600 lines
82 KiB
XML
600 lines
82 KiB
XML
|
<document id="3820181CDFC66B3428C8F5BDC7D46038" ID-CLB-Dataset="27879" ID-DOI="10.11646/zootaxa.4526.4.1" ID-GBIF-Dataset="69cd7636-9ec1-4c3b-98ce-d6ce90bc7042" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611755" ID-ZooBank="3CFFF3AC-C447-4FCE-B6F8-D2B7BAE8B678" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697705242" checkinUser="plazi" docAuthor="Valdés, Ángel, Lundsten, Lonny & Wilson, Nerida G." docDate="2018" docId="03BE87CCFFA5DE099B8E9A757DCE58E0" docLanguage="en" docName="zootaxa.4526.4.1.pdf" docOrigin="Zootaxa 4526 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tritonia nigritigris Valdés & Lundsten & Wilson 2018, sp. nov." docType="treatment" docUuid="F357AF10-798E-4339-BD7A-D473BDD9BA83" docUuidSource="ZooBank" docVersion="10" lastPageNumber="408" masterDocId="FF87FFB4FFA6DE0E9B199F7E7D365F43" masterDocTitle="Five new deep-sea species of nudibranchs (Gastropoda: Heterobranchia: Cladobranchia) from the Northeast Pacific" masterLastPageNumber="433" masterPageNumber="401" pageNumber="404" updateTime="1698714326354" updateUser="ExternalLinkService">
|
|||
|
|
<mods:mods id="C28747198D6409E74BA4BA9D611A8F31" xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo id="85EAB0FCA8BEB8349987A3513822D784">
|
|||
|
|
<mods:title id="FE3C58DD2B636C82B58B335EDB32A927">Five new deep-sea species of nudibranchs (Gastropoda: Heterobranchia: Cladobranchia) from the Northeast Pacific</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name id="3AFC720928313784ABB058F9BC6FA6A2" type="personal">
|
|||
|
|
<mods:role id="F82397E61DE75F8018C3654D5499821B">
|
|||
|
|
<mods:roleTerm id="8BCC7D401F5E6BEE1F4B62F3F4F54D17">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="B111E41F68D23C440E954BDA9D88D303">Valdés, Ángel</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="1BB2F17CAEFFF137781FC622F5F9C6E5" type="personal">
|
|||
|
|
<mods:role id="CB8B3A92921163D1EEF4BDA738B484AE">
|
|||
|
|
<mods:roleTerm id="49E63B9123902223D3BB664F792AE719">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="562941E05BD56D6C5F87F13E2F3E3B16">Lundsten, Lonny</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name id="0A7F365BB0B481FC96B685955B4FAA19" type="personal">
|
|||
|
|
<mods:role id="2C691E16BC5C853EEC5FDB68E7608CEB">
|
|||
|
|
<mods:roleTerm id="305E9AA508A9D29526222DEB234AF8B4">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="72F12CECA05504087F6218305B04806E">Wilson, Nerida G.</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource id="4D0F7FF8FCD6FFE2222F55D58F6E94FB">text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem id="F4548B6744D088CCCB6D4F844B202683" type="host">
|
|||
|
|
<mods:titleInfo id="8B9EEF52946349FED707D135662C29DA">
|
|||
|
|
<mods:title id="633985DB35C71D85002A46774CE96D11">Zootaxa</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part id="2B63C8B645F056CBD8DF5F48070FB677">
|
|||
|
|
<mods:date id="C70153593E3B2FDA5B910E31CDE081A4">2018</mods:date>
|
|||
|
|
<mods:detail id="7FBC7CAF41561403003566BE7911CE57" type="pubDate">
|
|||
|
|
<mods:number id="97E1BD77D6E266DE6E3001796BAD5D4C">2018-12-03</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail id="99C05AA9F68692DC3B4730FEF33E6740" type="volume">
|
|||
|
|
<mods:number id="4F679585E06D9ECE271181E63CD052CE">4526</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail id="00E5753646714625E9C2B5841CB62DE7" type="issue">
|
|||
|
|
<mods:number id="7282181EB65D93D2FC61A5BAE724B807">4</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent id="1F975D370B104BF1B79A466F80F49A5E" unit="page">
|
|||
|
|
<mods:start id="59ECA026E77104C9C5CE4759A13A03A4">401</mods:start>
|
|||
|
|
<mods:end id="48C734E5E375ABB82C20AF2CF5763D45">433</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:classification id="3E184A2320A657111D8A27667B87F813">journal article</mods:classification>
|
|||
|
|
<mods:identifier id="7B6CDBBF70F7B20E62469642EDB3FEC2" type="CLB-Dataset">27879</mods:identifier>
|
|||
|
|
<mods:identifier id="4B8C68FF17C804595D1312888E62B4BE" type="DOI">10.11646/zootaxa.4526.4.1</mods:identifier>
|
|||
|
|
<mods:identifier id="48F93461726FD2CE9F1F334BFA63483C" type="GBIF-Dataset">69cd7636-9ec1-4c3b-98ce-d6ce90bc7042</mods:identifier>
|
|||
|
|
<mods:identifier id="C4ADED2F829E5B738AA43BCCE9D8C684" type="ISSN">1175-5326</mods:identifier>
|
|||
|
|
<mods:identifier id="83C6C5356A8ABAD3BB84AA08F12051D7" type="Zenodo-Dep">2611755</mods:identifier>
|
|||
|
|
<mods:identifier id="76081F3967A25D2CCC5D3CB46459907B" type="ZooBank">3CFFF3AC-C447-4FCE-B6F8-D2B7BAE8B678</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment id="03BE87CCFFA5DE099B8E9A757DCE58E0" ID-DOI="http://doi.org/10.5281/zenodo.5971394" ID-GBIF-Taxon="154507775" ID-Zenodo-Dep="5971394" LSID="urn:lsid:zoobank.org:act:F357AF10-798E-4339-BD7A-D473BDD9BA83" httpUri="http://treatment.plazi.org/id/03BE87CCFFA5DE099B8E9A757DCE58E0" lastPageId="7" lastPageNumber="408" pageId="3" pageNumber="404">
|
|||
|
|
<subSubSection id="C30D6551FFA5DE0D9B8E9A757CEB5A66" box="[151,477,1291,1317]" pageId="3" pageNumber="404" type="nomenclature">
|
|||
|
|
<paragraph id="8BA836DAFFA5DE0D9B8E9A757CEB5A66" blockId="3.[151,477,1291,1350]" box="[151,477,1291,1317]" pageId="3" pageNumber="404">
|
|||
|
|
<heading id="D0E081B6FFA5DE0D9B8E9A757CEB5A66" bold="true" box="[151,477,1291,1317]" fontSize="11" level="1" pageId="3" pageNumber="404" reason="1">
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9B8E9A757CEB5A66" bold="true" box="[151,477,1291,1317]" pageId="3" pageNumber="404">
|
|||
|
|
<taxonomicName id="4C174D59FFA5DE0D9B8E9A757C4E5A66" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[151,376,1291,1317]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="404" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9B8E9A757C4E5A66" bold="true" box="[151,376,1291,1317]" italics="true" pageId="3" pageNumber="404">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA5DE0D9A999A757CEB5A66" box="[384,477,1291,1317]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA5DE0D9B8E9A537C445A05" box="[151,370,1325,1350]" pageId="3" pageNumber="404" type="description">
|
|||
|
|
<paragraph id="8BA836DAFFA5DE0D9B8E9A537C445A05" blockId="3.[151,477,1291,1350]" box="[151,370,1325,1350]" pageId="3" pageNumber="404">
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132C2A5FFFA5DE0D9B869A537C355A05" box="[159,259,1325,1350]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="3" pageNumber="404">Figs. 2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132C2A5FFFA5DE0D9A179A507C055A05" box="[270,307,1326,1350]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="3" pageNumber="404">3A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132C2A5FFFA5DE0D9A269A537C5C5A05" box="[319,362,1325,1350]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1206,1228]" captionStartId-1="7.[151,250,1113,1135]" captionTargetBox-0="[164,1416,187,1185]" captionTargetBox-1="[151,1436,193,1092]" captionTargetId-0="figure@6.[151,1436,175,1185]" captionTargetId-1="figure@7.[151,1436,193,1092]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Drawings of the internal anatomy of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. General view of the internal anatomy. B. Reproductive system. Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; cns, central nervous system; dd, deferent duct; dg, digestive gland; es, esophagus; fgc, female gland complex; ht, heart; i, intestine; pe, penis; pr, prostate; rs, reproductive system; sg, salivary gland; sr, seminal receptacle; vg, vagina." captionText-1="FIGURE 5. SEMs of the radula and jaws of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw." httpUri-0="https://zenodo.org/record/2611763/files/figure.png" httpUri-1="https://zenodo.org/record/2611765/files/figure.png" pageId="3" pageNumber="404">4–5</figureCitation>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA5DE0D9B8E9A0B7F0A5A95" pageId="3" pageNumber="404" type="materials_examined">
