treatments-xml/data/03/CF/87/03CF87C0FFF3FFB7FD60996C8F68FC27.xml

<document id="C7BFD07AB394C2E7CBE1D69890B9EB26" ID-DOI="10.1163/18759866-bja10018" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343383" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1694664001284" checkinUser="felipe" docAuthor="Cooper, David, Yamaguchi, Nobuyuki, Macdonald, David, Nanova, Olga, Yudin, Viktor, Dugmore, Andrew &amp; Kitchener, Andrew" docDate="2021" docId="03CF87C0FFF3FFB7FD60996C8F68FC27" docLanguage="en" docName="ContribZool.90.3.285-317.pdf" docOrigin="Contributions to Zoology 90 (1)" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Zebrus pallaoroi Kovacic, Sanda &amp; Vukic 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="300" masterDocId="FFF6FFB8FFFBFFA7FFC498328E50FFFC" masterDocTitle="Zebrus pallaoroi sp. nov.: a new species of goby (Actinopterygii: Gobiidae) from the Mediterranean Sea with a DNA-based phylogenetic analysis of the Gobius-lineage" masterLastPageNumber="317" masterPageNumber="285" pageNumber="293" updateTime="1695057095745" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FF2A338436F0F7F008E44529F4FFBC0A">Zebrus pallaoroi sp. nov.: a new species of goby (Actinopterygii: Gobiidae) from the Mediterranean Sea with a DNA-based phylogenetic analysis of the Gobius-lineage</mods:title>
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<taxonomicName id="4C664D55FFF3FFAFFD60996C8A23FE85" authority="Kovacic, Sanda &amp; Vukic" authorityName="Kovacic, Sanda &amp; Vukic" authorityYear="2021" box="[676,1139,350,384]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="293" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
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<emphasis id="B912EAC4FFF3FFAFFD60996C8D30FE85" bold="true" box="[676,864,350,377]" italics="true" pageId="8" pageNumber="293">Zebrus pallaoroi</emphasis>
Kovačić, Šanda &amp; Vukić
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<taxonomicNameLabel id="A22157BFFFF3FFAFFD6099B18CA8FE61" box="[676,760,387,413]" pageId="8" pageNumber="293">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BD936D6FFF3FFAFFD6099958ACBFDD1" blockId="8.[676,1188,422,1742]" pageId="8" pageNumber="293">
<materialsCitation id="3B0E3C8BFFF3FFAFFD6099958ACBFDD1" collectingDate="2016-08-24" collectionCode="R" collectorName="Sanda" country="Serbia" county="Adriatic Sea" latitude="42.485138" location="Male" longLatPrecision="1" longitude="18.670347" municipality="Boka Kotorska" pageId="8" pageNumber="293" specimenCode="V 144302" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54DD8874FFF3FFAFFD6099958D55FE3D" box="[676,773,423,449]" pageId="8" pageNumber="293">Holotype</typeStatus>
(
</emphasis>
<figureCitation id="135D2A53FFF3FFAFFCDB99958D3DFE3D" box="[799,877,423,449]" captionStart="figure 1" captionStartId="10.[128,195,1556,1578]" captionTargetBox="[223,1092,575,1537]" captionTargetId="figure-3@10.[223,1092,575,1538]" captionTargetPageId="10" captionText="figure 1 Preserved specimens.(A) Zebrus pallaoroi sp.nov., npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M.Kovačić. (B) Zebrus zebrus, nmp P6V 140912,neotype, female, 23.25 + 6.22mm, Îll Gross, Banyuls sur Mer, France (C) Millerigobius macrocephalus, nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France." figureDoi="http://doi.org/10.5281/zenodo.8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="8" pageNumber="293">fig. 1A</figureCitation>
<emphasis id="B912EAC4FFF3FFAFFCA999958D2CFE3D" box="[877,892,423,449]" italics="true" pageId="8" pageNumber="293">).</emphasis>
<location id="8EB9600DFFF3FFAFFC4B99958D99FE3D" LSID="urn:lsid:plazi:treatment:03CF87C0FFF3FFB7FD60996C8F68FC27:8EB9600DFFF3FFAFFC4B99958D99FE3D" box="[911,969,423,449]" country="Serbia" county="Adriatic Sea" latitude="42.485138" longLatPrecision="1" longitude="18.670347" municipality="Boka Kotorska" name="Male" pageId="8" pageNumber="293">Male</location>
, 31.81 + 
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, nmp P6 
<specimenCode id="DBC09EADFFF3FFAFFD3999F88D36FE19" box="[765,870,458,485]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="293" type="Museum">V 144302</specimenCode>
, 
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, 
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, 
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, 
<collectingCountry id="F3717646FFF3FFAFFC8C99DD8D83FDF5" box="[840,979,495,521]" name="Serbia" pageId="8" pageNumber="293">Montenegro</collectingCountry>
, 
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<collectingDate id="EF9CE9FEFFF3FFAFFC2299DD8ACFFDF4" box="[998,1183,494,521]" pageId="8" pageNumber="293" value="2016-08-24">24 August 2016</collectingDate>
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, 
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, 
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, collector 
<collectorName id="26935300FFF3FFAFFBFE9A278A2EFDC8" box="[1082,1150,533,564]" pageId="8" pageNumber="293">Šanda</collectorName>
<collectionCode id="ED77AE13FFF3FFAFFB409A218AC7FDD1" box="[1156,1175,531,557]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="293">R</collectionCode>
.
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<materialsCitation id="3B0E3C8BFFF3FFAFFD609A058A01FD64" collectingDate="2016-08-24" collectionCode="V" collectorName="Sanda" country="Serbia" county="Adriatic Sea" latitude="42.485138" location="Male" longLatPrecision="1" longitude="18.670347" municipality="Boka Kotorska" pageId="8" pageNumber="293" specimenCode="V 144300, V 144303" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54DD8874FFF3FFAFFD609A058D42FDAC" box="[676,786,567,592]" pageId="8" pageNumber="293">Paratypes</typeStatus>
.
</emphasis>
Male, 27.72 + 
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, nmp P6 
<specimenCode id="DBC09EADFFF3FFAFFB549A048CA3FD88" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="293" type="Museum">V 144300</specimenCode>
, and female, 26.44 + 
<quantity id="4C9E9B33FFF3FFAFFC389A688A0DFD89" box="[1020,1117,602,629]" metricMagnitude="-3" metricUnit="m" metricValue="6.39" pageId="8" pageNumber="293" unit="mm" value="6.39">6.39 mm</quantity>
, nmp P6 
<specimenCode id="DBC09EADFFF3FFAFFD7A9A4C8D73FD64" box="[702,803,638,664]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="293" type="Museum">V 144303</specimenCode>
, the same data as holotype
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.
</paragraph>
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<emphasis id="B912EAC4FFF3FFAFFD069A918A56FD40" box="[706,1030,674,700]" italics="true" pageId="8" pageNumber="293">
Diagnosis. 
<taxonomicName id="4C664D55FFF3FFAFFC819A908A56FD40" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[837,1030,674,700]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="293" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">Zebrus pallaoroi</taxonomicName>
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differs from the only congeneric species, 
<taxonomicName id="4C664D55FFF3FFAFFB499AF58CBFFCF8" authorityName="Risso" authorityYear="1827" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="293" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF3FFAFFB499AF58CBFFCF8" italics="true" pageId="8" pageNumber="293">Z. zebrus</emphasis>
</taxonomicName>
by each of the following characters: (1) snout longer than eye, its length 1.1‒1.2 of eye diameter; (2) posterior nostril short tube, about 0.8–0.9 of anterior nostril (
<figureCitation id="135D2A53FFF3FFAFFBBB9B678C93FC68" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="8" pageNumber="293">fig. 2A</figureCitation>
); (3) eyes moderately small, eye diameter is 
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head length; (4) left and right ventrolateral head ridges transversally connected on anterior part by short transversal ridge (
<figureCitation id="135D2A53FFF3FFAFFD309C3B8D1AFBDF" box="[756,842,1033,1059]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="8" pageNumber="293">fig. 2C</figureCitation>
), (5) anterior membrane in midline depth about 2/3 of spinous ray (
<figureCitation id="135D2A53FFF3FFAFFBB99C1F8C95FB97" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="8" pageNumber="293">fig. 2B</figureCitation>
); (6) head canal pores large, pore 
<emphasis id="B912EAC4FFF3FFAFFB8F9C608A08FB97" box="[1099,1112,1106,1131]" italics="true" pageId="8" pageNumber="293">α</emphasis>
diameter about half of the distance between pore 
<emphasis id="B912EAC4FFF3FFAFFD1A9CAB8CB9FB4E" box="[734,745,1177,1202]" italics="true" pageId="8" pageNumber="293">Ρ</emphasis>
and 
<emphasis id="B912EAC4FFF3FFAFFCE69CAB8D65FB4F" box="[802,821,1177,1203]" italics="true" pageId="8" pageNumber="293">Ρ1</emphasis>
; (7) suborbital sensory papillae row 
<emphasis id="B912EAC4FFF3FFAFFD139C8F8CBAFB2B" box="[727,746,1213,1239]" italics="true" pageId="8" pageNumber="293">5i</emphasis>
going downwards to or near the level of row 
<emphasis id="B912EAC4FFF3FFAFFD319CD28D53FB06" box="[757,771,1248,1274]" italics="true" pageId="8" pageNumber="293">d</emphasis>
, distance between row 
<emphasis id="B912EAC4FFF3FFAFFBD29CD38A79FB07" box="[1046,1065,1249,1275]" italics="true" pageId="8" pageNumber="293">5i</emphasis>
and row 
<emphasis id="B912EAC4FFF3FFAFFB509CD28AF2FB06" box="[1172,1186,1248,1274]" italics="true" pageId="8" pageNumber="293">d</emphasis>
absent or much smaller than length of row 
<emphasis id="B912EAC4FFF3FFAFFD609D1B8CE7FABF" box="[676,695,1321,1347]" italics="true" pageId="8" pageNumber="293">5i</emphasis>
; (8) body with ten to eleven vertical dark brown bands present along lateral side, first in front of the first dorsal fin, last at end of the second dorsal fin, at upper edge about equal or narrower than pale interspaces inbetween.
