" type="volume">
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<mods:number id="187393F73982DC1A599A1A90A068B0E6">4090</mods:number>
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<mods:number id="E2CEC57D581C85263A53C7DEF02CE4FC">1</mods:number>
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<mods:start id="1EE16D0691D41D05046ED874A411D357">1</mods:start>
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<mods:classification id="9BC7FFAA3A975F6D2AFA8A82D03907B4">journal article</mods:classification>
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<mods:identifier id="A184CE62A275F9EAB206BE7C18F33DD6" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<mods:identifier id="6FA3978C56BE382527F58B4AD09E140B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="45229D066879A83FB4C7EA0421779495" type="Zenodo-Dep">263299</mods:identifier>
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</mods:mods>
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<treatment id="039F87C4FA4BFF86CBAFB983FB63FA3A" ID-DOI="http://doi.org/10.5281/zenodo.6076279" ID-GBIF-Taxon="117911719" ID-Zenodo-Dep="6076279" LSID="urn:lsid:plazi:treatment:039F87C4FA4BFF86CBAFB983FB63FA3A" httpUri="http://treatment.plazi.org/id/039F87C4FA4BFF86CBAFB983FB63FA3A" lastPageNumber="47" pageId="46" pageNumber="47">
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<subSubSection id="C32C6559FA4BFF86CBAFB983FED5FEB9" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8B8936D2FA4BFF86CBAFB983FDC9FF18" blockId="46.[151,545,151,178]" box="[151,545,151,178]" pageId="46" pageNumber="47">
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<heading id="D0C181BEFA4BFF86CBAFB983FDC9FF18" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<emphasis id="B942EAC0FA4BFF86CBAFB983FDC9FF18" bold="true" box="[151,545,151,178]" pageId="46" pageNumber="47">
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Genus
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<taxonomicName id="4C364D51FA4BFF86CBD6B98CFDC9FF18" ID-CoL="7NVM5" authority="McLean, 1971" authorityName="McLean" authorityYear="1971" box="[238,545,151,178]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CBD6B98CFE81FF1B" bold="true" box="[238,361,151,177]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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McLean, 1971
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBAFB9C4FED5FEB9" blockId="46.[151,1436,222,275]" pageId="46" pageNumber="47">
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<typeStatus id="548D8870FA4BFF86CBAFB9C4FF21FF5E" box="[151,201,223,244]" pageId="46" pageNumber="47">Type</typeStatus>
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species:
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<taxonomicName id="4C364D51FA4BFF86CA17B9C5FD79FF5E" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[303,657,222,244]" class="Gastropoda" family="Drilliidae" genus="Cymatosyrinx" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="palmeri">
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<emphasis id="B942EAC0FA4BFF86CA17B9C5FDFDFF5E" box="[303,533,222,244]" italics="true" pageId="46" pageNumber="47">Cymatosyrinx palmeri</emphasis>
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Dall, 1919
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</taxonomicName>
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, by original designation (Recent, beach at the head of the Gulf, Gulf of California).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA4BFF86CBAFB824FE0DFE42" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="8B8936D2FA4BFF86CBAFB824FE0DFE42" blockId="46.[151,1437,319,560]" pageId="46" pageNumber="47">
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<emphasis id="B942EAC0FA4BFF86CBAFB824FEF9FEF2" bold="true" box="[151,273,319,344]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
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According to McLean (1971: 117), shell small to medium sized, surface glossy, body whorl short in comparison with the spire. Protoconch large, of two carinate whorls. Axials from suture-to-suture, but weak in sulcus. Spirals of incised grooves. Anal sinus on shoulder deep, U-shaped, bordered by a curved parietal callus. Outer lip nearly straight with a moderately deep stromboid notch; a varix is present ¼-turn from the edge of the lip. Anterior canal short, notched.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA4BFF86CBFFB8EEFEF1FD15" pageId="46" pageNumber="47" type="key">
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<paragraph id="8B8936D2FA4BFF86CBFFB8EEFD26FD9A" blockId="46.[151,1437,319,560]" pageId="46" pageNumber="47">
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<emphasis id="B942EAC0FA4BFF86CBFFB8EEFE54FDA7" bold="true" box="[199,444,500,525]" pageId="46" pageNumber="47">Key characteristics.</emphasis>
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The combination of the following characteristics is diagnostic of
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<taxonomicName id="4C364D51FA4BFF86CFD7B8EFFA8CFDA7" ID-CoL="7NVM5" box="[1263,1380,500,525]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CFD7B8EFFA8CFDA7" box="[1263,1380,500,525]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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</taxonomicName>
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and separates the genus from all other TWA
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<taxonomicName id="4C364D51FA4BFF86C963BB0CFD23FD9A" ID-CoL="7NHDF" box="[603,715,535,560]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
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:
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</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBAFBB7BFEF1FD15" blockId="46.[151,1437,607,813]" pageId="46" pageNumber="47">1. Axial ribs numerous, extend from suture-to-suture, depressed and recurved in the sulcus, and run to anterior fasciole on body whorl; ribs overridden by spiral grooves that are shallow on crests, deeper in intercostal spaces;</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA4BFF86CBAFBBD7FD8CFAC6" pageId="46" pageNumber="47" type="description">