|
|||
|
|
<paragraph id="8BA836DAFFA5DE0D9B8E9A0B7F0A5A95" blockId="3.[151,1437,1397,1962]" pageId="3" pageNumber="404">
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9B8E9A0B7C705ACD" bold="true" box="[151,326,1397,1422]" pageId="3" pageNumber="404">Type material.</emphasis>
|
|||
|
|
<materialsCitation id="3B7F3C87FFA5DE0D9A549A0B7F0A5A95" ID-GBIF-Occurrence="2029165590" collectingDate="2016-08-15" collectionCode="MH, COI" elevation="1733" pageId="3" pageNumber="404" specimenCode="LACM 3553" specimenCount="1" typeStatus="holotype">
|
|||
|
|
<typeStatus id="54AC8878FFA5DE0D9A549A0B7C8C5ACD" box="[333,442,1397,1422]" pageId="3" pageNumber="404" type="holotype">Holotype</typeStatus>
|
|||
|
|
: Guide Seamount, Pacific Ocean (37.036532, -123.330875),
|
|||
|
|
<quantity id="4CEF9B3FFFA5DE0D9F729A0879E05ACE" box="[1131,1238,1398,1422]" metricMagnitude="3" metricUnit="m" metricValue="1.7334" pageId="3" pageNumber="404" unit="m" value="1733.4">
|
|||
|
|
<elevation id="003AD1E9FFA5DE0D9F729A0879E05ACE" box="[1131,1238,1398,1422]" metricMagnitude="3" metricUnit="m" metricValue="1.7334" pageId="3" pageNumber="404" unit="m" value="1733.4">1733.4 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
depth, ROV
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9E749A087DC65AF1" italics="true" pageId="3" pageNumber="404">Doc Ricketts</emphasis>
|
|||
|
|
(dive 882),
|
|||
|
|
<date id="FFA9101AFFA5DE0D9A619AE77F3A5AF1" box="[376,524,1433,1458]" pageId="3" pageNumber="404" value="2016-08-15">
|
|||
|
|
<collectingDate id="EFEDE9F2FFA5DE0D9A619AE77F3A5AF1" box="[376,524,1433,1458]" pageId="3" pageNumber="404" value="2016-08-15">15 Aug 2016</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, 82 mm preserved length, dissected (
|
|||
|
|
<specimenCode id="DBB19EA1FFA5DE0D98B09AE479085AF1" box="[937,1086,1433,1458]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="404" type="Museum">LACM 3553</specimenCode>
|
|||
|
|
), GenBank accession numbers:
|
|||
|
|
<accessionNumber id="9444AB39FFA5DE0D9B8E9AC37C2E5A95" box="[151,280,1469,1494]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH756138" pageId="3" pageNumber="404">
|
|||
|
|
<collectionCode id="ED06AE1FFFA5DE0D9B8E9AC37DF25A95" box="[151,196,1469,1494]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="3" pageNumber="404">MH</collectionCode>
|
|||
|
|
756138
|
|||
|
|
</accessionNumber>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9A319AC07C6E5A96" box="[296,344,1470,1493]" italics="true" pageId="3" pageNumber="404">
|
|||
|
|
<collectionCode id="ED06AE1FFFA5DE0D9A319AC07C6E5A96" box="[296,344,1470,1493]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="3" pageNumber="404" type="Herbarium">COI</collectionCode>
|
|||
|
|
</emphasis>
|
|||
|
|
),
|
|||
|
|
<accessionNumber id="9444AB39FFA5DE0D9A699AC37CC75A95" box="[368,497,1469,1494]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH756133" pageId="3" pageNumber="404">
|
|||
|
|
<collectionCode id="ED06AE1FFFA5DE0D9A699AC37CAB5A95" box="[368,413,1469,1494]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="3" pageNumber="404">MH</collectionCode>
|
|||
|
|
756133
|
|||
|
|
</accessionNumber>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D99189AC37F1D5A95" box="[513,555,1469,1494]" italics="true" pageId="3" pageNumber="404">16S</emphasis>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA5DE0B9BDE9A9F7E965842" lastPageId="5" lastPageNumber="406" pageId="3" pageNumber="404" type="description">
|
|||
|
|
<paragraph id="8BA836DAFFA5DE0D9BDE9A9F7F9A58E9" blockId="3.[151,1437,1397,1962]" pageId="3" pageNumber="404">
|
|||
|
|
<emphasis id="B963EAC8FFA5DE0D9BDE9A9F7C6F5AB9" bold="true" box="[199,345,1505,1530]" pageId="3" pageNumber="404">Description.</emphasis>
|
|||
|
|
Body elongate, narrow (
|
|||
|
|
<figureCitation id="132C2A5FFFA5DE0D99769A9F7FFF5AB9" box="[623,713,1505,1530]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="3" pageNumber="404">Fig. 2A</figureCitation>
|
|||
|
|
). Notum outline with four invaginations, completely surrounded by numerous, densely packed glandular projections (
|
|||
|
|
<figureCitation id="132C2A5FFFA5DE0D99E2997B7E6E595D" box="[763,856,1541,1566]" captionStart="FIGURE 2" captionStartId="4.[151,250,1838,1860]" captionTargetBox="[151,1436,193,1817]" captionTargetId="figure@4.[151,1436,193,1817]" captionTargetPageId="4" captionText="FIGURE 2. Digital images of live animals showing color in life. A. Tritonia nigritigris sp. nov., holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B. Tritonia sp. (SIO-BIC M12395), lab photo. C. Dendronotus claguei sp. nov., holotype (LACM 3555), in situ. D. Ziminella vrijenhoeki sp. nov., holotype (SIO-BIC M12137), lab photo. E. Ziminella vrijenhoeki sp. nov., paratype (SIO-BIC M12135), lab photo. F. Cuthona methana sp. nov., holotype (SIO-BIC M12413), lab photo. G. Cuthona methana sp. nov., paratype (SIO-BIC M12412), lab photo. H. Aeolidia libitinaria sp. nov., holotype (SIO- BIC M12138), lab photo." httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="3" pageNumber="404">Fig. 2A</figureCitation>
|
|||
|
|
), giving the animal a ruffled appearance. Dorsum covered with numerous, densely packed elongate tubercles or projections, oriented in different directions, forming a zigzagging pattern in some areas. Velum wide, forming two triangular lateral projections, with several frontal irregular indentations. Rhinophores short and wide, with 9 simple lamellae, located on anterior end of notum. Rhinophoral sheaths elongate, with 12 branched papillae around distal edges. Foot smooth, rounded posteriorly, visible dorsally at posterior end and in notum invaginations. Genital and anal openings on right side of body. Background color of notum black. Dorsal tubercles primarily white to greyish-brown near posterior notum, giving the body a light appearance, becoming darker towards head, turning completely black on anterior notum; this gives a black and white appearance with a greyish transitional area mid-length, forming a wedge of black pigment. Notum edge primarily white, except for anterior-most end that becomes dark grey. Velum completely black. Rhinophoral sheaths black with grey edges. Rhinophores with black stalks and beige sheath papillae and clubs. Foot dark grey, surrounded by a thin white line.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF686652FFA2DE0A9B8E98507CAE589E" httpUri="https://zenodo.org/record/2611759/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1838,1860]" targetBox="[151,1436,193,1817]" targetPageId="4">
|
|||
|
|
<paragraph id="8BA836DAFFA2DE0A9B8E98507CAE589E" blockId="4.[151,1436,1838,2013]" pageId="4" pageNumber="405">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9B8E98507C225800" bold="true" box="[151,276,1838,1860]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
|
|||
|
|
