treatments-xml/data/19/10/75/1910752EFFA27912FF2352840F80F99A.xml

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<document id="26D4D2DA96C3F8E12EF504F12A3199A0" ID-DOI="10.11646/phytotaxa.257.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13673154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725402689584" checkinUser="felipe" docAuthor="Zhou, Jun-Liang, Su, Sheng-Yu, Su, Hong-Yan, Wang, Bo, Callac, Philippe, Guinberteau, Jacques &amp; Zhao, Rui-Lin" docDate="2016" docId="1910752EFFA27912FF2352840F80F99A" docLanguage="en" docName="phytotaxa.257.2.1.pdf" docOrigin="Phytotaxa 257 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.257.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Agaricus nigrogracilis R. L. Zhao 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="118" masterDocId="E5290D56FFB17901FFAB53300E1FFFE9" masterDocTitle="A description of eleven new species of Agaricus sections Xanthodermatei and Hondenses collected from Tibet and the surrounding areas" masterLastPageNumber="121" masterPageNumber="99" pageNumber="118" updateTime="1725423386967" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6B8292B73CE39F5DFFA133D884FB062C">A description of eleven new species of Agaricus sections Xanthodermatei and Hondenses collected from Tibet and the surrounding areas</mods:title>
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<mods:namePart id="6228A12E777EF5AAE3ADBA654CAC5480">Zhou, Jun-Liang</mods:namePart>
<mods:affiliation id="0F88D99341B4010175D9C2DE509073D2">State Key Laboratory of Mycology, Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China &amp; Institute of Microbiology, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:namePart id="9E72D33ABCF79EBB2354CF72CC163628">Su, Sheng-Yu</mods:namePart>
<mods:affiliation id="1CD9D58ACBC9594250C2DED4B68DCE72">Key Laboratory of Forest Disaster Warning and Control in Yunnan Province, College of Forestry, Southwest Forestry University, Kunming, Yunnan 650224, China</mods:affiliation>
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<mods:namePart id="1C52FA48E831EE36643EB4F31A748153">Su, Hong-Yan</mods:namePart>
<mods:affiliation id="62298A93ECF721F12D1D2025C1CDEC56">College of Agriculture and Biology, Dali University, Dali, Yunnan, 671003, China</mods:affiliation>
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<mods:namePart id="D7804FE51ADB2FCDC99545C4BC3F17E0">Wang, Bo</mods:namePart>
<mods:affiliation id="90F5E90D796B4E80FAB18C21DBBC4E2D">Sichuan Academy of Agaricultural sciences, Chengdu, Sichuan 610066, China</mods:affiliation>
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<mods:namePart id="20EC309A20A72D4C5B5338EA1BD9920A">Callac, Philippe</mods:namePart>
<mods:affiliation id="08AE75FFD16511CBA7BB49B0A8EC4517">MycSA, INRA, 33882 Villenave dOrnon, France</mods:affiliation>
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<mods:namePart id="2B60E12C65199D8B47A9A0A23DC71396">Guinberteau, Jacques</mods:namePart>
<mods:affiliation id="3560428A4A5BE7CCFB10583A6F8E4EE5">MycSA, INRA, 33882 Villenave dOrnon, France &amp; Center of Excellence in Fungal Research, and School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China &amp; World Agroforestry Centre, East and Central Asia, Kunming 650201, Yunnan, China</mods:affiliation>
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<mods:namePart id="FA6E5E2828B92AAD464EB15697537927">Zhao, Rui-Lin</mods:namePart>
<mods:affiliation id="88FC94942528D09DBB8F79FFE628CA29">State Key Laboratory of Mycology, Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="4FBE08E3006156122176829CD2F2E840">Phytotaxa</mods:title>
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<mods:date id="C89A9A70037197A7DF09712C1A985FD8">2016</mods:date>