</paragraph>
<paragraph id="8BD936D6FFF3FFABFD069E328C27FB73" blockId="8.[676,1188,422,1742]" lastBlockId="12.[129,640,171,1742]" lastPageId="12" lastPageNumber="297" pageId="8" pageNumber="293">
<emphasis id="B912EAC4FFF3FFAFFD069E328D10F9E5" box="[706,832,1536,1561]" italics="true" pageId="8" pageNumber="293">Description</emphasis>
(all morphometric values and meristics in the text are presented as 
<typeStatus id="54DD8874FFF3FFAFFBA39E168C85F99E" pageId="8" pageNumber="293">holotype</typeStatus>
first and 
<typeStatus id="54DD8874FFF3FFAFFCFA9E7A8DFEF99E" box="[830,942,1608,1634]" pageId="8" pageNumber="293">paratypes</typeStatus>
, if different, in parentheses; first the male 
<typeStatus id="54DD8874FFF3FFAFFC5D9E5E8DAEF97A" box="[921,1022,1644,1670]" pageId="8" pageNumber="293">paratype</typeStatus>
, then female). General morphology (
<figureCitation id="135D2A53FFF3FFAFFC619EBD8DA2F955" box="[933,1010,1679,1705]" captionStart="figure 1" captionStartId="10.[128,195,1556,1578]" captionTargetBox="[223,1092,575,1537]" captionTargetId="figure-3@10.[223,1092,575,1538]" captionTargetPageId="10" captionText="figure 1 Preserved specimens.(A) Zebrus pallaoroi sp.nov., npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M.Kovačić. (B) Zebrus zebrus, nmp P6V 140912,neotype, female, 23.25 + 6.22mm, Îll Gross, Banyuls sur Mer, France (C) Millerigobius macrocephalus, nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France." figureDoi="http://doi.org/10.5281/zenodo.8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="8" pageNumber="293">fig. 1A</figureCitation>
): Body proportions are given in 
<tableCitation id="C6E4036DFFF3FFAFFCBC9E818D99F931" box="[888,969,1715,1741]" captionStart="table 2" captionStartId="9.[98,158,165,187]" captionTargetPageId="9" captionText="table 2 Morphometric characters (as proportional measurements in %) of Zebrus pallaoroi sp.nov." httpUri="http://table.plazi.org/id/DF19665EFFF2FFAEFFA698978A4BFF47" pageId="8" pageNumber="293" tableUuid="DF19665EFFF2FFAEFFA698978A4BFF47">table 2</tableCitation>
. Body elongate, its depth at pelvic-fin origin 5.3 (5.4) in sl, at anal-fin origin 5.5 (5.9) in sl, laterally compressed posteriorly, with deep caudal peduncle, caudal peduncle depth 0.6 of caudal peduncle length (
<figureCitation id="135D2A53FFF0FFACFEF799098FD2FEA9" box="[307,386,315,341]" captionStart="figure 1" captionStartId="10.[128,195,1556,1578]" captionTargetBox="[223,1092,575,1537]" captionTargetId="figure-3@10.[223,1092,575,1538]" captionTargetPageId="10" captionText="figure 1 Preserved specimens.(A) Zebrus pallaoroi sp.nov., npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M.Kovačić. (B) Zebrus zebrus, nmp P6V 140912,neotype, female, 23.25 + 6.22mm, Îll Gross, Banyuls sur Mer, France (C) Millerigobius macrocephalus, nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France." figureDoi="http://doi.org/10.5281/zenodo.8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="11" pageNumber="296">fig. 1A</figureCitation>
). Head moderately large, the length 3.5 (3.2 and 3.4) in sl, width 3.9 (3.8 and 4.2) in sl, depth 6.4 (5.7 and 6.9) in sl, and depressed, its depth 1.6 (1.5 and 1.6) in width. Snout oblique, with a convex ridge along dorsal midline visible only in 
<typeStatus id="54DD8874FFF0FFACFBD5989E8AD0FF3A" box="[1041,1152,172,198]" pageId="11" pageNumber="296">paratypes</typeStatus>
, longer than eye, its length 1.2 (1.1 and 1.2) of eye diameter, 3.7 (4.2 and 3.8) in head length. Anterior nostril nasal tube with a single process from the posterior rim, posterior nostril short tube, 81% (85 and 88%) of the anterior nostril (
<figureCitation id="135D2A53FFF0FFACFD2199B18D6BFE61" box="[741,827,387,413]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="11" pageNumber="296">fig. 2A</figureCitation>
). Eyes dorsolateral, moderately small, eye diameter is 4.3 (4.7 and 4.6) in head length, slightly elevated above the dorsal profile. Interorbital moderately narrow, 1/2 of eye diameter. Cranial roof covered by dorsal axial musculature. Predorsal area about horizontal. Mouth oblique, jaws ending anteriorly nearly equally to lower lip. Upper lip broad, width nearly uniform, more or less as wide as the lateral preorbital area in female, wider in males. Mouth moderately large, posterior angle of jaws ending posteriorly below mideye. Cheek relatively narrow. Teeth in both jaws erect, caniniform; upper jaw with enlarged outer row, intermediate irregularly scattered small teeth and four enlarged median posterior teeth; lower jaw with an outer row of larger teeth, intermediate irregularly scattered small teeth and a short inner row of enlarged teeth laterally ending with one or two distinct canine teeth on each side. Tongue reduced. Chin with a small mental fold. Left and right ventrolateral head ridges transversally connected on the anterior part by a short transversal ridge (
<figureCitation id="135D2A53FFF7FFABFF4C9BF38E85FC27" box="[136,213,961,987]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="12" pageNumber="297">fig. 2C</figureCitation>
). Branchiostegal membranes fused to isthmus along the entire lateral margin of the isthmus, from immediately anterior to pectoral margin, gill openings restricted to pectoral-fin base. No spines on preopercle. Pectoral girdle without dermal flaps on anterior edge.
</paragraph>
<caption id="DF19665EFFF2FFAEFFA698978A4BFF47" ID-Table-UUID="DF19665EFFF2FFAEFFA698978A4BFF47" box="[98,1051,165,188]" httpUri="http://table.plazi.org/id/DF19665EFFF2FFAEFFA698978A4BFF47" pageId="9" pageNumber="294" startId="9.[98,158,165,187]" targetBox="[104,1149,208,1657]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="8BD936D6FFF2FFAEFFA698978A4BFF47" blockId="9.[98,1051,165,188]" box="[98,1051,165,188]" pageId="9" pageNumber="294">
table 2 Morphometric characters (as proportional measurements in %) of 
<taxonomicName id="4C664D55FFF2FFAEFCFB98978D82FF46" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[831,978,165,186]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="294" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF2FFAEFCFB98978D82FF46" box="[831,978,165,186]" italics="true" pageId="9" pageNumber="294">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF2FFAEFC1398948A4BFF47" box="[983,1051,166,187]" pageId="9" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD936D6FFF2FFAEFFAC98E28A7EF985" pageId="9" pageNumber="294">
<table id="F966C476FFF20058FFAC98E28A2DF985" box="[104,1149,208,1657]" colsContinueIn="10.[134,1179,208,457]" gridcols="4" gridrows="44" pageId="9" pageNumber="294">
<tr id="35563494FFF20058FFAC98E28A2DFF16" box="[104,1149,208,234]" gridrow="0" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC98E28FA3FF16" box="[104,499,208,234]" gridcol="0" gridrow="0" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFFAC98E28E8BFF16" bold="true" box="[104,219,208,234]" pageId="9" pageNumber="294">Specimen</emphasis>
</th>
<th id="76875DE8FFF20058FDF698E28CE2FF16" box="[562,690,208,234]" gridcol="1" gridrow="0" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFDF698E28CE2FF16" bold="true" box="[562,690,208,234]" pageId="9" pageNumber="294">P6V 144302</emphasis>
</th>
<th id="76875DE8FFF20058FCE198E28DF6FF16" box="[805,934,208,234]" gridcol="2" gridrow="0" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFCE198E28DF6FF16" bold="true" box="[805,934,208,234]" pageId="9" pageNumber="294">P6V 144300</emphasis>
</th>
<th id="76875DE8FFF20058FC3998E28A2DFF16" box="[1021,1149,208,234]" gridcol="3" gridrow="0" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFC3998E28A2DFF16" bold="true" box="[1021,1149,208,234]" pageId="9" pageNumber="294">P6V 144303</emphasis>
</th>
</tr>
<tr id="35563494FFF20058FFAC98C68A2DFEF2" box="[104,1149,244,270]" gridrow="1" pageId="9" pageNumber="294" rowspan-0="1">
<td id="76875DE8FFF20058FDF698C68CE2FEF2" box="[562,690,244,270]" gridcol="1" gridrow="1" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFDF698C68CC9FEF2" bold="true" box="[562,665,244,270]" pageId="9" pageNumber="294">holotype</emphasis>
</td>
<td id="76875DE8FFF20058FCE198C68DF6FEF2" box="[805,934,244,270]" gridcol="2" gridrow="1" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFCE198C68DDCFEF2" bold="true" box="[805,908,244,270]" pageId="9" pageNumber="294">paratype</emphasis>
</td>
<td id="76875DE8FFF20058FC3998C68A2DFEF2" box="[1021,1149,244,270]" gridcol="3" gridrow="1" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFC3998C68A34FEF2" bold="true" box="[1021,1124,244,270]" pageId="9" pageNumber="294">paratype</emphasis>
</td>
</tr>
<tr id="35563494FFF20058FFAC99168A2DFEC3" box="[104,1149,292,319]" gridrow="2" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99168FA3FEC3" box="[104,499,292,319]" gridcol="0" gridrow="2" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFFAC99168EC0FEC2" bold="true" box="[104,144,292,318]" pageId="9" pageNumber="294">Sex</emphasis>
</th>
<td id="76875DE8FFF20058FDF699168CE2FEC3" box="[562,690,292,319]" gridcol="1" gridrow="2" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFDF699178C3BFEC3" bold="true" box="[562,619,293,319]" pageId="9" pageNumber="294">male</emphasis>