Digital images of live animals showing color in life. A.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A9847985179285807" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[862,1054,1839,1860]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9847985179285807" box="[862,1054,1839,1860]" italics="true" pageId="4" pageNumber="405">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9F3C985079415800" bold="true" box="[1061,1143,1838,1859]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A9F3C985079415800" box="[1061,1143,1838,1859]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (LACM 3553), in situ, right side invaginations indicated with arrows. B.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A99DC98307E215820" authorityName="Cuvier" authorityYear="1797" box="[709,791,1870,1891]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A99DC98307E215820" box="[709,791,1870,1891]" italics="true" pageId="4" pageNumber="405">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. (SIO-BIC M12395), lab photo. C.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A9FBB983278415821" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1186,1399,1868,1890]" class="Gastropoda" family="Dendronotidae" genus="Dendronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="claguei" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9FBB983278415821" box="[1186,1399,1868,1890]" italics="true" pageId="4" pageNumber="405">Dendronotus claguei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9E6498337DF558C2" bold="true" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A9E6498337DF558C2" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (LACM 3555), in situ. D.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A992598157E2058C2" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[572,790,1899,1921]" class="Gastropoda" family="Paracoryphellidae" genus="Ziminella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="vrijenhoeki" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A992598157E2058C2" box="[572,790,1899,1921]" italics="true" pageId="4" pageNumber="405">Ziminella vrijenhoeki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A980498127E5858C2" bold="true" box="[797,878,1900,1921]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A980498127E5858C2" box="[797,878,1900,1921]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (SIO-BIC
|
|||
|
|
<accessionNumber id="9444AB39FFA2DE0A9F5F9815799658C2" box="[1094,1184,1899,1921]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M12137" pageId="4" pageNumber="405" type="EnaNcbi">M12137</accessionNumber>
|
|||
|
|
), lab photo. E.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A9E2298157C3F58E3" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" class="Gastropoda" family="Paracoryphellidae" genus="Ziminella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="vrijenhoeki" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9E2298157C3F58E3" italics="true" pageId="4" pageNumber="405">Ziminella vrijenhoeki</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9A0998F47C5758DC" bold="true" box="[272,353,1930,1951]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A9A0998F47C5758DC" box="[272,353,1930,1951]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, paratype (SIO-BIC
|
|||
|
|
<accessionNumber id="9444AB39FFA2DE0A992198F47FA758E3" box="[568,657,1930,1952]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M12135" pageId="4" pageNumber="405" type="EnaNcbi">M12135</accessionNumber>
|
|||
|
|
), lab photo. F.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A983298F47ED658E3" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[811,992,1930,1952]" class="Gastropoda" family="Fionidae" genus="Cuthona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="methana" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A983298F47ED658E3" box="[811,992,1930,1952]" italics="true" pageId="4" pageNumber="405">Cuthona methana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A98FE98F4790F58DC" bold="true" box="[999,1081,1930,1951]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A98FE98F4790F58DC" box="[999,1081,1930,1951]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (SIO-BIC M12413), lab photo. G.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A9BE298D77C8658FC" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[251,432,1961,1983]" class="Gastropoda" family="Fionidae" genus="Cuthona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="methana" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9BE298D77C8658FC" box="[251,432,1961,1983]" italics="true" pageId="4" pageNumber="405">Cuthona methana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9AAE98D77F3F58FD" bold="true" box="[439,521,1961,1982]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A9AAE98D77F3F58FD" box="[439,521,1961,1982]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, paratype (SIO-BIC M12412), lab photo. H.
|
|||
|
|
<taxonomicName id="4C174D59FFA2DE0A98C098D779AA58FC" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[985,1180,1961,1983]" class="Gastropoda" family="Aeolidiidae" genus="Aeolidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="405" phylum="Mollusca" rank="species" species="libitinaria" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A98C098D779AA58FC" box="[985,1180,1961,1983]" italics="true" pageId="4" pageNumber="405">Aeolidia libitinaria</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA2DE0A9FBA98D779C258FD" bold="true" box="[1187,1268,1961,1982]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA2DE0A9FBA98D779C258FD" box="[1187,1268,1961,1982]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype (SIO- BIC M12138), lab photo.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF686652FFA3DE0B9B8E9BC57F8E5A6E" httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1211,1233]" targetBox="[151,1437,193,1189]" targetPageId="5">
|
|||
|
|
<paragraph id="8BA836DAFFA3DE0B9B8E9BC57F8E5A6E" blockId="5.[151,1436,1211,1325]" pageId="5" pageNumber="406">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9B8E9BC57C245B93" bold="true" box="[151,274,1211,1233]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
|
|||
|
|
Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B9BAB9BA57C475BB3" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[178,369,1243,1264]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="406" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9BAB9BA57C475BB3" box="[178,369,1243,1264]" italics="true" pageId="5" pageNumber="406">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9A619BA47CFE5BAC" bold="true" box="[376,456,1242,1263]" pageId="5" pageNumber="406">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA3DE0B9A619BA47CFE5BAC" box="[376,456,1242,1263]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(LACM 3553), lateral view. B. Paratype of
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B98939BA479695BB3" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[906,1119,1242,1264]" class="Gastropoda" family="Dendronotidae" genus="Dendronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="406" phylum="Mollusca" rank="species" species="claguei" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B98939BA479695BB3" box="[906,1119,1242,1264]" italics="true" pageId="5" pageNumber="406">Dendronotus claguei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9F7C9BA479835BAC" bold="true" box="[1125,1205,1242,1263]" pageId="5" pageNumber="406">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA3DE0B9F7C9BA479835BAC" box="[1125,1205,1242,1263]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(LACM 3555), lateral view. C–D.