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<mods:url id="270C7DA0E055FF305CAA808719794E2B">http://dx.doi.org/10.11646/phytotaxa.257.2.1</mods:url>
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<mods:identifier id="891F8F01EF1A934AAF4DE0766298A5FB" type="DOI">10.11646/phytotaxa.257.2.1</mods:identifier>
<mods:identifier id="DFFECF2454B9AD9563C3EE5E55F3F8F0" type="ISSN">1179-3163</mods:identifier>
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<treatment id="1910752EFFA27912FF2352840F80F99A" LSID="urn:lsid:plazi:treatment:1910752EFFA27912FF2352840F80F99A" httpUri="http://treatment.plazi.org/id/1910752EFFA27912FF2352840F80F99A" lastPageNumber="118" pageId="19" pageNumber="118">
<subSubSection id="D9A397B3FFA27912FF2352840FCFFE1A" pageId="19" pageNumber="118" type="nomenclature">
<paragraph id="9106C438FFA27912FF2352840D18FE26" blockId="19.[136,775,436,463]" box="[136,775,436,463]" pageId="19" pageNumber="118">
<heading id="CA4E7354FFA27912FF2352840D18FE26" box="[136,775,436,463]" fontSize="11" level="2" pageId="19" pageNumber="118" reason="3">
<taxonomicName id="56B9BFBBFFA27912FF2352840C18FE26" ID-CoL="5TN8R" authority="R.L. Zhao" authorityName="R. L. Zhao" authorityYear="2016" box="[136,519,436,463]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="19" pageNumber="118" phylum="Basidiomycota" rank="species" species="nigrogracilis" status="sp. nov.">
<emphasis id="A3CD182AFFA27912FF2352840F94FE26" bold="true" box="[136,395,436,463]" italics="true" pageId="19" pageNumber="118">Agaricus nigrogracilis</emphasis>
R.L. Zhao
</taxonomicName>
,
<emphasis id="A3CD182AFFA27912FDB952860C78FE26" box="[530,615,438,463]" italics="true" pageId="19" pageNumber="118">
<taxonomicNameLabel id="B8FEA551FFA27912FDB952860C78FE26" box="[530,615,438,463]" pageId="19" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="0982D8BDFFA27912FDDD52860CA4FE26" box="[630,699,438,463]" captionStart="FIGURE 6" captionStartId="15.[136,229,1899,1920]" captionTargetBox="[158,1428,192,1874]" captionTargetId="figure-14@15.[158,1430,190,1874]" captionTargetPageId="15" captionText="FIGURE 6. AF. Morphology of A. melanocapus (ZRL2011037 HOLOTYPE): AD Basidiomata, C shows yellow staining at the base of stipe; E. Basidiospores; F. Pileipellis hyphae. GK. Morphology of A. nigrogracilis (ZRL2012014 HOLOTYPE): G. Basidiospores; H. Pileipellis hyphae; IK. Basidiomata. Scale bar AD = 20 mm; IK = 10 mm; EH = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13673166" httpUri="https://zenodo.org/record/13673166/files/figure.png" pageId="19" pageNumber="118">Fig. 6</figureCitation>
: GK)
</heading>
</paragraph>
<paragraph id="9106C438FFA27912FF2352EA0FCFFE1A" blockId="19.[136,464,474,499]" box="[136,464,474,499]" pageId="19" pageNumber="118">MycoBank NO.: MB 812317</paragraph>
</subSubSection>
<subSubSection id="D9A397B3FFA27912FF2351120C8CFBE6" pageId="19" pageNumber="118" type="diagnosis">
<paragraph id="9106C438FFA27912FF2351120C09FD6A" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">Diagnosis: this species can be recognized by its small basidiomata, pileus covered with brown squamulose, and stiped surface that is white fibrillose to fibrillose squamulose. No discoloration occurs on cutting, and elongated basidiospores are present (5.25.9 × 2.93.4 μm).</paragraph>
<paragraph id="9106C438FFA27912FF1651BE0C91FD22" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">
Etymology: the word
<taxonomicName id="56B9BFBBFFA27912FE1951BE0C51FD4E" authorityName="R. L. Zhao" authorityYear="2016" box="[434,590,654,679]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="19" pageNumber="118" phylum="Basidiomycota" rank="species" species="nigrogracilis">“nigrogracilis</taxonomicName>
” refers to the black squamules on the pileus and slender habit; from nigrare (Latin) be black, and gracilis (Latin) slender.