</td>
<td id="76875DE8FFF20058FCE199168DF6FEC3" box="[805,934,292,319]" gridcol="2" gridrow="2" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFCE199178D0EFEC3" bold="true" box="[805,862,293,319]" pageId="9" pageNumber="294">male</emphasis>
</td>
<td id="76875DE8FFF20058FC3999168A2DFEC3" box="[1021,1149,292,319]" gridcol="3" gridrow="2" pageId="9" pageNumber="294">
<emphasis id="B912EAC4FFF2FFAEFC3999168A1CFEC2" bold="true" box="[1021,1100,292,318]" pageId="9" pageNumber="294">female</emphasis>
</td>
</tr>
<tr id="35563494FFF20058FFAC99678A2DFE93" box="[104,1149,341,367]" gridrow="3" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99678FA3FE93" box="[104,499,341,367]" gridcol="0" gridrow="3" pageId="9" pageNumber="294">Standard length (sl) in mm</th>
<td id="76875DE8FFF20058FDF699678CE2FE93" box="[562,690,341,367]" gridcol="1" gridrow="3" pageId="9" pageNumber="294">31.81</td>
<td id="76875DE8FFF20058FCE199678DF6FE93" box="[805,934,341,367]" gridcol="2" gridrow="3" pageId="9" pageNumber="294">27.72</td>
<td id="76875DE8FFF20058FC3999678A2DFE93" box="[1021,1149,341,367]" gridcol="3" gridrow="3" pageId="9" pageNumber="294">26.44</td>
</tr>
<tr id="35563494FFF20058FFAC99478A2DFE73" box="[104,1149,373,399]" gridrow="4" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99478A2DFE73" box="[104,1149,373,399]" colspan="4" colspanRight="3" gridcol="0" gridrow="4" pageId="9" pageNumber="294">% of standard length</th>
</tr>
<tr id="35563494FFF20058FFAC99A78A2DFE53" box="[104,1149,405,431]" gridrow="5" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99A78FA3FE53" box="[104,499,405,431]" gridcol="0" gridrow="5" pageId="9" pageNumber="294">1st spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF699A78CE2FE53" box="[562,690,405,431]" gridcol="1" gridrow="5" pageId="9" pageNumber="294">10.0</td>
<td id="76875DE8FFF20058FCE199A78DF6FE53" box="[805,934,405,431]" gridcol="2" gridrow="5" pageId="9" pageNumber="294">12.6</td>
<td id="76875DE8FFF20058FC3999A78A2DFE53" box="[1021,1149,405,431]" gridcol="3" gridrow="5" pageId="9" pageNumber="294">12.2</td>
</tr>
<tr id="35563494FFF20058FFAC99878A2DFE2C" box="[104,1149,437,464]" gridrow="6" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99878FA3FE2C" box="[104,499,437,464]" gridcol="0" gridrow="6" pageId="9" pageNumber="294">2nd spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF699878CE2FE2C" box="[562,690,437,464]" gridcol="1" gridrow="6" pageId="9" pageNumber="294">11.8</td>
<td id="76875DE8FFF20058FCE199878DF6FE2C" box="[805,934,437,464]" gridcol="2" gridrow="6" pageId="9" pageNumber="294">13.5</td>
<td id="76875DE8FFF20058FC3999878A2DFE2C" box="[1021,1149,437,464]" gridcol="3" gridrow="6" pageId="9" pageNumber="294">12.7</td>
</tr>
<tr id="35563494FFF20058FFAC99E48A2DFE0C" box="[104,1149,470,496]" gridrow="7" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99E48FA3FE0C" box="[104,499,470,496]" gridcol="0" gridrow="7" pageId="9" pageNumber="294">3rd spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF699E48CE2FE0C" box="[562,690,470,496]" gridcol="1" gridrow="7" pageId="9" pageNumber="294">12.7</td>
<td id="76875DE8FFF20058FCE199E48DF6FE0C" box="[805,934,470,496]" gridcol="2" gridrow="7" pageId="9" pageNumber="294">14.4</td>
<td id="76875DE8FFF20058FC3999E48A2DFE0C" box="[1021,1149,470,496]" gridcol="3" gridrow="7" pageId="9" pageNumber="294">14.3</td>
</tr>
<tr id="35563494FFF20058FFAC99C48A2DFDEC" box="[104,1149,502,528]" gridrow="8" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC99C48FA3FDEC" box="[104,499,502,528]" gridcol="0" gridrow="8" pageId="9" pageNumber="294">4th spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF699C48CE2FDEC" box="[562,690,502,528]" gridcol="1" gridrow="8" pageId="9" pageNumber="294">12.2</td>
<td id="76875DE8FFF20058FCE199C48DF6FDEC" box="[805,934,502,528]" gridcol="2" gridrow="8" pageId="9" pageNumber="294">14.0</td>
<td id="76875DE8FFF20058FC3999C48A2DFDEC" box="[1021,1149,502,528]" gridcol="3" gridrow="8" pageId="9" pageNumber="294">13.3</td>
</tr>
<tr id="35563494FFF20058FFAC9A248A2DFDCD" box="[104,1149,534,561]" gridrow="9" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9A248FA3FDCD" box="[104,499,534,561]" gridcol="0" gridrow="9" pageId="9" pageNumber="294">5th spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF69A248CE2FDCD" box="[562,690,534,561]" gridcol="1" gridrow="9" pageId="9" pageNumber="294">12.1</td>
<td id="76875DE8FFF20058FCE19A248DF6FDCD" box="[805,934,534,561]" gridcol="2" gridrow="9" pageId="9" pageNumber="294">12.8</td>
<td id="76875DE8FFF20058FC399A248A2DFDCD" box="[1021,1149,534,561]" gridcol="3" gridrow="9" pageId="9" pageNumber="294">11.5</td>
</tr>
<tr id="35563494FFF20058FFAC9A048A2DFDAD" box="[104,1149,566,593]" gridrow="10" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9A048FA3FDAD" box="[104,499,566,593]" gridcol="0" gridrow="10" pageId="9" pageNumber="294">6th spine length of first dorsal fin</th>
<td id="76875DE8FFF20058FDF69A048CE2FDAD" box="[562,690,566,593]" gridcol="1" gridrow="10" pageId="9" pageNumber="294">7.9</td>
<td id="76875DE8FFF20058FCE19A048DF6FDAD" box="[805,934,566,593]" gridcol="2" gridrow="10" pageId="9" pageNumber="294">9.0</td>
<td id="76875DE8FFF20058FC399A048A2DFDAD" box="[1021,1149,566,593]" gridcol="3" gridrow="10" pageId="9" pageNumber="294">8.2</td>
</tr>
<tr id="35563494FFF20058FFAC9A658A2DFD8D" box="[104,1149,599,625]" gridrow="11" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9A658FA3FD8D" box="[104,499,599,625]" gridcol="0" gridrow="11" pageId="9" pageNumber="294">Anal fin base</th>
<td id="76875DE8FFF20058FDF69A658CE2FD8D" box="[562,690,599,625]" gridcol="1" gridrow="11" pageId="9" pageNumber="294">19.0</td>
<td id="76875DE8FFF20058FCE19A658DF6FD8D" box="[805,934,599,625]" gridcol="2" gridrow="11" pageId="9" pageNumber="294">19.7</td>
<td id="76875DE8FFF20058FC399A658A2DFD8D" box="[1021,1149,599,625]" gridcol="3" gridrow="11" pageId="9" pageNumber="294">18.8</td>
</tr>
<tr id="35563494FFF20058FFAC9A458A2DFD6E" box="[104,1149,631,658]" gridrow="12" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9A458FA3FD6E" box="[104,499,631,658]" gridcol="0" gridrow="12" pageId="9" pageNumber="294">Anal fin spine length</th>
<td id="76875DE8FFF20058FDF69A458CE2FD6E" box="[562,690,631,658]" gridcol="1" gridrow="12" pageId="9" pageNumber="294">7.7</td>
<td id="76875DE8FFF20058FCE19A458DF6FD6E" box="[805,934,631,658]" gridcol="2" gridrow="12" pageId="9" pageNumber="294">8.2</td>
<td id="76875DE8FFF20058FC399A458A2DFD6E" box="[1021,1149,631,658]" gridcol="3" gridrow="12" pageId="9" pageNumber="294">8.4</td>
</tr>
<tr id="35563494FFF20058FFAC9AA58A2DFD4E" box="[104,1149,663,690]" gridrow="13" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9AA58FA3FD4E" box="[104,499,663,690]" gridcol="0" gridrow="13" pageId="9" pageNumber="294">Body depth at anal fin origin</th>
<td id="76875DE8FFF20058FDF69AA58CE2FD4E" box="[562,690,663,690]" gridcol="1" gridrow="13" pageId="9" pageNumber="294">18.2</td>
<td id="76875DE8FFF20058FCE19AA58DF6FD4E" box="[805,934,663,690]" gridcol="2" gridrow="13" pageId="9" pageNumber="294">16.8</td>
<td id="76875DE8FFF20058FC399AA58A2DFD4E" box="[1021,1149,663,690]" gridcol="3" gridrow="13" pageId="9" pageNumber="294">17.0</td>
</tr>
<tr id="35563494FFF20058FFAC9A858A2DFD2E" box="[104,1149,695,722]" gridrow="14" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9A858FA3FD2E" box="[104,499,695,722]" gridcol="0" gridrow="14" pageId="9" pageNumber="294">Body depth at pelvic fin origin</th>
<td id="76875DE8FFF20058FDF69A858CE2FD2E" box="[562,690,695,722]" gridcol="1" gridrow="14" pageId="9" pageNumber="294">18.9</td>
<td id="76875DE8FFF20058FCE19A858DF6FD2E" box="[805,934,695,722]" gridcol="2" gridrow="14" pageId="9" pageNumber="294">18.4</td>
<td id="76875DE8FFF20058FC399A858A2DFD2E" box="[1021,1149,695,722]" gridcol="3" gridrow="14" pageId="9" pageNumber="294">18.6</td>
</tr>
<tr id="35563494FFF20058FFAC9AEA8A2DFD0F" box="[104,1149,728,755]" gridrow="15" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9AEA8FA3FD0F" box="[104,499,728,755]" gridcol="0" gridrow="15" pageId="9" pageNumber="294">Body width at anal fin origin</th>
<td id="76875DE8FFF20058FDF69AEA8CE2FD0F" box="[562,690,728,755]" gridcol="1" gridrow="15" pageId="9" pageNumber="294">12.5</td>
<td id="76875DE8FFF20058FCE19AEA8DF6FD0F" box="[805,934,728,755]" gridcol="2" gridrow="15" pageId="9" pageNumber="294">11.7</td>
<td id="76875DE8FFF20058FC399AEA8A2DFD0F" box="[1021,1149,728,755]" gridcol="3" gridrow="15" pageId="9" pageNumber="294">11.6</td>
</tr>
<tr id="35563494FFF20058FFAC9ACA8A2DFCEF" box="[104,1149,760,787]" gridrow="16" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9ACA8FA3FCEF" box="[104,499,760,787]" gridcol="0" gridrow="16" pageId="9" pageNumber="294">Body width at pelvic fin origin</th>
<td id="76875DE8FFF20058FDF69ACA8CE2FCEF" box="[562,690,760,787]" gridcol="1" gridrow="16" pageId="9" pageNumber="294">17.7</td>