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B9A0C9B877C515A4D" authorityName="Cuvier" authorityYear="1797" box="[277,359,1273,1294]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="406" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9A0C9B877C515A4D" box="[277,359,1273,1294]" italics="true" pageId="5" pageNumber="406">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA836DAFFA3DE0B9BDE9A277F595909" blockId="5.[151,1437,1369,2009]" pageId="5" pageNumber="406">
|
|||
|
|
Digestive system with large, muscular buccal bulb (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B98359A277EBD5A31" box="[812,907,1369,1394]" captionStart="FIGURE 4" captionStartId="6.[151,250,1206,1228]" captionTargetBox="[164,1416,187,1185]" captionTargetId="figure@6.[151,1436,175,1185]" captionTargetPageId="6" captionText="FIGURE 4. Drawings of the internal anatomy of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. General view of the internal anatomy. B. Reproductive system. Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; cns, central nervous system; dd, deferent duct; dg, digestive gland; es, esophagus; fgc, female gland complex; ht, heart; i, intestine; pe, penis; pr, prostate; rs, reproductive system; sg, salivary gland; sr, seminal receptacle; vg, vagina." httpUri="https://zenodo.org/record/2611763/files/figure.png" pageId="5" pageNumber="406">Fig. 4A</figureCitation>
|
|||
|
|
). Esophagus wide, connecting anteriorly into buccal bulb, where two large, flat salivary glands connect. Intestine emerging dorsally from left side of digestive gland, forming a loop towards the right side of body where it opens into the anus (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9F209ADF79A55AF9" box="[1081,1171,1441,1466]" captionStart="FIGURE 3" captionStartId="5.[151,250,1211,1233]" captionTargetBox="[151,1437,193,1189]" captionTargetId="figure@5.[151,1437,193,1189]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the external morphology of preserved specimens of Tritonioidea and Dendronotoidea. A. Holotype of Tritonia nigritigris sp. nov. (LACM 3553), lateral view. B. Paratype of Dendronotus claguei sp. nov. (LACM 3555), lateral view. C–D. Tritonia sp. (SIO-BIC M12395), C. dorsal view of the head, D. lateral view. Abbreviations: a, anus; pe, penis. Arrow indicates damage on the right side of the body." httpUri="https://zenodo.org/record/2611761/files/figure.png" pageId="5" pageNumber="406">Fig. 3A</figureCitation>
|
|||
|
|
). Radular formula 44 × 50.1.
|
|||
|
|
<quantity id="4CEF9B3FFFA3DE0B9BD49ABB7C315A9D" box="[205,263,1477,1502]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="406" unit="in" value="50.0">50 in</quantity>
|
|||
|
|
<typeStatus id="54AC8878FFA3DE0B9A179ABB7C425A9D" box="[270,372,1477,1502]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
|
|||
|
|
. Rachidian teeth narrow, with a large, sharp central cusp about twice as large as tooth bases and 3 irregular denticles on each side of central cusp (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B99D49A977E1F5941" box="[717,809,1513,1538]" captionStart="FIGURE 5" captionStartId="7.[151,250,1113,1135]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@7.[151,1436,193,1092]" captionTargetPageId="7" captionText="FIGURE 5. SEMs of the radula and jaws of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw." httpUri="https://zenodo.org/record/2611765/files/figure.png" pageId="5" pageNumber="406">Fig. 5A</figureCitation>
|
|||
|
|
). Lateral teeth hook-shaped, with sharp, elongate cusps lacking denticles (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9A7C99737C8B5965" box="[357,445,1549,1574]" captionStart="FIGURE 5" captionStartId="7.[151,250,1113,1135]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@7.[151,1436,193,1092]" captionTargetPageId="7" captionText="FIGURE 5. SEMs of the radula and jaws of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw." httpUri="https://zenodo.org/record/2611765/files/figure.png" pageId="5" pageNumber="406">Fig. 5B</figureCitation>
|
|||
|
|
). Outer lateral teeth very elongate with reduced bases (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9F34997379B15965" box="[1069,1159,1549,1574]" captionStart="FIGURE 5" captionStartId="7.[151,250,1113,1135]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@7.[151,1436,193,1092]" captionTargetPageId="7" captionText="FIGURE 5. SEMs of the radula and jaws of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw." httpUri="https://zenodo.org/record/2611765/files/figure.png" pageId="5" pageNumber="406">Fig. 5C</figureCitation>
|
|||
|
|
). Jaws elongate, with no differentiate masticatory border (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9911994F7F545909" box="[520,610,1585,1610]" captionStart="FIGURE 5" captionStartId="7.[151,250,1113,1135]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@7.[151,1436,193,1092]" captionTargetPageId="7" captionText="FIGURE 5. SEMs of the radula and jaws of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw." httpUri="https://zenodo.org/record/2611765/files/figure.png" pageId="5" pageNumber="406">Fig. 5D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA836DAFFA3DE0B9BDE992B7E965842" blockId="5.[151,1437,1369,2009]" pageId="5" pageNumber="406">
|
|||
|
|
Reproductive system with large, highly convoluted ampulla forming several loops (
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9FBA992B7836592D" box="[1187,1280,1621,1646]" captionStart="FIGURE 4" captionStartId="6.[151,250,1206,1228]" captionTargetBox="[164,1416,187,1185]" captionTargetId="figure@6.[151,1436,175,1185]" captionTargetPageId="6" captionText="FIGURE 4. Drawings of the internal anatomy of the holotype of Tritonia nigritigris sp. nov. (LACM 3553). A. General view of the internal anatomy. B. Reproductive system. Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; cns, central nervous system; dd, deferent duct; dg, digestive gland; es, esophagus; fgc, female gland complex; ht, heart; i, intestine; pe, penis; pr, prostate; rs, reproductive system; sg, salivary gland; sr, seminal receptacle; vg, vagina." httpUri="https://zenodo.org/record/2611763/files/figure.png" pageId="5" pageNumber="406">Fig. 4B</figureCitation>
|
|||
|
|
), opening on female gland complex where the prostate connects. Prostate narrow, elongate, forming a single loop and expanding into a wide and muscular deferent duct, which connects with a simple, conical penis. Vagina elongate, narrow, expanding abruptly in its distal 1/4
|
|||
|
|
<superScript id="7C629B92FFA3DE0B993F99BF7F04598C" attach="left" box="[550,562,1729,1743]" fontSize="6" pageId="5" pageNumber="406">th</superScript>
|
|||
|
|
and opening into the spherical bursa copulatrix. The seminal receptacle is a small spherical evagination located close to mid-length in the vagina.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA3DE089BDE9872795B5ACE" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406" type="biology_ecology">
|
|||
|
|
<paragraph id="8BA836DAFFA3DE0B9BDE98727DDF589A" blockId="5.[151,1437,1369,2009]" pageId="5" pageNumber="406">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9BDE98727C115866" bold="true" box="[199,295,1804,1829]" pageId="5" pageNumber="406">Biology.</emphasis>
|
|||
|
|
This species was collected at
|
|||
|
|
<quantity id="4CEF9B3FFFA3DE0B999198737FC35867" box="[648,757,1805,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.7334" pageId="5" pageNumber="406" unit="m" value="1733.4">1733.4 m</quantity>
|
|||
|
|
depth on the Guide Seamount, about 55 nautical miles off Davenport,
|
|||
|
|
<collectingRegion id="49D3F838FFA3DE0B9A39984E7CA4580A" box="[288,402,1840,1865]" country="United States of America" name="California" pageId="5" pageNumber="406">California</collectingRegion>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132C2A5FFFA3DE0B9ABF984E7CDA580A" box="[422,492,1840,1865]" captionStart="FIGURE 1" captionStartId="2.[151,250,1630,1652]" captionTargetBox="[151,1436,195,1606]" captionTargetId="figure@2.[151,1436,195,1606]" captionTargetPageId="2" captionText="FIGURE 1. Distribution map of the collection localities in this paper including depth ranges." httpUri="https://zenodo.org/record/2611757/files/figure.png" pageId="5" pageNumber="406">Fig. 1</figureCitation>
|
|||
|
|
). The specimen was collected on a steep section of volcanoclastic rock. Dense populations of