</paragraph>
<paragraph id="9106C438FFA27912FF1651E60AD8FCFA" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">
Material examined:
<emphasis id="A3CD182AFFA27912FE0151E60CB0FD06" bold="true" box="[426,687,726,751]" pageId="19" pageNumber="118">
<collectingCountry id="E9AE84A8FFA27912FE0151E60FEAFD06" box="[426,501,726,751]" name="China" pageId="19" pageNumber="118">China</collectingCountry>
,
<collectingRegion id="537D0ADAFFA27912FDAD51E60CB0FD06" box="[518,687,726,751]" country="China" name="Yunnan" pageId="19" pageNumber="118">Yunnan Prov.</collectingRegion>
</emphasis>
, Kunming, Yeya Lake Forest Park,
<geoCoordinate id="F48DA2FFFFA27912FBC951E60AEBFD06" box="[1122,1268,726,751]" degrees="25" direction="north" minutes="07" orientation="latitude" pageId="19" pageNumber="118" precision="15" seconds="15" value="25.120834">N 25°0715″</geoCoordinate>
,
<geoCoordinate id="F48DA2FFFFA27912FAA351E60BBAFD06" box="[1288,1445,726,751]" degrees="102" direction="east" minutes="51" orientation="longitude" pageId="19" pageNumber="118" precision="15" seconds="47" value="102.86305">E 102°5147″</geoCoordinate>
, elevation
<quantity id="564169DDFFA27912FF5D51CA0F6FFCFA" box="[246,368,762,787]" metricMagnitude="3" metricUnit="m" metricValue="2.07894" pageId="19" pageNumber="118" unit="m" value="2078.94">2078.94 m</quantity>
,
<date id="E507E2F8FFA27912FED751CA0C0BFCFA" box="[380,532,762,787]" pageId="19" pageNumber="118" value="2012-06-30">30 June 2012</date>
, Rui-Lin Zhao (
<emphasis id="A3CD182AFFA27912FD6351CA0D45FCFA" box="[712,858,762,787]" italics="true" pageId="19" pageNumber="118">ZRL2012014</emphasis>
, HMAS 275735,
<typeStatus id="4E027A9AFFA27912FB8951CA0AA4FCFA" box="[1058,1211,762,787]" pageId="19" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
).
</paragraph>
<paragraph id="9106C438FFA27912FF16502E0C8CFBE6" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">
<emphasis id="A3CD182AFFA27912FF16502E0F1AFCDE" bold="true" box="[189,261,798,823]" pageId="19" pageNumber="118">Pileus</emphasis>
<quantity id="564169DDFFA27912FEA4502E0F7EFCDE" box="[271,353,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="118" unit="mm" value="35.0">35 mm</quantity>
in diam., convex to plano-convex; surface innately fibrillose, becoming appressed squamulose towards the margin, brown (oac 662) with a white background; edge decurved.
<emphasis id="A3CD182AFFA27912FBA950720A7EFCB2" bold="true" box="[1026,1121,834,859]" pageId="19" pageNumber="118">Context</emphasis>
<quantity id="564169DDFFA27912FBCC50720AB7FCB2" box="[1127,1192,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="118" unit="mm" value="3.0">3 mm</quantity>
thick at the disc, white and fleshy.
<emphasis id="A3CD182AFFA27912FEA550560F63FC96" bold="true" box="[270,380,870,895]" pageId="19" pageNumber="118">Lamellae</emphasis>
free and crowded; 45 series of plural-lamellulae,
<quantity id="564169DDFFA27912FC7950560A09FC96" box="[978,1046,870,895]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="118" unit="mm" value="4.0">4 mm</quantity>
broad, pink to brown; edge entire.
<emphasis id="A3CD182AFFA27912FF2350BA0EDAFC4A" bold="true" box="[136,197,906,931]" pageId="19" pageNumber="118">Stipe</emphasis>
<quantity id="564169DDFFA27912FF6050BA0F13FC4A" box="[203,268,906,931]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="118" unit="mm" value="5.0">5 mm</quantity>
(apex)
<quantity id="564169DDFFA27912FEC250BA0FA9FC4A" box="[361,438,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="118" unit="mm" value="10.0">10 mm</quantity>
(base) ×
<quantity id="564169DDFFA27912FDB550BA0C72FC4A" box="[542,621,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="19" pageNumber="118" unit="mm" value="75.0">75 mm</quantity>
(length); long clavate, surface white, dry, silky, and fibrillose to fibrillosesquamulose; hollow.
<emphasis id="A3CD182AFFA27912FEDD509E0FC4FC2E" bold="true" box="[374,475,942,967]" pageId="19" pageNumber="118">Annulus</emphasis>
single, membranous, subperonate,
<quantity id="564169DDFFA27912FCC2509E0DDFFC2E" box="[873,960,942,967]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="118" unit="mm" value="2.5">2.5 mm</quantity>
in diam., with a distinct brown edge; upper and lower surfaces smooth and white.
<emphasis id="A3CD182AFFA27912FD9450E20C60FC02" bold="true" box="[575,639,978,1003]" pageId="19" pageNumber="118">Odor</emphasis>
not iodine-like, but pleasant. Changing slightly yellow on touching the surface of the stipe; not discoloring on cutting.