<td id="76875DE8FFF20058FCE19ACA8DF6FCEF" box="[805,934,760,787]" gridcol="2" gridrow="16" pageId="9" pageNumber="294">18.4</td>
<td id="76875DE8FFF20058FC399ACA8A2DFCEF" box="[1021,1149,760,787]" gridcol="3" gridrow="16" pageId="9" pageNumber="294">17.6</td>
</tr>
<tr id="35563494FFF20058FFAC9B2A8A2DFCCF" box="[104,1149,792,819]" gridrow="17" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9B2A8FA3FCCF" box="[104,499,792,819]" gridcol="0" gridrow="17" pageId="9" pageNumber="294">Caudal fin length</th>
<td id="76875DE8FFF20058FDF69B2A8CE2FCCF" box="[562,690,792,819]" gridcol="1" gridrow="17" pageId="9" pageNumber="294">26.8</td>
<td id="76875DE8FFF20058FCE19B2A8DF6FCCF" box="[805,934,792,819]" gridcol="2" gridrow="17" pageId="9" pageNumber="294">24.6</td>
<td id="76875DE8FFF20058FC399B2A8A2DFCCF" box="[1021,1149,792,819]" gridcol="3" gridrow="17" pageId="9" pageNumber="294">24.2</td>
</tr>
<tr id="35563494FFF20058FFAC9B0B8A2DFCAF" box="[104,1149,825,851]" gridrow="18" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9B0B8FA3FCAF" box="[104,499,825,851]" gridcol="0" gridrow="18" pageId="9" pageNumber="294">Caudal peduncle depth</th>
<td id="76875DE8FFF20058FDF69B0B8CE2FCAF" box="[562,690,825,851]" gridcol="1" gridrow="18" pageId="9" pageNumber="294">12.6</td>
<td id="76875DE8FFF20058FCE19B0B8DF6FCAF" box="[805,934,825,851]" gridcol="2" gridrow="18" pageId="9" pageNumber="294">12.7</td>
<td id="76875DE8FFF20058FC399B0B8A2DFCAF" box="[1021,1149,825,851]" gridcol="3" gridrow="18" pageId="9" pageNumber="294">10.7</td>
</tr>
<tr id="35563494FFF20058FFAC9B6B8A2DFC8F" box="[104,1149,857,883]" gridrow="19" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9B6B8FA3FC8F" box="[104,499,857,883]" gridcol="0" gridrow="19" pageId="9" pageNumber="294">Caudal peduncle length</th>
<td id="76875DE8FFF20058FDF69B6B8CE2FC8F" box="[562,690,857,883]" gridcol="1" gridrow="19" pageId="9" pageNumber="294">21.0</td>
<td id="76875DE8FFF20058FCE19B6B8DF6FC8F" box="[805,934,857,883]" gridcol="2" gridrow="19" pageId="9" pageNumber="294">20.2</td>
<td id="76875DE8FFF20058FC399B6B8A2DFC8F" box="[1021,1149,857,883]" gridcol="3" gridrow="19" pageId="9" pageNumber="294">18.9</td>
</tr>
<tr id="35563494FFF20058FFAC9B4B8A2DFC68" box="[104,1149,889,916]" gridrow="20" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9B4B8FA3FC68" box="[104,499,889,916]" gridcol="0" gridrow="20" pageId="9" pageNumber="294">Eye diameter</th>
<td id="76875DE8FFF20058FDF69B4B8CE2FC68" box="[562,690,889,916]" gridcol="1" gridrow="20" pageId="9" pageNumber="294">7.7</td>
<td id="76875DE8FFF20058FCE19B4B8DF6FC68" box="[805,934,889,916]" gridcol="2" gridrow="20" pageId="9" pageNumber="294">7.0</td>
<td id="76875DE8FFF20058FC399B4B8A2DFC68" box="[1021,1149,889,916]" gridcol="3" gridrow="20" pageId="9" pageNumber="294">6.7</td>
</tr>
<tr id="35563494FFF20058FFAC9BAB8A2DFC48" box="[104,1149,921,948]" gridrow="21" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9BAB8FA3FC48" box="[104,499,921,948]" gridcol="0" gridrow="21" pageId="9" pageNumber="294">First dorsal fin base</th>
<td id="76875DE8FFF20058FDF69BAB8CE2FC48" box="[562,690,921,948]" gridcol="1" gridrow="21" pageId="9" pageNumber="294">13.1</td>
<td id="76875DE8FFF20058FCE19BAB8DF6FC48" box="[805,934,921,948]" gridcol="2" gridrow="21" pageId="9" pageNumber="294">13.4</td>
<td id="76875DE8FFF20058FC399BAB8A2DFC48" box="[1021,1149,921,948]" gridcol="3" gridrow="21" pageId="9" pageNumber="294">14.2</td>
</tr>
<tr id="35563494FFF20058FFAC9B888A2DFC29" box="[104,1149,954,981]" gridrow="22" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9B888FA3FC29" box="[104,499,954,981]" gridcol="0" gridrow="22" pageId="9" pageNumber="294">Head depth</th>
<td id="76875DE8FFF20058FDF69B888CE2FC29" box="[562,690,954,981]" gridcol="1" gridrow="22" pageId="9" pageNumber="294">15.7</td>
<td id="76875DE8FFF20058FCE19B888DF6FC29" box="[805,934,954,981]" gridcol="2" gridrow="22" pageId="9" pageNumber="294">17.5</td>
<td id="76875DE8FFF20058FC399B888A2DFC29" box="[1021,1149,954,981]" gridcol="3" gridrow="22" pageId="9" pageNumber="294">14.6</td>
</tr>
<tr id="35563494FFF20058FFAC9BE88A2DFC09" box="[104,1149,986,1013]" gridrow="23" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9BE88FA3FC09" box="[104,499,986,1013]" gridcol="0" gridrow="23" pageId="9" pageNumber="294">Head length</th>
<td id="76875DE8FFF20058FDF69BE88CE2FC09" box="[562,690,986,1013]" gridcol="1" gridrow="23" pageId="9" pageNumber="294">28.5</td>
<td id="76875DE8FFF20058FCE19BE88DF6FC09" box="[805,934,986,1013]" gridcol="2" gridrow="23" pageId="9" pageNumber="294">31.2</td>
<td id="76875DE8FFF20058FC399BE88A2DFC09" box="[1021,1149,986,1013]" gridcol="3" gridrow="23" pageId="9" pageNumber="294">29.0</td>
</tr>
<tr id="35563494FFF20058FFAC9BC88A2DFBE9" box="[104,1149,1018,1045]" gridrow="24" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9BC88FA3FBE9" box="[104,499,1018,1045]" gridcol="0" gridrow="24" pageId="9" pageNumber="294">Head width</th>
<td id="76875DE8FFF20058FDF69BC88CE2FBE9" box="[562,690,1018,1045]" gridcol="1" gridrow="24" pageId="9" pageNumber="294">25.5</td>
<td id="76875DE8FFF20058FCE19BC88DF6FBE9" box="[805,934,1018,1045]" gridcol="2" gridrow="24" pageId="9" pageNumber="294">26.0</td>
<td id="76875DE8FFF20058FC399BC88A2DFBE9" box="[1021,1149,1018,1045]" gridcol="3" gridrow="24" pageId="9" pageNumber="294">23.9</td>
</tr>
<tr id="35563494FFF20058FFAC9C288A2DFBC9" box="[104,1149,1050,1077]" gridrow="25" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9C288FA3FBC9" box="[104,499,1050,1077]" gridcol="0" gridrow="25" pageId="9" pageNumber="294">Pectoral fin length</th>
<td id="76875DE8FFF20058FDF69C288CE2FBC9" box="[562,690,1050,1077]" gridcol="1" gridrow="25" pageId="9" pageNumber="294">29.9</td>
<td id="76875DE8FFF20058FCE19C288DF6FBC9" box="[805,934,1050,1077]" gridcol="2" gridrow="25" pageId="9" pageNumber="294">26.0</td>
<td id="76875DE8FFF20058FC399C288A2DFBC9" box="[1021,1149,1050,1077]" gridcol="3" gridrow="25" pageId="9" pageNumber="294">26.1</td>
</tr>
<tr id="35563494FFF20058FFAC9C098A2DFBA9" box="[104,1149,1083,1109]" gridrow="26" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9C098FA3FBA9" box="[104,499,1083,1109]" gridcol="0" gridrow="26" pageId="9" pageNumber="294">Pelvic fin length</th>
<td id="76875DE8FFF20058FDF69C098CE2FBA9" box="[562,690,1083,1109]" gridcol="1" gridrow="26" pageId="9" pageNumber="294">21.0</td>
<td id="76875DE8FFF20058FCE19C098DF6FBA9" box="[805,934,1083,1109]" gridcol="2" gridrow="26" pageId="9" pageNumber="294">20.9</td>
<td id="76875DE8FFF20058FC399C098A2DFBA9" box="[1021,1149,1083,1109]" gridcol="3" gridrow="26" pageId="9" pageNumber="294">20.7</td>
</tr>
<tr id="35563494FFF20058FFAC9C698A2DFB8A" box="[104,1149,1115,1142]" gridrow="27" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9C698FA3FB8A" box="[104,499,1115,1142]" gridcol="0" gridrow="27" pageId="9" pageNumber="294">Pelvic to anus</th>
<td id="76875DE8FFF20058FDF69C698CE2FB8A" box="[562,690,1115,1142]" gridcol="1" gridrow="27" pageId="9" pageNumber="294">26.2</td>
<td id="76875DE8FFF20058FCE19C698DF6FB8A" box="[805,934,1115,1142]" gridcol="2" gridrow="27" pageId="9" pageNumber="294">24.8</td>
<td id="76875DE8FFF20058FC399C698A2DFB8A" box="[1021,1149,1115,1142]" gridcol="3" gridrow="27" pageId="9" pageNumber="294">27.2</td>
</tr>
<tr id="35563494FFF20058FFAC9C498A2DFB6A" box="[104,1149,1147,1174]" gridrow="28" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9C498FA3FB6A" box="[104,499,1147,1174]" gridcol="0" gridrow="28" pageId="9" pageNumber="294">Second dorsal fin base</th>
<td id="76875DE8FFF20058FDF69C498CE2FB6A" box="[562,690,1147,1174]" gridcol="1" gridrow="28" pageId="9" pageNumber="294">30.8</td>
<td id="76875DE8FFF20058FCE19C498DF6FB6A" box="[805,934,1147,1174]" gridcol="2" gridrow="28" pageId="9" pageNumber="294">27.5</td>
<td id="76875DE8FFF20058FC399C498A2DFB6A" box="[1021,1149,1147,1174]" gridcol="3" gridrow="28" pageId="9" pageNumber="294">26.4</td>
</tr>
<tr id="35563494FFF20058FFAC9CAE8A2DFB4A" box="[104,1149,1180,1206]" gridrow="29" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9CAE8FA3FB4A" box="[104,499,1180,1206]" gridcol="0" gridrow="29" pageId="9" pageNumber="294">Second dorsal fin spine length</th>
<td id="76875DE8FFF20058FDF69CAE8CE2FB4A" box="[562,690,1180,1206]" gridcol="1" gridrow="29" pageId="9" pageNumber="294">11.3</td>
<td id="76875DE8FFF20058FCE19CAE8DF6FB4A" box="[805,934,1180,1206]" gridcol="2" gridrow="29" pageId="9" pageNumber="294">13.3</td>
<td id="76875DE8FFF20058FC399CAE8A2DFB4A" box="[1021,1149,1180,1206]" gridcol="3" gridrow="29" pageId="9" pageNumber="294">11.0</td>
</tr>
<tr id="35563494FFF20058FFAC9C8E8A2DFB2A" box="[104,1149,1212,1238]" gridrow="30" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9C8E8FA3FB2A" box="[104,499,1212,1238]" gridcol="0" gridrow="30" pageId="9" pageNumber="294">Snout to origin of first dorsal fin</th>
<td id="76875DE8FFF20058FDF69C8E8CE2FB2A" box="[562,690,1212,1238]" gridcol="1" gridrow="30" pageId="9" pageNumber="294">36.0</td>