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B9A55982B7C89582F" authorityName="Wright" authorityYear="1869" box="[332,447,1877,1900]" class="Anthozoa" family="Isididae" genus="Keratoisis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="406" phylum="Cnidaria" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9A55982B7C89582F" box="[332,447,1877,1900]" italics="true" pageId="5" pageNumber="406">Keratoisis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. (isidid octocoral) were observed in close proximity. Other organisms included comatulid and hyocrinid crinoids, primnoid octocorals,
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B981F98067E9058D2" box="[774,934,1912,1937]" italics="true" pageId="5" pageNumber="406">Evoloplosoma</emphasis>
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B98B49806791558D2" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[941,1059,1912,1937]" class="Gastropoda" family="Dendronotidae" genus="Dendronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="406" phylum="Mollusca" rank="species" species="claguei">
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B98C89806791558D2" box="[977,1059,1912,1937]" italics="true" pageId="5" pageNumber="406">claguei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(sea star), additional unidentified sea stars,
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B9A1598E37C6158F7" box="[268,343,1949,1972]" class="Bivalvia" family="Limidae" genus="Acesta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Limoida" pageId="5" pageNumber="406" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B9A1598E37C6158F7" box="[268,343,1949,1972]" italics="true" pageId="5" pageNumber="406">Acesta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. (bivalve),
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B991D98E07F6558F6" authorityName="Bowerbank" authorityYear="1862" box="[516,595,1950,1973]" class="Hexactinellida" family="Farreidae" genus="Farrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hexactinosida" pageId="5" pageNumber="406" phylum="Porifera" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA3DE0B991D98E07F6558F6" box="[516,595,1950,1973]" italics="true" pageId="5" pageNumber="406">Farrea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. (hexactinellid sponge), and large and abundant
|
|||
|
|
<taxonomicName id="4C174D59FFA3DE0B9FAE98E2787058F6" box="[1207,1350,1948,1973]" class="Actinopterygii" family="Macrouridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="family">Macrouridae</taxonomicName>
|
|||
|
|
(rattail fishes).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF686652FFA0DE089B8E9BC879515A6A" httpUri="https://zenodo.org/record/2611763/files/figure.png" pageId="6" pageNumber="407" startId="6.[151,250,1206,1228]" targetBox="[164,1416,187,1185]" targetPageId="6">
|
|||
|
|
<paragraph id="8BA836DAFFA0DE089B8E9BC879515A6A" blockId="6.[151,1436,1206,1321]" pageId="6" pageNumber="407">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089B8E9BC87C255B8F" bold="true" box="[151,275,1206,1228]" pageId="6" pageNumber="407">FIGURE 4.</emphasis>
|
|||
|
|
Drawings of the internal anatomy of the holotype of
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE08982C9BC67EC25B8E" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[821,1012,1208,1229]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE08982C9BC67EC25B8E" box="[821,1012,1208,1229]" italics="true" pageId="6" pageNumber="407">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA0DE0898E29BC9797A5B8F" bold="true" box="[1019,1100,1207,1228]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA0DE0898E29BC9797A5B8F" box="[1019,1100,1207,1228]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(LACM 3553). A. General view of the internal anatomy. B. Reproductive system. Abbreviations: am, ampulla; bb, buccal bulb; bc, bursa copulatrix; cns, central nervous system; dd, deferent duct; dg, digestive gland; es, esophagus; fgc, female gland complex; ht, heart; i, intestine; pe, penis; pr, prostate; rs, reproductive system; sg, salivary gland; sr, seminal receptacle; vg, vagina.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA836DAFFA0DE089BDE9A2E795B5ACE" blockId="6.[151,1437,1360,2033]" pageId="6" pageNumber="407">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089BDE9A2E7CF85A2A" bold="true" box="[199,462,1360,1385]" pageId="6" pageNumber="407">Phylogenetic position.</emphasis>
|
|||
|
|
Based on available sequence data and the phylogenetic analyses conducted,
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089E5B9A2F7C3F5ACF" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089E5B9A2F7C3F5ACF" italics="true" pageId="6" pageNumber="407">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089A169A0A7C5F5ACE" bold="true" box="[271,361,1396,1421]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA0DE089A169A0A7C5F5ACE" box="[271,361,1396,1421]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is sister to the Antarctic species
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE0899C09A0A793D5ACE" authority="Eliot, 1907" authorityName="Eliot" authorityYear="1907" box="[729,1035,1396,1421]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="belli">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE0899C09A0A7EBE5ACE" box="[729,904,1396,1421]" italics="true" pageId="6" pageNumber="407">Tritoniella belli</emphasis>
|
|||
|
|
Eliot, 1907
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132C2A5FFFA0DE089F029A0A79695ACE" box="[1051,1119,1396,1421]" captionStart="FIGURE 6" captionStartId="8.[151,250,1949,1971]" captionTargetBox="[391,1173,195,1901]" captionTargetId="figure@8.[388,1173,195,1901]" captionTargetPageId="8" captionText="FIGURE 6. Bayesian consensus phylogenetic analyses of Tritonioidea and Dendronotoidea, with branches collapsed for clarity. The phylogenetic position of the species described herein (in bold) are represented in grey ovals. Posterior probability and maximum likelihood bootstrap values indicated for each branch." httpUri="https://zenodo.org/record/2611767/files/figure.png" pageId="6" pageNumber="407">Fig. 6</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA0DE089BDE9AE67FCA5A96" pageId="6" pageNumber="407" type="etymology">
|
|||
|
|
<paragraph id="8BA836DAFFA0DE089BDE9AE67FCA5A96" blockId="6.[151,1437,1360,2033]" pageId="6" pageNumber="407">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089BDE9AE67C785AF2" bold="true" box="[199,334,1432,1457]" pageId="6" pageNumber="407">Etymology.</emphasis>
|
|||
|
|
The species name is derived from the unique coloration of the
|
|||
|
|
<typeStatus id="54AC8878FFA0DE089F059AE679B75AF2" box="[1052,1153,1432,1457]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
|
|||
|
|
, in particular the broken, black stripes, which are reminiscent of those of a tiger.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA0DE089BDE9A9E7C3659B6" pageId="6" pageNumber="407" type="discussion">
|
|||
|
|
<paragraph id="8BA836DAFFA0DE089BDE9A9E7C3659B6" blockId="6.[151,1437,1360,2033]" pageId="6" pageNumber="407">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089BDE9A9E7C0D5ABA" bold="true" box="[199,315,1504,1529]" pageId="6" pageNumber="407">Remarks.</emphasis>
|
|||
|
|
The most common species in relatively deep-waters of the eastern Pacific is
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089FA39A9F7C03595E" authority="(Pallas, 1788)" baseAuthorityName="Pallas" baseAuthorityYear="1788" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="tetraquetra">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089FA39A9F78AA5ABB" box="[1210,1436,1505,1528]" italics="true" pageId="6" pageNumber="407">Tritonia tetraquetra</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089B86997A7C1B595E" author="Pallas, P. S." box="[159,301,1540,1565]" pageId="6" pageNumber="407" pagination="229 - 249" refId="ref15780" refString="Pallas, P. S. (1788) Marina varia nova et rariora. Nova Acta Academiae Scientiarum Imperialis Petropolitanea, 2, 229 - 249." type="journal article" year="1788">Pallas, 1788</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, previously recognized as
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089978997B7EB6595E" authority="Bergh, 1894" authorityName="Bergh" authorityYear="1894" box="[609,896,1540,1565]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="diomedea">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089978997B7FDC595E" box="[609,746,1540,1565]" italics="true" pageId="6" pageNumber="407">T. diomedea</emphasis>