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<paragraph id="8B8936D2FA4BFF86CBAFBBD7FC5BFD4F" blockId="46.[151,1437,607,813]" box="[151,947,716,741]" pageId="46" pageNumber="47">2. Protoconch of about 2–2¼ smooth, translucent, keeled whorls; and</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBAFBBEBFEE5FC87" blockId="46.[151,1437,607,813]" pageId="46" pageNumber="47">3. Varix broad and hump-like positioned about ⅓-turn from the edge of the outer lip; marked with a dark color patch.</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBFFBA46FD20FC4A" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
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<emphasis id="B942EAC0FA4BFF86CBFFBA46FE2EFCDE" bold="true" box="[199,454,861,884]" pageId="46" pageNumber="47">Nomenclatural notes.</emphasis>
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This genus was erected by McLean for species that otherwise resembled
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<taxonomicName id="4C364D51FA4BFF86CFC1BA47FA7DFCDF" box="[1273,1429,860,885]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CFC1BA47FA7DFCDF" box="[1273,1429,860,885]" italics="true" pageId="46" pageNumber="47">Clathrodrillia</emphasis>
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</taxonomicName>
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, but differ in possessing a protoconch that is carinate from the tip of the first whorl, and a radula with a compressed, not elongate, lateral tooth. Dall erected separate genera,
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<taxonomicName id="4C364D51FA4BFF86C825BABFFBF6FC16" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[797,1054,932,957]" class="Gastropoda" family="Drilliidae" genus="Elaeocyma" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86C825BABFFC72FC17" box="[797,922,932,957]" italics="true" pageId="46" pageNumber="47">Elaeocyma</emphasis>
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Dall, 1918
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</taxonomicName>
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; and
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<taxonomicName id="4C364D51FA4BFF86CF65BABFFAF2FC16" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[1117,1306,932,957]" class="Gastropoda" family="Drilliidae" genus="Kylix" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CF65BABFFB71FC17" box="[1117,1177,932,957]" italics="true" pageId="46" pageNumber="47">Kylix</emphasis>
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Dall, 1919
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</taxonomicName>
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for eastern Pacific drilliids with partially keeled protoconchs.
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</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBFFBAF7FAA6FBDA" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
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TWA species of
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<taxonomicName id="4C364D51FA4BFF86CAB9BAF7FE1EFBAF" box="[385,502,1004,1029]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CAB9BAF7FE1EFBAF" box="[385,502,1004,1029]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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</taxonomicName>
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are similar in appearance to
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<taxonomicName id="4C364D51FA4BFF86C802BAF7FB77FBAF" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[826,1183,1004,1029]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="palmeri">
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<emphasis id="B942EAC0FA4BFF86C802BAF7FBE5FBAF" box="[826,1037,1004,1029]" italics="true" pageId="46" pageNumber="47">Calliclava palmeri</emphasis>
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(Dall, 1919)
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</taxonomicName>
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, the
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<typeStatus id="548D8870FA4BFF86CFEDBAF6FAEDFBAE" box="[1237,1285,1005,1028]" pageId="46" pageNumber="47">type</typeStatus>
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of the genus. Another eastern Pacific species,
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<taxonomicName id="4C364D51FA4BFF86C92DBD14FBC5FB82" authority="McLean & Poorman, 1971" authorityName="McLean & Poorman" authorityYear="1971" box="[533,1069,1039,1064]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="subtilis">
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<emphasis id="B942EAC0FA4BFF86C92DBD14FD0EFB82" box="[533,742,1039,1064]" italics="true" pageId="46" pageNumber="47">Calliclava subtilis</emphasis>
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McLean & Poorman, 1971
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</taxonomicName>
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, is also similar. Ten species of
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<taxonomicName id="4C364D51FA4BFF86CBAFBD2FFEE4FBE7" box="[151,268,1076,1101]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86CBAFBD2FFEE4FBE7" box="[151,268,1076,1101]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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</taxonomicName>
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are listed in the on-line database, World Registry of Marine Species as of the date of this research, all from the eastern Pacific (WoRMS, 2016). The
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<taxonomicName id="4C364D51FA4BFF86C99BBD4CFCF0FBDA" box="[675,792,1111,1136]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86C99BBD4CFCF0FBDA" box="[675,792,1111,1136]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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</taxonomicName>
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herein are the first to be described from the TWA.