<td id="76875DE8FFF20058FCE19C8E8DF6FB2A" box="[805,934,1212,1238]" gridcol="2" gridrow="30" pageId="9" pageNumber="294">37.2</td>
<td id="76875DE8FFF20058FC399C8E8A2DFB2A" box="[1021,1149,1212,1238]" gridcol="3" gridrow="30" pageId="9" pageNumber="294">37.0</td>
</tr>
<tr id="35563494FFF20058FFAC9CEE8A2DFB0B" box="[104,1149,1244,1271]" gridrow="31" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9CEE8FA3FB0B" box="[104,499,1244,1271]" gridcol="0" gridrow="31" pageId="9" pageNumber="294">Snout to origin of second dorsal fin</th>
<td id="76875DE8FFF20058FDF69CEE8CE2FB0B" box="[562,690,1244,1271]" gridcol="1" gridrow="31" pageId="9" pageNumber="294">56.3</td>
<td id="76875DE8FFF20058FCE19CEE8DF6FB0B" box="[805,934,1244,1271]" gridcol="2" gridrow="31" pageId="9" pageNumber="294">56.8</td>
<td id="76875DE8FFF20058FC399CEE8A2DFB0B" box="[1021,1149,1244,1271]" gridcol="3" gridrow="31" pageId="9" pageNumber="294">56.8</td>
</tr>
<tr id="35563494FFF20058FFAC9CCE8A2DFAEB" box="[104,1149,1276,1303]" gridrow="32" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9CCE8FA3FAEB" box="[104,499,1276,1303]" gridcol="0" gridrow="32" pageId="9" pageNumber="294">Snout to vertical of anal fin origin</th>
<td id="76875DE8FFF20058FDF69CCE8CE2FAEB" box="[562,690,1276,1303]" gridcol="1" gridrow="32" pageId="9" pageNumber="294">61.2</td>
<td id="76875DE8FFF20058FCE19CCE8DF6FAEB" box="[805,934,1276,1303]" gridcol="2" gridrow="32" pageId="9" pageNumber="294">61.1</td>
<td id="76875DE8FFF20058FC399CCE8A2DFAEB" box="[1021,1149,1276,1303]" gridcol="3" gridrow="32" pageId="9" pageNumber="294">61.3</td>
</tr>
<tr id="35563494FFF20058FFAC9D2E8A2DFACB" box="[104,1149,1308,1335]" gridrow="33" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9D2E8FA3FACB" box="[104,499,1308,1335]" gridcol="0" gridrow="33" pageId="9" pageNumber="294">Snout to vertical of anus</th>
<td id="76875DE8FFF20058FDF69D2E8CE2FACB" box="[562,690,1308,1335]" gridcol="1" gridrow="33" pageId="9" pageNumber="294">58.5</td>
<td id="76875DE8FFF20058FCE19D2E8DF6FACB" box="[805,934,1308,1335]" gridcol="2" gridrow="33" pageId="9" pageNumber="294">56.5</td>
<td id="76875DE8FFF20058FC399D2E8A2DFACB" box="[1021,1149,1308,1335]" gridcol="3" gridrow="33" pageId="9" pageNumber="294">56.4</td>
</tr>
<tr id="35563494FFF20058FFAC9D0F8A2DFAA4" box="[104,1149,1341,1368]" gridrow="34" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9D0F8FA3FAA4" box="[104,499,1341,1368]" gridcol="0" gridrow="34" pageId="9" pageNumber="294">Snout to vertical of pelvic fin origin</th>
<td id="76875DE8FFF20058FDF69D0F8CE2FAA4" box="[562,690,1341,1368]" gridcol="1" gridrow="34" pageId="9" pageNumber="294">31.3</td>
<td id="76875DE8FFF20058FCE19D0F8DF6FAA4" box="[805,934,1341,1368]" gridcol="2" gridrow="34" pageId="9" pageNumber="294">31.0</td>
<td id="76875DE8FFF20058FC399D0F8A2DFAA4" box="[1021,1149,1341,1368]" gridcol="3" gridrow="34" pageId="9" pageNumber="294">29.5</td>
</tr>
<tr id="35563494FFF20058FFAC9D6F8A2DFA8B" box="[104,1149,1373,1399]" gridrow="35" pageId="9" pageNumber="294" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76875DE8FFF20058FFAC9D6F8FA3FA8B" box="[104,499,1373,1399]" gridcol="0" gridrow="35" pageId="9" pageNumber="294">% of caudal peduncle</th>
</tr>
<tr id="35563494FFF20058FFAC9D4F8A2DFA64" box="[104,1149,1405,1432]" gridrow="36" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9D4F8FA3FA64" box="[104,499,1405,1432]" gridcol="0" gridrow="36" pageId="9" pageNumber="294">Caudal peduncle depth</th>
<td id="76875DE8FFF20058FDF69D4F8CE2FA64" box="[562,690,1405,1432]" gridcol="1" gridrow="36" pageId="9" pageNumber="294">59.9</td>
<td id="76875DE8FFF20058FCE19D4F8DF6FA64" box="[805,934,1405,1432]" gridcol="2" gridrow="36" pageId="9" pageNumber="294">62.6</td>
<td id="76875DE8FFF20058FC399D4F8A2DFA64" box="[1021,1149,1405,1432]" gridcol="3" gridrow="36" pageId="9" pageNumber="294">56.8</td>
</tr>
<tr id="35563494FFF20058FFAC9DAC8A2DFA44" box="[104,1149,1438,1464]" gridrow="37" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9DAC8A2DFA44" box="[104,1149,1438,1464]" colspan="4" colspanRight="3" gridcol="0" gridrow="37" pageId="9" pageNumber="294">% of head length</th>
</tr>
<tr id="35563494FFF20058FFAC9D8C8A2DFA25" box="[104,1149,1470,1497]" gridrow="38" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9D8C8FA3FA25" box="[104,499,1470,1497]" gridcol="0" gridrow="38" pageId="9" pageNumber="294">Cheek depth</th>
<td id="76875DE8FFF20058FDF69D8C8CE2FA25" box="[562,690,1470,1497]" gridcol="1" gridrow="38" pageId="9" pageNumber="294">27.1</td>
<td id="76875DE8FFF20058FCE19D8C8DF6FA25" box="[805,934,1470,1497]" gridcol="2" gridrow="38" pageId="9" pageNumber="294">22.5</td>
<td id="76875DE8FFF20058FC399D8C8A2DFA25" box="[1021,1149,1470,1497]" gridcol="3" gridrow="38" pageId="9" pageNumber="294">22.9</td>
</tr>
<tr id="35563494FFF20058FFAC9DEC8A2DFA05" box="[104,1149,1502,1529]" gridrow="39" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9DEC8FA3FA05" box="[104,499,1502,1529]" gridcol="0" gridrow="39" pageId="9" pageNumber="294">Eye diameter</th>
<td id="76875DE8FFF20058FDF69DEC8CE2FA05" box="[562,690,1502,1529]" gridcol="1" gridrow="39" pageId="9" pageNumber="294">23.2</td>
<td id="76875DE8FFF20058FCE19DEC8DF6FA05" box="[805,934,1502,1529]" gridcol="2" gridrow="39" pageId="9" pageNumber="294">21.4</td>
<td id="76875DE8FFF20058FC399DEC8A2DFA05" box="[1021,1149,1502,1529]" gridcol="3" gridrow="39" pageId="9" pageNumber="294">21.7</td>
</tr>
<tr id="35563494FFF20058FFAC9DCC8A2DF9E5" box="[104,1149,1534,1561]" gridrow="40" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9DCC8FA3F9E5" box="[104,499,1534,1561]" gridcol="0" gridrow="40" pageId="9" pageNumber="294">Head depth</th>
<td id="76875DE8FFF20058FDF69DCC8CE2F9E5" box="[562,690,1534,1561]" gridcol="1" gridrow="40" pageId="9" pageNumber="294">55.2</td>
<td id="76875DE8FFF20058FCE19DCC8DF6F9E5" box="[805,934,1534,1561]" gridcol="2" gridrow="40" pageId="9" pageNumber="294">56.1</td>
<td id="76875DE8FFF20058FC399DCC8A2DF9E5" box="[1021,1149,1534,1561]" gridcol="3" gridrow="40" pageId="9" pageNumber="294">50.1</td>
</tr>
<tr id="35563494FFF20058FFAC9E2D8A2DF9C6" box="[104,1149,1567,1594]" gridrow="41" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9E2D8FA3F9C6" box="[104,499,1567,1594]" gridcol="0" gridrow="41" pageId="9" pageNumber="294">Head width</th>
<td id="76875DE8FFF20058FDF69E2D8CE2F9C6" box="[562,690,1567,1594]" gridcol="1" gridrow="41" pageId="9" pageNumber="294">89.5</td>
<td id="76875DE8FFF20058FCE19E2D8DF6F9C6" box="[805,934,1567,1594]" gridcol="2" gridrow="41" pageId="9" pageNumber="294">83.4</td>
<td id="76875DE8FFF20058FC399E2D8A2DF9C6" box="[1021,1149,1567,1594]" gridcol="3" gridrow="41" pageId="9" pageNumber="294">82.2</td>
</tr>
<tr id="35563494FFF20058FFAC9E0D8A2DF9A6" box="[104,1149,1599,1626]" gridrow="42" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9E0D8FA3F9A6" box="[104,499,1599,1626]" gridcol="0" gridrow="42" pageId="9" pageNumber="294">Postorbital length</th>
<td id="76875DE8FFF20058FDF69E0D8CE2F9A6" box="[562,690,1599,1626]" gridcol="1" gridrow="42" pageId="9" pageNumber="294">55.3</td>
<td id="76875DE8FFF20058FCE19E0D8DF6F9A6" box="[805,934,1599,1626]" gridcol="2" gridrow="42" pageId="9" pageNumber="294">56.6</td>
<td id="76875DE8FFF20058FC399E0D8A2DF9A6" box="[1021,1149,1599,1626]" gridcol="3" gridrow="42" pageId="9" pageNumber="294">57.4</td>
</tr>
<tr id="35563494FFF20058FFAC9E6D8A2DF985" box="[104,1149,1631,1657]" gridrow="43" pageId="9" pageNumber="294">
<th id="76875DE8FFF20058FFAC9E6D8FA3F985" box="[104,499,1631,1657]" gridcol="0" gridrow="43" pageId="9" pageNumber="294">Snout length</th>
<td id="76875DE8FFF20058FDF69E6D8CE2F985" box="[562,690,1631,1657]" gridcol="1" gridrow="43" pageId="9" pageNumber="294">26.8</td>
<td id="76875DE8FFF20058FCE19E6D8DF6F985" box="[805,934,1631,1657]" gridcol="2" gridrow="43" pageId="9" pageNumber="294">23.9</td>
<td id="76875DE8FFF20058FC399E6D8A2DF985" box="[1021,1149,1631,1657]" gridcol="3" gridrow="43" pageId="9" pageNumber="294">26.0</td>
</tr>
</table>
</paragraph>
<caption id="DF19665EFFF1FFADFF4498978A2CFF47" box="[128,1148,165,188]" pageId="10" pageNumber="295" startId="10.[128,188,165,187]" targetBox="[134,1179,208,457]" targetIsTable="true" targetPageId="10" targetType="table">
<paragraph id="8BD936D6FFF1FFADFF4498978A2CFF47" blockId="10.[128,1148,165,188]" box="[128,1148,165,188]" pageId="10" pageNumber="295">
table 2 Morphometric characters (as proportional measurements in %) of 
<taxonomicName id="4C664D55FFF1FFADFC9A98978DA0FF46" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[862,1008,165,186]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="295" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF1FFADFC9A98978DA0FF46" box="[862,1008,165,186]" italics="true" pageId="10" pageNumber="295">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF1FFADFC3198948A69FF47" box="[1013,1081,166,187]" pageId="10" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B912EAC4FFF1FFADFB8098948A3EFF47" box="[1092,1134,166,187]" italics="true" pageId="10" pageNumber="295">Cont</emphasis>
.)