</paragraph>
</subSubSection>
<subSubSection id="D9A397B3FFA27912FF16572A0F80F99A" pageId="19" pageNumber="118" type="description">
<paragraph id="9106C438FFA27912FF16572A0B36FBDA" blockId="19.[136,1452,546,1651]" box="[189,1321,1050,1075]" pageId="19" pageNumber="118">
<emphasis id="A3CD182AFFA27912FF16572A0FF6FBDA" bold="true" box="[189,489,1050,1075]" pageId="19" pageNumber="118">Macrochemical reactions</emphasis>
KOH reaction no change or slightly reddish. Schäffers reaction negative.
</paragraph>
<paragraph id="9106C438FFA27912FF16570E0F4FFAC6" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">
<emphasis id="A3CD182AFFA27912FF16570E0F7EFBBE" bold="true" box="[189,353,1086,1111]" pageId="19" pageNumber="118">Basidiospores</emphasis>
(
<figureCitation id="0982D8BDFFA27912FEDB570E0FCFFBBE" box="[368,464,1086,1111]" captionStart="FIGURE 6" captionStartId="15.[136,229,1899,1920]" captionTargetBox="[158,1428,192,1874]" captionTargetId="figure-14@15.[158,1430,190,1874]" captionTargetPageId="15" captionText="FIGURE 6. AF. Morphology of A. melanocapus (ZRL2011037 HOLOTYPE): AD Basidiomata, C shows yellow staining at the base of stipe; E. Basidiospores; F. Pileipellis hyphae. GK. Morphology of A. nigrogracilis (ZRL2012014 HOLOTYPE): G. Basidiospores; H. Pileipellis hyphae; IK. Basidiomata. Scale bar AD = 20 mm; IK = 10 mm; EH = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13673166" httpUri="https://zenodo.org/record/13673166/files/figure.png" pageId="19" pageNumber="118">Fig. 6-G</figureCitation>
) (4.6) 5.25.9 × 2.93.4 μm [x = 5.4 ± 0.3 × 3.2 ± 0.1 μm, Q = 1.52.0, Qm = 1.7 ± 0.1, n = 20 basidiospores]; elongated and occasionally ellipsoid, smooth, thick-walled, and brown.
<emphasis id="A3CD182AFFA27912FB2457520AF7FB92" bold="true" box="[1167,1256,1122,1147]" pageId="19" pageNumber="118">Basidia</emphasis>
13.116.4 × 4.5 5.9 μm, hyaline, smooth, clavate, and 4-spored.
<emphasis id="A3CD182AFFA27912FD0E57B60D50FB76" bold="true" box="[677,847,1158,1183]" pageId="19" pageNumber="118">Cheilocystidia</emphasis>
not observed.
<emphasis id="A3CD182AFFA27912FC5F57B60A80FB76" bold="true" box="[1012,1183,1158,1183]" pageId="19" pageNumber="118">Pleurocystidia</emphasis>
absent.
<emphasis id="A3CD182AFFA27912FB5057B60B71FB76" bold="true" box="[1275,1390,1158,1183]" pageId="19" pageNumber="118">Pileipellis</emphasis>
(
<figureCitation id="0982D8BDFFA27912FAD557B60EABFB2A" captionStart="FIGURE 6" captionStartId="15.[136,229,1899,1920]" captionTargetBox="[158,1428,192,1874]" captionTargetId="figure-14@15.[158,1430,190,1874]" captionTargetPageId="15" captionText="FIGURE 6. AF. Morphology of A. melanocapus (ZRL2011037 HOLOTYPE): AD Basidiomata, C shows yellow staining at the base of stipe; E. Basidiospores; F. Pileipellis hyphae. GK. Morphology of A. nigrogracilis (ZRL2012014 HOLOTYPE): G. Basidiospores; H. Pileipellis hyphae; IK. Basidiomata. Scale bar AD = 20 mm; IK = 10 mm; EH = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13673166" httpUri="https://zenodo.org/record/13673166/files/figure.png" pageId="19" pageNumber="118">Fig. 6-H</figureCitation>
) a cutis composed of hyphae 3.27.8 μm wide, cylindrical, branched, and smooth, some of which are constricted at the branch and contain light brown vacuolar pigments.