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE0899EB997A7EB6595E" author="Bergh, R." box="[754,896,1540,1565]" pageId="6" pageNumber="407" pagination="125 - 233" refId="ref13305" refString="Bergh, R. (1894) Die Opisthobranchien. Reports on the dredging operations off the west coast of Central America to the Galapagos, to the west coast of Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried on by the U. S. Fish Commission steamer " Albatross ", during 1891, Lieut. Commander Z. L. Tanner, U. S. N., commanding. Bulletin of the Museum of Comparative Zoology, 25, 125 - 233." type="journal article" year="1894">Bergh, 1894</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089895997B797A595E" author="Martynov, A. V." box="[908,1100,1540,1565]" pageId="6" pageNumber="407" pagination="267 - 294" refId="ref15496" refString="Martynov, A. V. (2006) Nudipleura. In: Kantor, Yu. I. & Sysoev, A. V. (Eds.), Marine and brackish water Gastropoda of Russia and adjacent countries: An illustrated catalogue. KMK Scientific Press, Moscow, pp. 267 - 294, pls. 131 - 138." type="book chapter" year="2006">Martynov (2006)</bibRefCitation>
|
|||
|
|
considered that
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089E1E997A78AA595E" author="Pallas, P. S." box="[1287,1436,1540,1565]" pageId="6" pageNumber="407" pagination="229 - 249" refId="ref15780" refString="Pallas, P. S. (1788) Marina varia nova et rariora. Nova Acta Academiae Scientiarum Imperialis Petropolitanea, 2, 229 - 249." type="journal article" year="1788">Pallas (1788)</bibRefCitation>
|
|||
|
|
original description of
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089AB099577F4C5903" authorityName="Pallas" authorityYear="1788" box="[425,634,1577,1600]" class="Gastropoda" family="Aeolididae" genus="Limax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="tetraquetra">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089AB099577F4C5903" box="[425,634,1577,1600]" italics="true" pageId="6" pageNumber="407">Limax tetraquetra</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the Kuril Islands corresponded to a specimen of the species commonly identified as
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089AB099337F055926" authorityName="Bergh" authorityYear="1894" box="[425,563,1612,1637]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="diomedea">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089AB099337F055926" box="[425,563,1612,1637]" italics="true" pageId="6" pageNumber="407">T. diomedea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE08996999327E875926" author="Behrens, D. W. & Hermosillo, A." box="[624,945,1612,1637]" pageId="6" pageNumber="407" refId="ref13266" refString="Behrens, D. W. & Hermosillo, A. (2005) Eastern Pacific nudibranchs. A guide to the Opisthobranchs from Alaska to Central America. Sea Challengers, Monterey, California, 137 pp." type="book" year="2005">Behrens & Hermosillo 2005</bibRefCitation>
|
|||
|
|
) and therefore proposed to use the name
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089E9E99337C2359CB" authorityName="Pallas" authorityYear="1788" class="Gastropoda" family="Aeolididae" genus="Limax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="tetraquetra">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089E9E99337C2359CB" italics="true" pageId="6" pageNumber="407">T. tetraquetra</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
for this species. Here we cautiously follow
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089823990E792259CA" author="Martynov, A. V." box="[826,1044,1648,1673]" pageId="6" pageNumber="407" pagination="267 - 294" refId="ref15496" refString="Martynov, A. V. (2006) Nudipleura. In: Kantor, Yu. I. & Sysoev, A. V. (Eds.), Marine and brackish water Gastropoda of Russia and adjacent countries: An illustrated catalogue. KMK Scientific Press, Moscow, pp. 267 - 294, pls. 131 - 138." type="book chapter" year="2006">Martynov’s (2006)</bibRefCitation>
|
|||
|
|
opinion, but clarification of the taxonomic status of this species requires comprehensive examination of specimens across its broad range (from the Kuril Islands to
|
|||
|
|
<collectingCountry id="F300764AFFA0DE089A5599C67C9F5993" box="[332,425,1720,1744]" name="Panama" pageId="6" pageNumber="407">Panama</collectingCountry>
|
|||
|
|
), because as in other nominal species with similar ranges, multiple cryptic species may be involved.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA0DE099BDE987F79CE5954" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="reference_group">
|
|||
|
|
<paragraph id="8BA836DAFFA0DE089BDE987F7ED258B2" blockId="6.[151,1437,1360,2033]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089BDE987F7CAA585B" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[199,412,1793,1816]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089BDE987F7CAA585B" box="[199,412,1793,1816]" italics="true" pageId="6" pageNumber="407">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089ABD987E7CC9585A" bold="true" box="[420,511,1792,1817]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA0DE089ABD987E7CC9585A" box="[420,511,1792,1817]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is externally different from other species of
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE0898E7987F796E585B" ID-CoL="7PJZ6" authorityName="Cuvier" authorityYear="1797" box="[1022,1112,1793,1816]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE0898E7987F796E585B" box="[1022,1112,1793,1816]" italics="true" pageId="6" pageNumber="407">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
described to date. The most similar species morphologically is the deep-water species
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089833985B79D0587E" ID-CoL="5953N" authority="Valdes et al., 2016" authorityName="Valdes" authorityYear="2016" box="[810,1254,1828,1853]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="newfoundlandica">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089833985B793E587E" box="[810,1032,1828,1853]" italics="true" pageId="6" pageNumber="407">T. newfoundlandica</emphasis>
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089F09985A79D0587E" author="Valdes, A. & Murillo, F. J. & McCarthy, J. B. & Yedinak, N." box="[1040,1254,1828,1853]" pageId="6" pageNumber="407" pagination="303 - 319" refId="ref17189" refString="Valdes, A., Murillo, F. J., McCarthy, J. B. & Yedinak, N. (2016) New deep-water records and species of North Atlantic nudibranchs (Mollusca, Gastropoda: Heterobranchia) with the description of a new species. Journal of the Marine Biological Association of the United Kingdom, 97, 303 - 319. https: // doi. org / 10.1017 / S 0025315416000394" type="journal article" year="2016">
|
|||
|
|
Valdés
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089F7B985B79AD587E" box="[1122,1179,1828,1853]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
|||
|
|
, 2016
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, described from the North Atlantic (
|
|||
|
|
<bibRefCitation id="EF864B2BFFA0DE089A6D98367F735822" author="Valdes, A. & Murillo, F. J. & McCarthy, J. B. & Yedinak, N." box="[372,581,1864,1889]" pageId="6" pageNumber="407" pagination="303 - 319" refId="ref17189" refString="Valdes, A., Murillo, F. J., McCarthy, J. B. & Yedinak, N. (2016) New deep-water records and species of North Atlantic nudibranchs (Mollusca, Gastropoda: Heterobranchia) with the description of a new species. Journal of the Marine Biological Association of the United Kingdom, 97, 303 - 319. https: // doi. org / 10.1017 / S 0025315416000394" type="journal article" year="2016">
|
|||
|
|
Valdés
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089AD198377F375822" box="[456,513,1864,1889]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
|||
|
|
2016
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). Both
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE08998E98377E135823" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[663,805,1865,1888]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE08998E98377E135823" box="[663,805,1865,1888]" italics="true" pageId="6" pageNumber="407">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA0DE08983598367EB15822" bold="true" box="[812,903,1864,1889]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA0DE08983598367EB15822" box="[812,903,1864,1889]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE0898A6983779AA5822" ID-CoL="5953N" box="[959,1180,1864,1889]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="newfoundlandica">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE0898A6983779AA5822" box="[959,1180,1864,1889]" italics="true" pageId="6" pageNumber="407">T. newfoundlandica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have narrow, elongate bodies with the anterior end wider, tapering posteriorly into a pointy, triangular end. However, whereas