</paragraph>
</caption>
<paragraph id="8BD936D6FFF1FFADFF4298E28A1AFE35" pageId="10" pageNumber="295">
<table id="F966C476FFF10058FF4298E28ACBFE35" box="[134,1179,208,457]" colsContinueFrom="9.[104,1149,208,1657]" gridcols="4" gridrows="7" pageId="10" pageNumber="295">
<tr id="35563494FFF10058FF4298E28ACBFF16" box="[134,1179,208,234]" gridrow="0" pageId="10" pageNumber="295">
<th id="76875DE8FFF10058FF4298E28F01FF16" box="[134,337,208,234]" gridcol="0" gridrow="0" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFF4298E28EA9FF16" bold="true" box="[134,249,208,234]" pageId="10" pageNumber="295">Specimen</emphasis>
</th>
<th id="76875DE8FFF10058FD9598E28C80FF16" box="[593,720,208,234]" gridcol="1" gridrow="0" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFD9598E28C80FF16" bold="true" box="[593,720,208,234]" pageId="10" pageNumber="295">P6V 144302</emphasis>
</th>
<th id="76875DE8FFF10058FC8098E28D94FF16" box="[836,964,208,234]" gridcol="2" gridrow="0" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFC8098E28D94FF16" bold="true" box="[836,964,208,234]" pageId="10" pageNumber="295">P6V 144300</emphasis>
</th>
<th id="76875DE8FFF10058FBD898E28ACBFF16" box="[1052,1179,208,234]" gridcol="3" gridrow="0" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFBD898E28ACBFF16" bold="true" box="[1052,1179,208,234]" pageId="10" pageNumber="295">P6V 144303</emphasis>
</th>
</tr>
<tr id="35563494FFF10058FF4298C68ACBFEF2" box="[134,1179,244,270]" gridrow="1" pageId="10" pageNumber="295" rowspan-0="1">
<td id="76875DE8FFF10058FD9598C68C80FEF2" box="[593,720,244,270]" gridcol="1" gridrow="1" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFD9598C68CE8FEF2" bold="true" box="[593,696,244,270]" pageId="10" pageNumber="295">holotype</emphasis>
</td>
<td id="76875DE8FFF10058FC8098C68D94FEF2" box="[836,964,244,270]" gridcol="2" gridrow="1" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFC8098C68DFBFEF2" bold="true" box="[836,939,244,270]" pageId="10" pageNumber="295">paratype</emphasis>
</td>
<td id="76875DE8FFF10058FBD898C68ACBFEF2" box="[1052,1179,244,270]" gridcol="3" gridrow="1" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFBD898C68AD3FEF2" bold="true" box="[1052,1155,244,270]" pageId="10" pageNumber="295">paratype</emphasis>
</td>
</tr>
<tr id="35563494FFF10058FF4299168ACBFEC3" box="[134,1179,292,319]" gridrow="2" pageId="10" pageNumber="295">
<th id="76875DE8FFF10058FF4299168F01FEC3" box="[134,337,292,319]" gridcol="0" gridrow="2" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFF4299168EFEFEC2" bold="true" box="[134,174,292,318]" pageId="10" pageNumber="295">Sex</emphasis>
</th>
<td id="76875DE8FFF10058FD9599168C80FEC3" box="[593,720,292,319]" gridcol="1" gridrow="2" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFD9599178CDAFEC3" bold="true" box="[593,650,293,319]" pageId="10" pageNumber="295">male</emphasis>
</td>
<td id="76875DE8FFF10058FC8099168D94FEC3" box="[836,964,292,319]" gridcol="2" gridrow="2" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFC8099178D2DFEC3" bold="true" box="[836,893,293,319]" pageId="10" pageNumber="295">male</emphasis>
</td>
<td id="76875DE8FFF10058FBD899168ACBFEC3" box="[1052,1179,292,319]" gridcol="3" gridrow="2" pageId="10" pageNumber="295">
<emphasis id="B912EAC4FFF1FFADFBD899168A3BFEC2" bold="true" box="[1052,1131,292,318]" pageId="10" pageNumber="295">female</emphasis>
</td>
</tr>
<tr id="35563494FFF10058FF42997F8ACBFE94" box="[134,1179,333,360]" gridrow="3" pageId="10" pageNumber="295" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76875DE8FFF10058FF42997F8F01FE94" box="[134,337,333,360]" gridcol="0" gridrow="3" pageId="10" pageNumber="295">% of eye diameter</th>
</tr>
<tr id="35563494FFF10058FF42995F8ACBFE74" box="[134,1179,365,392]" gridrow="4" pageId="10" pageNumber="295">
<th id="76875DE8FFF10058FF42995F8F01FE74" box="[134,337,365,392]" gridcol="0" gridrow="4" pageId="10" pageNumber="295">Interorbital width</th>
<td id="76875DE8FFF10058FD95995F8C80FE74" box="[593,720,365,392]" gridcol="1" gridrow="4" pageId="10" pageNumber="295">51.4</td>
<td id="76875DE8FFF10058FC80995F8D94FE74" box="[836,964,365,392]" gridcol="2" gridrow="4" pageId="10" pageNumber="295">49.7</td>
<td id="76875DE8FFF10058FBD8995F8ACBFE74" box="[1052,1179,365,392]" gridcol="3" gridrow="4" pageId="10" pageNumber="295">51.5</td>
</tr>
<tr id="35563494FFF10058FF4299BC8ACBFE54" box="[134,1179,398,424]" gridrow="5" pageId="10" pageNumber="295">
<th id="76875DE8FFF10058FF4299BC8ACBFE54" box="[134,1179,398,424]" colspan="4" colspanRight="3" gridcol="0" gridrow="5" pageId="10" pageNumber="295">% of pelvic to anus</th>
</tr>
<tr id="35563494FFF10058FF42999C8ACBFE35" box="[134,1179,430,457]" gridrow="6" pageId="10" pageNumber="295">
<th id="76875DE8FFF10058FF42999C8F01FE35" box="[134,337,430,457]" gridcol="0" gridrow="6" pageId="10" pageNumber="295">Pelvic fin length</th>
<td id="76875DE8FFF10058FD95999C8C80FE35" box="[593,720,430,457]" gridcol="1" gridrow="6" pageId="10" pageNumber="295">80.3</td>
<td id="76875DE8FFF10058FC80999C8D94FE35" box="[836,964,430,457]" gridcol="2" gridrow="6" pageId="10" pageNumber="295">84.3</td>
<td id="76875DE8FFF10058FBD8999C8ACBFE35" box="[1052,1179,430,457]" gridcol="3" gridrow="6" pageId="10" pageNumber="295">75.9</td>
</tr>
</table>
</paragraph>
<caption id="DF19665EFFF1FFADFF449E268F3CF963" ID-DOI="http://doi.org/10.5281/zenodo.8343385" ID-Zenodo-Dep="8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="10" pageNumber="295" startId="10.[128,195,1556,1578]" targetBox="[223,1092,575,1537]" targetPageId="10" targetType="figure">
<paragraph id="8BD936D6FFF1FFADFF449E268F3CF963" blockId="10.[128,1159,1556,1695]" pageId="10" pageNumber="295">
figure 1 Preserved specimens. (A) 
<taxonomicName id="4C664D55FFF1FFADFE219E268C28F9D5" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[485,632,1556,1577]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="295" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF1FFADFE219E268C28F9D5" box="[485,632,1556,1577]" italics="true" pageId="10" pageNumber="295">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF1FFADFDB99E278C92F9D6" box="[637,706,1557,1578]" pageId="10" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
, npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M. Kovačić. (B) 
<taxonomicName id="4C664D55FFF1FFADFC0D9E038A16F9BA" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[969,1094,1585,1606]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="295" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF1FFADFC0D9E038A16F9BA" box="[969,1094,1585,1606]" italics="true" pageId="10" pageNumber="295">Zebrus zebrus</emphasis>
</taxonomicName>
, nmp P6V 140912, neotype, female, 23.25 + 6.22 mm, Îll Gross, Banyuls sur Mer, France (C) 
<emphasis id="B912EAC4FFF1FFADFC379E7D8FD7F97D" italics="true" pageId="10" pageNumber="295">
<taxonomicName id="4C664D55FFF1FFADFC379E7D8FD3F97D" baseAuthorityName="Kolombatovic" baseAuthorityYear="1891" family="Gobiidae" genus="Millerigobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="295" phylum="Chordata" rank="species" species="macrocephalus">Millerigobius macrocephalus</taxonomicName>
,
</emphasis>
nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France.
</paragraph>
</caption>
<caption id="DF19665EFFF0FFACFFA69DD98F97F928" ID-DOI="http://doi.org/10.5281/zenodo.8343387" ID-Zenodo-Dep="8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="11" pageNumber="296" startId="11.[98,165,1515,1537]" targetBox="[193,1062,520,1497]" targetPageId="11" targetType="figure">
<paragraph id="8BD936D6FFF0FFACFFA69DD98F97F928" blockId="11.[98,1156,1515,1748]" pageId="11" pageNumber="296">
figure 2 
<taxonomicName id="4C664D55FFF0FFACFF1F9DD98F3EF9FC" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[219,366,1515,1536]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="296" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF0FFACFF1F9DD98F3EF9FC" box="[219,366,1515,1536]" italics="true" pageId="11" pageNumber="296">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF0FFACFEB79DDE8FE7F9FD" box="[371,439,1516,1537]" pageId="11" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. 
<emphasis id="B912EAC4FFF0FFACFE529E538C47F98A" box="[406,535,1633,1654]" italics="true" pageId="11" pageNumber="296">
<taxonomicName id="4C664D55FFF0FFACFE529E538C43F98A" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[406,531,1633,1654]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="296" phylum="Chordata" rank="species" species="zebrus">Zebrus zebrus</taxonomicName>
:
</emphasis>
nmp P6V 142593, male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ.