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</paragraph>
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<paragraph id="8B8936D2FA4BFF86CBFFBD67FAA8FB77" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
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<emphasis id="B942EAC0FA4BFF86CBFFBD67FE68FB3E" bold="true" box="[199,384,1148,1173]" pageId="46" pageNumber="47">Similar genera.</emphasis>
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The genus
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<taxonomicName id="4C364D51FA4BFF86C93CBD67FD91FB3F" box="[516,633,1148,1173]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86C93CBD67FD91FB3F" box="[516,633,1148,1173]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
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</taxonomicName>
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is most easily distinguished by its fully carinate protoconch. Although otherwise close in appearance, only the second protoconch whorl of
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<taxonomicName id="4C364D51FA4BFF86C8FCBD84FB2FFB12" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[964,1223,1183,1208]" class="Gastropoda" family="Drilliidae" genus="Elaeocyma" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86C8FCBD84FBA9FB12" box="[964,1089,1183,1208]" italics="true" pageId="46" pageNumber="47">Elaeocyma</emphasis>
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Dall, 1918
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</taxonomicName>
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is carinate. Other drilliid genera with carinate protoconchs include
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<taxonomicName id="4C364D51FA4BFF86C985BDDFFCCDFB77" box="[701,805,1220,1245]" class="Gastropoda" family="Drilliidae" genus="Imaclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA4BFF86C985BDDFFCCDFB77" box="[701,805,1220,1245]" italics="true" pageId="46" pageNumber="47">Imaclava</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C364D51FA4BFF86C85BBDDFFC76FB77" box="[867,926,1220,1245]" class="Gastropoda" family="Drilliidae" genus="Kylix" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86C85BBDDFFC76FB77" box="[867,926,1220,1245]" italics="true" pageId="46" pageNumber="47">Kylix</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but none have a carinate first whorl.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8936D2FA4BFF86CBFFBDFCFD8CFAC6" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C364D51FA4BFF86CBFFBDFCFE09FAAA" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[199,481,1255,1280]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86CBFFBDFCFE8BFAAA" box="[199,355,1255,1280]" italics="true" pageId="46" pageNumber="47">Clathrodrillia</emphasis>
|
|||
|
|
Dall, 1918
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs in possessing a smooth, not carinate protoconch.
|
|||
|
|
<taxonomicName id="4C364D51FA4BFF86CF5CBDFCFA74FAAA" authority="Woodring, 1928" authorityName="Woodring" authorityYear="1928" box="[1124,1436,1255,1280]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86CF5CBDFCFB37FAAA" box="[1124,1247,1255,1280]" italics="true" pageId="46" pageNumber="47">Agladrillia</emphasis>
|
|||
|
|
Woodring, 1928
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs in possessing only a carinate second protoconch whorl, a varix positioned ½-turn from the edge of the outer lip, and axials obsolete from the varix to the outer lip.
|
|||
|
|
<taxonomicName id="4C364D51FA4BFF86C834BC34FC6FFAE2" box="[780,903,1327,1352]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86C834BC34FC6FFAE2" box="[780,903,1327,1352]" italics="true" pageId="46" pageNumber="47">Agladrillia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
also has a more acutely “pinched” anterior, as opposed to the tapered one of
|
|||
|
|
<taxonomicName id="4C364D51FA4BFF86CAD0BC4FFDB4FAC7" box="[488,604,1364,1389]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86CAD0BC4FFDB4FAC7" box="[488,604,1364,1389]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32C6559FA4BFF86CBFFBC6CFB63FA3A" box="[199,1163,1399,1424]" pageId="46" pageNumber="47" type="distribution">
|
|||
|
|
<paragraph id="8B8936D2FA4BFF86CBFFBC6CFB63FA3A" blockId="46.[151,1437,860,1424]" box="[199,1163,1399,1424]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86CBFFBC6CFE88FA3A" bold="true" box="[199,352,1399,1424]" pageId="46" pageNumber="47">Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="4C364D51FA4BFF86CA5FBC6CFE34FA3A" box="[359,476,1399,1424]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B942EAC0FA4BFF86CA5FBC6CFE34FA3A" box="[359,476,1399,1424]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is only known from off the northern coast of South
|
|||
|
|
<collectingCountry id="F3217642FA4BFF86CF1BBC6CFB6FFA3A" box="[1059,1159,1399,1424]" name="United States of America" pageId="46" pageNumber="47">America</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|