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089E9E98137C6E58EA" ID-CoL="5953N" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="newfoundlandica">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089E9E98137C6E58EA" italics="true" pageId="6" pageNumber="407">T. newfoundlandica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is uniformly black, with the rhinophores, rhinophoral sheaths, velar appendages, and secondary gills brownish-red and the dorsum is smooth, lacking tubercles. On the contrary
|
|||
|
|
<taxonomicName id="4C174D59FFA0DE089F3198CB7980588F" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1064,1206,1973,1996]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="407" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089F3198CB7980588F" box="[1064,1206,1973,1996]" italics="true" pageId="6" pageNumber="407">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA0DE089FA798CA782E588E" bold="true" box="[1214,1304,1972,1997]" pageId="6" pageNumber="407">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA0DE089FA798CA782E588E" box="[1214,1304,1972,1997]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
has a more complex coloration of black, white and grey, and the dorsum is tuberculate.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF686652FFA1DE099B8E9B277EA35BCD" httpUri="https://zenodo.org/record/2611765/files/figure.png" pageId="7" pageNumber="408" startId="7.[151,250,1113,1135]" targetBox="[151,1436,193,1092]" targetPageId="7">
|
|||
|
|
<paragraph id="8BA836DAFFA1DE099B8E9B277EA35BCD" blockId="7.[151,1436,1113,1166]" pageId="7" pageNumber="408">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099B8E9B277C245B2C" bold="true" box="[151,274,1113,1135]" pageId="7" pageNumber="408">FIGURE 5.</emphasis>
|
|||
|
|
SEMs of the radula and jaws of the holotype of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999E09B247E8E5B2C" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[761,952,1114,1135]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999E09B247E8E5B2C" box="[761,952,1114,1135]" italics="true" pageId="7" pageNumber="408">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0998A49B24793B5B2C" bold="true" box="[957,1037,1114,1135]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE0998A49B24793B5B2C" box="[957,1037,1114,1135]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(LACM 3553). A. Rachidian and inner lateral radular teeth. B. Lateral radular teeth. C. Outer radular teeth. D. Jaw.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BA836DAFFA1DE099BDE9BC479CE5954" blockId="7.[151,1437,1210,1955]" pageId="7" pageNumber="408">
|
|||
|
|
The only two other deep-water species of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999B99BC27FCF5B90" ID-CoL="7PJZ6" authorityName="Cuvier" authorityYear="1797" box="[672,761,1212,1235]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999B99BC27FCF5B90" box="[672,761,1212,1235]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(found below
|
|||
|
|
<quantity id="4CEF9B3FFFA1DE0998869BC47ED15B91" box="[927,999,1210,1235]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="408" unit="m" value="500.0">500 m</quantity>
|
|||
|
|
depth) known to date are
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099E149BC27C175BB4" ID-CoL="5952V" authority="Bergh, 1899" authorityName="Bergh" authorityYear="1899" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="ingolfiana">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E149BC278AB5B90" box="[1293,1437,1210,1235]" italics="true" pageId="7" pageNumber="408">T. ingolfiana</emphasis>
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE099B8E9BA07C175BB4" author="Bergh, R." box="[151,289,1246,1271]" pageId="7" pageNumber="408" pagination="1 - 46" refId="ref13401" refString="Bergh, R. (1899) Nudibranchiate Gasteropoda. In: The Danish Ingolf-Expedition. Bianco Luno, Copenhagen, 2 (3), 1 - 46." type="journal article" year="1899">Bergh, 1899</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
described by
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE099AA49BA07F645BB4" author="Bergh, R." box="[445,594,1246,1271]" pageId="7" pageNumber="408" pagination="1 - 46" refId="ref13401" refString="Bergh, R. (1899) Nudibranchiate Gasteropoda. In: The Danish Ingolf-Expedition. Bianco Luno, Copenhagen, 2 (3), 1 - 46." type="journal article" year="1899">Bergh (1899)</bibRefCitation>
|
|||
|
|
from the mid-Atlantic ridge based on a single specimen collected at
|
|||
|
|
<quantity id="4CEF9B3FFFA1DE099E4C9BA178AA5BB4" box="[1365,1436,1247,1271]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="7" pageNumber="408" unit="m" value="887.0">887 m</quantity>
|
|||
|
|
depth, and
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099A019A7A7F535A58" ID-CoL="59525" authority="Bouchet, 1977" authorityName="Bouchet" authorityYear="1977" box="[280,613,1282,1307]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="episcopalis">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A019A7A7C835A58" box="[280,437,1282,1307]" italics="true" pageId="7" pageNumber="408">T. episcopalis</emphasis>
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE099AA69A7C7F535A58" author="Bouchet, P." box="[447,613,1282,1307]" pageId="7" pageNumber="408" pagination="28 - 66" refId="ref13573" refString="Bouchet, P. (1977) Opisthobranches de profondeur de l'Ocean Atlantique: II-Notaspidea et Nudibranchiata. Journal of Molluscan Studies, 43, 28 - 66." type="journal article" year="1977">Bouchet, 1977</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
described from
|
|||
|
|
<quantity id="4CEF9B3FFFA1DE09983C9A7C7EF55A59" box="[805,963,1282,1307]" metricMagnitude="3" metricUnit="m" metricValue="1.6024999999999998" metricValueMax="2.17" metricValueMin="1.035" pageId="7" pageNumber="408" unit="m" value="1602.5" valueMax="2170.0" valueMin="1035.0">1035–2170 m</quantity>
|
|||
|
|
depth also in the Atlantic.
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE099E1A9A7C78AA5A58" author="Bergh, R." box="[1283,1436,1282,1307]" pageId="7" pageNumber="408" pagination="1 - 46" refId="ref13401" refString="Bergh, R. (1899) Nudibranchiate Gasteropoda. In: The Danish Ingolf-Expedition. Bianco Luno, Copenhagen, 2 (3), 1 - 46." type="journal article" year="1899">Bergh (1899)</bibRefCitation>
|
|||
|
|
described his preserved specimen
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999029A567F9D5A7C" ID-CoL="5952V" authorityName="Bergh" authorityYear="1899" box="[539,683,1318,1343]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="ingolfiana">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999029A567F9D5A7C" box="[539,683,1318,1343]" italics="true" pageId="7" pageNumber="408">T. ingolfiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as grayish blue with yellow rhinophores. This is different from
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099E9E9A567C3F5A20" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E9E9A567C3F5A20" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A099A347C5F5A20" bold="true" box="[272,361,1354,1379]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099A099A347C5F5A20" box="[272,361,1354,1379]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, which has a black to white body and the rhinophores, gills and velum are much lighter even in the preserved specimens. Other differences include the shape of the penis, which is very elongate and tightly packed in the penial sheath of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099AEA9AEA7FB55AE8" ID-CoL="5952V" authorityName="Bergh" authorityYear="1899" box="[499,643,1426,1451]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="ingolfiana">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099AEA9AEA7FB55AE8" box="[499,643,1426,1451]" italics="true" pageId="7" pageNumber="408">T. ingolfiana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE09998B9AEC7E945AE8" author="Bergh, R." box="[658,930,1426,1451]" pageId="7" pageNumber="408" pagination="1 - 46" refId="ref13401" refString="Bergh, R. (1899) Nudibranchiate Gasteropoda. In: The Danish Ingolf-Expedition. Bianco Luno, Copenhagen, 2 (3), 1 - 46." type="journal article" year="1899">Bergh 1899: pl. 3 fig. 9</bibRefCitation>
|
|||
|
|
), whereas the penis of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099FB39AEA780E5AE8" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1194,1336,1428,1451]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FB39AEA780E5AE8" box="[1194,1336,1428,1451]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E589AEC78B75AE8" bold="true" box="[1345,1409,1426,1451]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099E589AEC78B75AE8" box="[1345,1409,1426,1451]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is much shorter and straight.