</paragraph>
</caption>
<paragraph id="8BD936D6FFF7FFABFF5B9CA88D29FB97" blockId="12.[129,640,171,1742]" lastBlockId="12.[676,1187,171,1741]" pageId="12" pageNumber="297">
<emphasis id="B912EAC4FFF7FFABFF5B9CA88E83FB4F" box="[159,211,1178,1203]" italics="true" pageId="12" pageNumber="297">Fins.</emphasis>
First dorsal fin vi, second dorsal fin I+10 (I+10–11); anal fin I+8 (I+8–9); branched caudal-fin rays 16 (15), segmented 18 (17), pectoral-fin rays 17 left fin, right fin cut (in all 
<typeStatus id="54DD8874FFF7FFABFF6F9D1B8E8DFABF" box="[171,221,1321,1347]" pageId="12" pageNumber="297">type</typeStatus>
material), pelvic fins I+5/5+I. 
<collectingRegion id="49A2F834FFF7FFABFD9E9D1A8C2FFABE" box="[602,639,1320,1346]" country="Cayman Islands" name="East End" pageId="12" pageNumber="297">Fin</collectingRegion>
morphometrics in proportion to standard body length given in 
<tableCitation id="C6E4036DFFF7FFABFEB99D428F9FFA76" box="[381,463,1392,1418]" captionStart="table 2" captionStartId="9.[98,158,165,187]" captionTargetPageId="9" captionText="table 2 Morphometric characters (as proportional measurements in %) of Zebrus pallaoroi sp.nov." httpUri="http://table.plazi.org/id/DF19665EFFF2FFAEFFA698978A4BFF47" pageId="12" pageNumber="297" tableUuid="DF19665EFFF2FFAEFFA698978A4BFF47">table 2</tableCitation>
. Spines of first dorsal fin not elongate or filamentous, spines ii-v about equal, the third spine longest; spines v and vi of first dorsal fin reaching the origin of the second dorsal fin when folded down. Origin of first dorsal fin behind vertical at the pectoral-fin base. Interdorsal space with a well-developed membranous connection between dorsal fins. The second dorsal fin originates above the anus in the female and above urogenital papilla in males, with the longest rays hardly reaching the base of uppermost caudal-fin rays in males, and not reaching in the female. Origin of anal fin below vertical of the first segmented ray of the second dorsal fin in males, or second in the female. Anal fin with last ray origin below penultimate ray of the second dorsal fin in the female, below third from the end in males. Posterior tip of anal fin not reaching the base of lowermost caudal-fin rays. Pectoral-fin rays all branched. Pectoral fin with three uppermost rays bifid, partially free from the membrane, and three more with shorter free tips, first ray free tip 0.6–0.7 of entire first ray length. Pectoral fins nearly extending posteriorly to below origin of the second dorsal fin. Pelvic fins disc complete, rounded, with ray 5 longest, all rays branched, anterior membrane well-developed, in midline depth about 2/3 (exactly 65‒68% in the 
<typeStatus id="54DD8874FFF7FFABFD0A9BAC8D50FC44" box="[718,768,926,952]" pageId="12" pageNumber="297">type</typeStatus>
material) of spinous ray and without lateral lobes, but with tips of spinous ray merely visible (
<figureCitation id="135D2A53FFF7FFABFC939BD78DF5FC03" box="[855,933,997,1023]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="12" pageNumber="297">fig. 2B</figureCitation>
). Pelvic fins posterior edge before anus in both sexes. Caudal fin rounded, shorter than the head, 1.1 (1.2 and 1.3) in head length.
</paragraph>
<paragraph id="8BD936D6FFF7FFABFD069C478D5CF931" blockId="12.[676,1187,171,1741]" pageId="12" pageNumber="297">
<emphasis id="B912EAC4FFF7FFABFD069C478D5CFB73" box="[706,780,1141,1167]" italics="true" pageId="12" pageNumber="297">Scales.</emphasis>
Body with ctenoid scales. Scales in lateral series 32 (male 
<typeStatus id="54DD8874FFF7FFABFC7E9CAB8A4DFB4F" box="[954,1053,1177,1203]" pageId="12" pageNumber="297">paratype</typeStatus>
32, female 
<typeStatus id="54DD8874FFF7FFABFD609C8F8D57FB2B" box="[676,775,1213,1239]" pageId="12" pageNumber="297">paratype</typeStatus>
left 33 and right side damaged for the count) left, with one more row of small scales on caudal fin; in transverse series 11 (male 
<typeStatus id="54DD8874FFF7FFABFCD49D1B8D23FABF" box="[784,883,1321,1347]" pageId="12" pageNumber="297">paratype</typeStatus>
left 10 and 11 right side, female 
<typeStatus id="54DD8874FFF7FFABFCCA9D7F8D21FA9B" box="[782,881,1357,1383]" pageId="12" pageNumber="297">paratype</typeStatus>
10); circumpeduncular scales 12. Head with cheek and opercle naked. Predorsal area and first dorsal fin base naked to the last spine, with the upper edge of the scaled area from behind upper part of pectoral axilla backward and up to the last first dorsal-fin spine. Prepectoral and breast naked. Belly naked anteriorly, with cycloid scales posteriorly and laterally. Uppermost and lowermost scales of caudal peduncle not enlarged.
</paragraph>
<caption id="DF19665EFFF6FFAAFFA69ADE8F86FCBF" ID-DOI="http://doi.org/10.5281/zenodo.8343389" ID-Zenodo-Dep="8343389" httpUri="https://zenodo.org/record/8343389/files/figure.png" pageId="13" pageNumber="298" startId="13.[98,165,748,770]" targetBox="[195,1063,166,730]" targetPageId="13" targetType="figure">
<paragraph id="8BD936D6FFF6FFAAFFA69ADE8F86FCBF" blockId="13.[98,1138,748,835]" pageId="13" pageNumber="298">
figure 3 
<taxonomicName id="4C664D55FFF6FFAAFF1F9ADE8F3EFCFD" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[219,366,748,769]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="298" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF6FFAAFF1F9ADE8F3EFCFD" box="[219,366,748,769]" italics="true" pageId="13" pageNumber="298">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF6FFAAFEB79ADF8FE7FCFE" box="[371,439,749,770]" pageId="13" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
nmp P6V 144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: head lateral line sensory papillae and canal pores. drawing by m. kovačiĆ.
</paragraph>
</caption>
<paragraph id="8BD936D6FFF6FFA9FF459B4B8E81FBDF" blockId="13.[98,610,889,1706]" lastBlockId="14.[128,640,171,1706]" lastPageId="14" lastPageNumber="299" pageId="13" pageNumber="298">
<emphasis id="B912EAC4FFF6FFAAFF459B4B8F01FC68" box="[129,337,889,916]" italics="true" pageId="13" pageNumber="298">Lateral line system(</emphasis>
<figureCitation id="135D2A53FFF6FFAAFE959B4B8FD1FC68" box="[337,385,889,916]" captionStart="figure 3" captionStartId="13.[98,165,748,770]" captionTargetBox="[195,1063,166,730]" captionTargetId="figure-456@13.[193,1065,164,732]" captionTargetPageId="13" captionText="figure 3 Zebrus pallaoroi sp. nov.nmp P6V 144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: head lateral line sensory papillae and canal pores. drawing by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343389" httpUri="https://zenodo.org/record/8343389/files/figure.png" pageId="13" pageNumber="298">fig.3</figureCitation>
<emphasis id="B912EAC4FFF6FFAAFE459B488FC0FC68" box="[385,400,890,916]" italics="true" pageId="13" pageNumber="298">).</emphasis>
Head with anterior and posterior oculoscapular canals and preopercular canal with pores 
<emphasis id="B912EAC4FFF6FFAAFE689BF08E28FC03" italics="true" pageId="13" pageNumber="298">σ, λ, κ, ω,α, β, Ρ,Ρ1, Ρ2</emphasis>
,and 
<emphasis id="B912EAC4FFF6FFAAFF749BD48EB4FC03" box="[176,228,997,1023]" italics="true" pageId="13" pageNumber="298">γ, δ, Ε</emphasis>
, respectively.Pores large, with pore 
<emphasis id="B912EAC4FFF6FFAAFFA69C388E3FFBDF" box="[98,111,1034,1059]" italics="true" pageId="13" pageNumber="298">α</emphasis>
diameter about half of the distance between pore 
<emphasis id="B912EAC4FFF6FFAAFF5A9C1C8EF9FBBB" box="[158,169,1070,1095]" italics="true" pageId="13" pageNumber="298">Ρ</emphasis>
and 
<emphasis id="B912EAC4FFF6FFAAFF229C1C8EA8FBBB" box="[230,248,1069,1095]" italics="true" pageId="13" pageNumber="298">Ρ1</emphasis>
. Rows of head sensory papillae were counted on all type material. 
<emphasis id="B912EAC4FFF6FFAAFE329C638EC4FB73" italics="true" pageId="13" pageNumber="298">Preorbital rows</emphasis>
: snout with four median preorbital series, upper row 
<emphasis id="B912EAC4FFF6FFAAFF259CA88EBBFB4F" box="[225,235,1178,1203]" italics="true" pageId="13" pageNumber="298">r</emphasis>
(3) middorsally from pore 
<emphasis id="B912EAC4FFF6FFAAFDE69CA88C7EFB4F" box="[546,558,1178,1203]" italics="true" pageId="13" pageNumber="298">σ</emphasis>
and above posterior nostrils, 
<emphasis id="B912EAC4FFF6FFAAFEBA9C8F8FDFFB2B" box="[382,399,1213,1239]" italics="true" pageId="13" pageNumber="298">s1</emphasis>
(3‒4) at posterior nostril, row 
<emphasis id="B912EAC4FFF6FFAAFF369CD38F56FB07" box="[242,262,1249,1275]" italics="true" pageId="13" pageNumber="298">s2</emphasis>
(2–3) at anterior nostril, and vertical row 
<emphasis id="B912EAC4FFF6FFAAFF299D378F51FAE3" box="[237,257,1285,1311]" italics="true" pageId="13" pageNumber="298">s3</emphasis>
(2) above the upper lip. Lateral series 
<emphasis id="B912EAC4FFF6FFAAFF6C9D1B8EE3FABE" box="[168,179,1321,1346]" italics="true" pageId="13" pageNumber="298">c</emphasis>
in four parts: superior 
<emphasis id="B912EAC4FFF6FFAAFE709D1B8F9AFABF" box="[436,458,1321,1347]" italics="true" pageId="13" pageNumber="298">c2</emphasis>
(3 + 3 – 3 + 4) as two rows between the posterior nostril and anterior nasal tube, middle 
<emphasis id="B912EAC4FFF6FFAAFE6A9D438F90FA76" box="[430,448,1392,1418]" italics="true" pageId="13" pageNumber="298">c1</emphasis>
(5‒6) as two rows or as a cluster of papillae just below the base of the anterior nasal tube, inferior rows: upper horizontal 
<emphasis id="B912EAC4FFF6FFAAFEE29DEF8F6CFA0A" box="[294,316,1500,1526]" italics="true" pageId="13" pageNumber="298">c2</emphasis>
(5‒6) above the upper lip and lower horizontal 
<emphasis id="B912EAC4FFF6FFAAFE959E338F33F9E6" box="[337,355,1536,1562]" italics="true" pageId="13" pageNumber="298">c1</emphasis>
(3) between the upper lip and row 
<emphasis id="B912EAC4FFF6FFAAFF2A9E168EA5F9C2" box="[238,245,1572,1598]" italics="true" pageId="13" pageNumber="298">1</emphasis>
. 