<emphasis id="A3CD182AFFA27912FCA857FE0D77FB0E" bold="true" box="[771,872,1230,1255]" pageId="19" pageNumber="118">Annulus</emphasis>
composed of hyphae 26 μm in diam., smooth, long clavoid, hyaline and branched, not constricted at septa; hyphae of the annulus margin contained brown to dark brown vacular pigments.
</paragraph>
<paragraph id="9106C438FFA27912FF16560A0FC0FABA" blockId="19.[136,1452,546,1651]" box="[189,479,1338,1363]" pageId="19" pageNumber="118">
<emphasis id="A3CD182AFFA27912FF16560A0F06FABA" bold="true" box="[189,281,1338,1363]" pageId="19" pageNumber="118">Habitat</emphasis>
solitary in forest.
</paragraph>
<paragraph id="9106C438FFA27912FF16566E0F80F99A" blockId="19.[136,1452,546,1651]" pageId="19" pageNumber="118">
<emphasis id="A3CD182AFFA27912FF16566E0F16FA9E" bold="true" box="[189,265,1374,1399]" pageId="19" pageNumber="118">Notes:</emphasis>
Molecular analysis based on ITS sequence indicates that this is a distinct species in section
<taxonomicName id="56B9BFBBFFA27912FB44566E0BBCFA9E" box="[1263,1443,1374,1399]" pageId="19" pageNumber="118" rank="section" section="Xanthodermatei">
<emphasis id="A3CD182AFFA27912FB44566E0BBCFA9E" box="[1263,1443,1374,1399]" italics="true" pageId="19" pageNumber="118">Xanthodermatei</emphasis>
</taxonomicName>
; however, the phylogenetic analysis using multi-gene sequences suggests that this species is a member of section
<taxonomicName id="56B9BFBBFFA27912FF2356960F1CFA56" box="[136,259,1446,1471]" pageId="19" pageNumber="118" rank="section" section="Hondenses">
<emphasis id="A3CD182AFFA27912FF2356960F1CFA56" box="[136,259,1446,1471]" italics="true" pageId="19" pageNumber="118">Hondenses</emphasis>
</taxonomicName>
. We performed the sequencing and analysis a second time with the same results, such that we were unable to clearly place this species in any section. Correspondingly, the morphology of this species was also quite different from that of the known species in those sections: it lacked color changes after flesh exposure and had a pleasant odor and a negative KOH reaction.
<taxonomicName id="56B9BFBBFFA27912FE7255220CD2F9C2" box="[473,717,1554,1579]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="19" pageNumber="118" phylum="Basidiomycota" rank="species" species="bukavuensis">
<emphasis id="A3CD182AFFA27912FE7255220CD2F9C2" box="[473,717,1554,1579]" italics="true" pageId="19" pageNumber="118">Agaricus bukavuensis</emphasis>
</taxonomicName>
is a species lacking a color change on cutting, but it has a bulbous stipe and larger basidiospores (5.46.9 × 44.9 μm) than those of
<taxonomicName id="56B9BFBBFFA27912FCD855060A3BF9A6" authorityName="R. L. Zhao" authorityYear="2016" box="[883,1060,1590,1615]" class="Agaricomycetes" family="Agaricaceae" genus="Agaricus" kingdom="Fungi" order="Agaricales" pageId="19" pageNumber="118" phylum="Basidiomycota" rank="species" species="nigrogracilis">
<emphasis id="A3CD182AFFA27912FCD855060A3BF9A6" box="[883,1060,1590,1615]" italics="true" pageId="19" pageNumber="118">A. nigrogracilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="F528B9C9FFA27912FB9F55060EDCF99A" author="Heinemann, P. &amp; Goossens-Fontana, M." pageId="19" pageNumber="118" pagination="99 - 119" refId="ref14614" refString="Heinemann, P. &amp; Goossens-Fontana, M. (1956) Agaricus I. In: Heinemann, P. (Ed.) Flore Iconographique des Champignons du Congo. Fasc., Bruxelles, pp. 99 - 119." type="book chapter" year="1956">Heinemann &amp; Goossens-Fontana 1956</bibRefCitation>
,
<bibRefCitation id="F528B9C9FFA27912FF65556A0F8DF99A" author="Heinemann, P." box="[206,402,1626,1651]" pageId="19" pageNumber="118" pagination="6 - 37" refId="ref14657" refString="Heinemann, P. (1978) Essai d'une cle de determination des genres Agaricus et Micropsalliota. Sydowia 30: 6 - 37." type="journal article" year="1978">Heinemann 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>