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099AD59AC67F985A8C" ID-CoL="59525" authorityName="Bouchet" authorityYear="1977" box="[460,686,1462,1487]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="episcopalis">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099AD59AC67F985A8C" box="[460,686,1462,1487]" italics="true" pageId="7" pageNumber="408">Tritonia episcopalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as well as
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE09982D9AC679255A8C" ID-CoL="5953N" box="[820,1043,1462,1487]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="newfoundlandica">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE09982D9AC679255A8C" box="[820,1043,1462,1487]" italics="true" pageId="7" pageNumber="408">T. newfoundlandica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have a characteristic scoop in the penis, which is absent in
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099AAA9AA27F765AB0" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[435,576,1500,1523]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099AAA9AA27F765AB0" box="[435,576,1500,1523]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999519AA47FB15AB0" bold="true" box="[584,647,1498,1523]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE0999519AA47FB15AB0" box="[584,647,1498,1523]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
The external coloration of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0998A29AA279635AB0" ID-CoL="59525" authorityName="Bouchet" authorityYear="1977" box="[955,1109,1498,1523]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="episcopalis">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0998A29AA279635AB0" box="[955,1109,1498,1523]" italics="true" pageId="7" pageNumber="408">T. episcopalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is bright violet with the gills of the same color and the rhinophores red, whereas in
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999EF997E7EB55954" ID-CoL="7TFDV" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[758,899,1536,1559]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999EF997E7EB55954" box="[758,899,1536,1559]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0998939A807EFF5954" bold="true" box="[906,969,1534,1559]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE0998939A807EFF5954" box="[906,969,1534,1559]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
the body is black to white.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30D6551FFA1DE099BDE995D7DCE58E0" pageId="7" pageNumber="408" type="description">
|
|||
|
|
<paragraph id="8BA836DAFFA1DE099BDE995D7DCE58E0" blockId="7.[151,1437,1210,1955]" pageId="7" pageNumber="408">
|
|||
|
|
The sister species of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099AA7995A7FA35978" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[446,661,1572,1595]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099AA7995A7FA35978" box="[446,661,1572,1595]" italics="true" pageId="7" pageNumber="408">Tritonia nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099987995D7FD6597F" bold="true" box="[670,736,1571,1596]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099987995D7FD6597F" box="[670,736,1571,1596]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999EB995D7E9C597F" box="[754,938,1571,1596]" italics="true" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999EB995D7E90597F" authorityName="Eliot" authorityYear="1907" box="[754,934,1571,1596]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="belli">Tritoniella belli</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
is very distinct externally.
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099FF3995D78AA597F" authorityName="Eliot" authorityYear="1907" box="[1258,1436,1571,1596]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="belli">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FF3995D78AA597F" box="[1258,1436,1571,1596]" italics="true" pageId="7" pageNumber="408">Tritoniella belli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
ranges in color from uniform milky-white or translucent to yellow or orange in specimens (
|
|||
|
|
<bibRefCitation id="EF864B2BFFA1DE099F9599387802591C" author="Wagele, H." box="[1164,1332,1606,1631]" pageId="7" pageNumber="408" pagination="235 - 243" refId="ref17461" refString="Wagele, H. (1989 b) On the anatomy and zoogeography of Tritoniella belli Eliot, 1907 (Opisthobranchia, Nudibranchia) and the synonymy of T. sinuata Eliot, 1907. Polar Biology, 9, 235 - 243. https: // doi. org / 10.1007 / BF 00263771" type="journal article" year="1989">Wägele 1989b</bibRefCitation>
|
|||
|
|
). As in
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099E9E99367C3F59C0" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E9E99367C3F59C0" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A1699157C7959C7" bold="true" box="[271,335,1643,1668]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099A1699157C7959C7" box="[271,335,1643,1668]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
the mantle edge is wavy and can be smooth or have finger-like processes. However, specimens of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099B8E99EE7DD159E4" authorityName="Eliot" authorityYear="1907" box="[151,231,1678,1703]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="belli">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099B8E99EE7DD159E4" box="[151,231,1678,1703]" italics="true" pageId="7" pageNumber="408">T. belli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have a smooth dorsum typically with a longitudinal median ridge, whereas
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099F6B99EE783259E4" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1138,1284,1680,1703]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099F6B99EE783259E4" box="[1138,1284,1680,1703]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E1699F0786559E4" bold="true" box="[1295,1363,1678,1703]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099E1699F0786559E4" box="[1295,1363,1678,1703]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
has a tuberculate dorsum and no ridge. In the phylogenetic analyses presented here, both
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099F5A99CA79E65988" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1091,1232,1716,1739]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099F5A99CA79E65988" box="[1091,1232,1716,1739]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FCE99CD7821598F" bold="true" box="[1239,1303,1715,1740]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099FCE99CD7821598F" box="[1239,1303,1715,1740]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099E5799CA78AC598F" authorityName="Eliot" authorityYear="1907" box="[1358,1434,1715,1740]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="belli">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E5799CA78AC598F" box="[1358,1434,1715,1740]" italics="true" pageId="7" pageNumber="408">T. belli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
form a monophyletic group sister to
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE09995899A67FAC59AC" authorityName="Cuvier" authorityYear="1797" box="[577,666,1752,1775]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE09995899A67FAC59AC" box="[577,666,1752,1775]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, supporting the separation of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0998E199A8795859AC" authorityName="Eliot" authorityYear="1907" box="[1016,1134,1750,1775]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0998E199A8795859AC" box="[1016,1134,1750,1775]" italics="true" pageId="7" pageNumber="408">Tritoniella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099FA199A6782459AC" authorityName="Cuvier" authorityYear="1797" box="[1208,1298,1752,1775]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FA199A6782459AC" box="[1208,1298,1752,1775]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Under this scenario,
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099A1A99827CA65850" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[259,400,1788,1811]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A1A99827CA65850" box="[259,400,1788,1811]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A8E99857CE15857" bold="true" box="[407,471,1787,1812]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099A8E99857CE15857" box="[407,471,1787,1812]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
should be considered a member of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE09987099857EE95857" authorityName="Eliot" authorityYear="1907" box="[873,991,1787,1812]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE09987099857EE95857" box="[873,991,1787,1812]" italics="true" pageId="7" pageNumber="408">Tritoniella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. However, the phylogeny of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099E3E998278B45850" authorityName="Cuvier" authorityYear="1797" box="[1319,1410,1788,1811]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E3E998278B45850" box="[1319,1410,1788,1811]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is poorly understood and several critical species have not yet been sequenced. Also, morphologically,
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099FE8985E78485874" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1265,1406,1824,1847]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FE8985E78485874" box="[1265,1406,1824,1847]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099E9C98607D8E581F" pageId="7" pageNumber="408" rank="species">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E9C98607D8E581F" bold="true" pageId="7" pageNumber="408">n. sp.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
appears more similar to some species of
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE0999AB983A7E3A5818" authorityName="Cuvier" authorityYear="1797" box="[690,780,1860,1883]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE0999AB983A7E3A5818" box="[690,780,1860,1883]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
than to
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE09986E983A7EFC581F" authorityName="Eliot" authorityYear="1907" box="[887,970,1859,1884]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="belli">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE09986E983A7EFC581F" box="[887,970,1859,1884]" italics="true" pageId="7" pageNumber="408">T. belli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
making difficult to find morphological synapomorphies for
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099A9A98187CCF583C" authorityName="Eliot" authorityYear="1907" box="[387,505,1894,1919]" class="Gastropoda" family="Tritoniidae" genus="Tritoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099A9A98187CCF583C" box="[387,505,1894,1919]" italics="true" pageId="7" pageNumber="408">Tritoniella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Until all these issues are resolved, we prefer to maintain
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099FB898167807583C" authority="Valdés & Lundsten & Wilson, 2018" authorityName="Valdés & Lundsten & Wilson" authorityYear="2018" box="[1185,1329,1896,1919]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="species" species="nigritigris" status="sp. nov.">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099FB898167807583C" box="[1185,1329,1896,1919]" italics="true" pageId="7" pageNumber="408">T. nigritigris</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099E229818784A583C" bold="true" box="[1339,1404,1894,1919]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="A25057B3FFA1DE099E229818784A583C" box="[1339,1404,1894,1919]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
in
|
|||
|
|
<taxonomicName id="4C174D59FFA1DE099B8E98F27DC658E0" authorityName="Cuvier" authorityYear="1797" box="[151,240,1932,1955]" class="Gastropoda" family="Tritoniidae" genus="Tritonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="408" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B963EAC8FFA1DE099B8E98F27DC658E0" box="[151,240,1932,1955]" italics="true" pageId="7" pageNumber="408">Tritonia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|