<emphasis id="B912EAC4FFF6FFAAFEC79E168FFDF9C2" box="[259,429,1572,1598]" italics="true" pageId="13" pageNumber="298">Suborbital rows</emphasis>
: no row 
<emphasis id="B912EAC4FFF6FFAAFDD09E178C72F9C2" box="[532,546,1573,1598]" italics="true" pageId="13" pageNumber="298">a</emphasis>
. Row 
<emphasis id="B912EAC4FFF6FFAAFFA69E7A8E3FF99E" box="[98,111,1608,1634]" italics="true" pageId="13" pageNumber="298">b</emphasis>
(8) longitudinal, short, ends anteriorly near row 
<emphasis id="B912EAC4FFF6FFAAFF579E5E8EF0F97A" box="[147,160,1644,1670]" italics="true" pageId="13" pageNumber="298">4</emphasis>
and below pupil or pupil posterior edge and posteriorly distant from pore 
<emphasis id="B912EAC4FFF6FFAAFDC59EA28C5CF956" box="[513,524,1680,1706]" italics="true" pageId="13" pageNumber="298">δ</emphasis>
. Seven transverse suborbital rows of sensory papillae, four suborbital rows in front of row 
<emphasis id="B912EAC4FFF6FFAAFBE99BAF8A6AFC4B" box="[1069,1082,925,951]" italics="true" pageId="13" pageNumber="298">b</emphasis>
, three above row 
<emphasis id="B912EAC4FFF6FFAAFCC39BF38D49FC27" box="[775,793,961,987]" italics="true" pageId="13" pageNumber="298">b,</emphasis>
two transverse suborbital rows below row 
<emphasis id="B912EAC4FFF6FFAAFCCD9BD78D46FC03" box="[777,790,997,1023]" italics="true" pageId="13" pageNumber="298">b</emphasis>
. Row 
<emphasis id="B912EAC4FFF6FFAAFCA69BD78D39FC03" box="[866,873,997,1023]" italics="true" pageId="13" pageNumber="298">1</emphasis>
vertical, upper edges of rows 
<quantity id="4C9E9B33FFF6FFAAFD069C3B8D55FBDF" box="[706,773,1033,1060]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="12.7" metricValueMin="5.08" pageId="13" pageNumber="298" unit="in" value="3.5" valueMax="5.0" valueMin="2.0">
<emphasis id="B912EAC4FFF6FFAAFD069C3B8CB8FBD8" box="[706,744,1033,1060]" italics="true" pageId="13" pageNumber="298">2‒5</emphasis>
in
</quantity>
the level and close to orbit in the female 
<typeStatus id="54DD8874FFF6FFAAFD1D9C1C8D6BFBB4" box="[729,827,1070,1096]" pageId="13" pageNumber="298">paratype</typeStatus>
or, in the 
<typeStatus id="54DD8874FFF6FFAAFC759C1F8A44FBBB" box="[945,1044,1069,1095]" pageId="13" pageNumber="298">holotype</typeStatus>
and male 
<typeStatus id="54DD8874FFF6FFAAFD419C638CB9FB97" box="[645,745,1105,1131]" pageId="13" pageNumber="298">paratype</typeStatus>
, rows 
<emphasis id="B912EAC4FFF6FFAAFCFE9C638D31FB97" box="[826,865,1105,1131]" italics="true" pageId="13" pageNumber="298">2–4</emphasis>
well separated from the lower border of orbit, row 
<emphasis id="B912EAC4FFF6FFAAFC779C478DEFFB73" box="[947,959,1141,1167]" italics="true" pageId="13" pageNumber="298">5</emphasis>
divided by row 
<emphasis id="B912EAC4FFF6FFAAFBB39C478AD4FB73" box="[1143,1156,1141,1167]" italics="true" pageId="13" pageNumber="298">b</emphasis>
into upper (
<emphasis id="B912EAC4FFF6FFAAFCCD9CAB8D4FFB4F" box="[777,799,1177,1203]" italics="true" pageId="13" pageNumber="298">5s</emphasis>
) and lower segments (
<emphasis id="B912EAC4FFF6FFAAFBDE9CAB8A7DFB4F" box="[1050,1069,1177,1203]" italics="true" pageId="13" pageNumber="298">5i</emphasis>
), row 
<emphasis id="B912EAC4FFF6FFAAFBB69CAB8AD5FB4F" box="[1138,1157,1177,1203]" italics="true" pageId="13" pageNumber="298">5i</emphasis>
going downwards to or near the level of row 
<emphasis id="B912EAC4FFF6FFAAFBB59C8F8A2FFB2B" box="[1137,1151,1213,1239]" italics="true" pageId="13" pageNumber="298">d</emphasis>
, the distance between row 
<emphasis id="B912EAC4FFF6FFAAFC6F9CD38DEEFB07" box="[939,958,1249,1275]" italics="true" pageId="13" pageNumber="298">5i</emphasis>
and row 
<emphasis id="B912EAC4FFF6FFAAFBE29CD28A64FB06" box="[1062,1076,1248,1274]" italics="true" pageId="13" pageNumber="298">d</emphasis>
absent or much smaller than the length of row 
<emphasis id="B912EAC4FFF6FFAAFBF89D378A04FAE3" box="[1084,1108,1285,1311]" italics="true" pageId="13" pageNumber="298">5i,</emphasis>
row 
<emphasis id="B912EAC4FFF6FFAAFD419D1B8CC8FABF" box="[645,664,1321,1347]" italics="true" pageId="13" pageNumber="298">5i</emphasis>
opposite row 
<emphasis id="B912EAC4FFF6FFAAFCF19D1A8D19FABE" box="[821,841,1320,1346]" italics="true" pageId="13" pageNumber="298">6i</emphasis>
, row 
<emphasis id="B912EAC4FFF6FFAAFC419D1A8DC2FABE" box="[901,914,1320,1346]" italics="true" pageId="13" pageNumber="298">6</emphasis>
above and below row 
<emphasis id="B912EAC4FFF6FFAAFD419D7E8CC7FA9A" box="[645,663,1356,1382]" italics="true" pageId="13" pageNumber="298">b,</emphasis>
row 
<emphasis id="B912EAC4FFF6FFAAFD089D7E8CB0FA9A" box="[716,736,1356,1382]" italics="true" pageId="13" pageNumber="298">6i</emphasis>
extending for a few papillae below the level of row 
<emphasis id="B912EAC4FFF6FFAAFCC59D428D5FFA76" box="[769,783,1392,1418]" italics="true" pageId="13" pageNumber="298">d</emphasis>
,row 
<emphasis id="B912EAC4FFF6FFAAFC809D438D00FA77" box="[836,848,1393,1419]" italics="true" pageId="13" pageNumber="298">7</emphasis>
above row 
<emphasis id="B912EAC4FFF6FFAAFC079D428D85FA76" box="[963,981,1392,1418]" italics="true" pageId="13" pageNumber="298">b,</emphasis>
at pore 
<emphasis id="B912EAC4FFF6FFAAFBE09D438A61FA76" box="[1060,1073,1393,1418]" italics="true" pageId="13" pageNumber="298">α</emphasis>
(
<emphasis id="B912EAC4FFF6FFAAFBF99D428A14FA76" box="[1085,1092,1392,1418]" italics="true" pageId="13" pageNumber="298">1</emphasis>
:8‒10, 
<emphasis id="B912EAC4FFF6FFAAFD419DA68CC1FA52" box="[645,657,1428,1454]" italics="true" pageId="13" pageNumber="298">2</emphasis>
: 6‒8, 
<emphasis id="B912EAC4FFF6FFAAFD109DA68C8FFA52" box="[724,735,1428,1454]" italics="true" pageId="13" pageNumber="298">3</emphasis>
: 6‒7, 
<emphasis id="B912EAC4FFF6FFAAFCE59DA68D7DFA52" box="[801,813,1428,1454]" italics="true" pageId="13" pageNumber="298">4</emphasis>
: 7‒9, 
<emphasis id="B912EAC4FFF6FFAAFCAB9DA68D2BFA52" box="[879,891,1428,1454]" italics="true" pageId="13" pageNumber="298">5</emphasis>
: 3 + 5 and 4 + 4, 
<emphasis id="B912EAC4FFF6FFAAFBF49DA68A6DFA52" box="[1072,1085,1428,1454]" italics="true" pageId="13" pageNumber="298">6</emphasis>
: 3 + 10 and 4 + 9, 
<emphasis id="B912EAC4FFF6FFAAFD329D8A8D52FA2E" box="[758,770,1464,1490]" italics="true" pageId="13" pageNumber="298">7</emphasis>
: 1). Row 
<emphasis id="B912EAC4FFF6FFAAFCA19D8A8D23FA2E" box="[869,883,1464,1490]" italics="true" pageId="13" pageNumber="298">d</emphasis>
(18‒22) with supralabial part and the part on cheek connected, ending backward below posterior part of the eye. 
<emphasis id="B912EAC4FFF6FFAAFD419E168D83F9C1" box="[645,979,1572,1598]" italics="true" pageId="13" pageNumber="298">Preoperculo-mandibular rows</emphasis>
: external row 
<emphasis id="B912EAC4FFF6FFAAFD419E7A8CC1F99D" box="[645,657,1608,1633]" italics="true" pageId="13" pageNumber="298">e</emphasis>
(21 + 14–26 + 16) and internal row 
<emphasis id="B912EAC4FFF6FFAAFBED9E7A8A60F99D" box="[1065,1072,1608,1633]" italics="true" pageId="13" pageNumber="298">i</emphasis>
(8 + 7– 10 + 9) divided into anterior and posterior sections, mental row 
<emphasis id="B912EAC4FFF6FFAAFC569EBD8DC9F955" box="[914,921,1679,1705]" italics="true" pageId="13" pageNumber="298">f</emphasis>
longitudinal (4‒7). 
<emphasis id="B912EAC4FFF5FFA9FF4598998F07FF39" box="[129,343,171,197]" italics="true" pageId="14" pageNumber="299">Oculoscapular rows</emphasis>
: anterior longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FF4598E28EC5FF15" box="[129,149,207,233]" italics="true" pageId="14" pageNumber="299">x1</emphasis>
(7–10) ends anteriorly well behind pore 
<emphasis id="B912EAC4FFF5FFA9FDA898E28C29FF16" box="[620,633,208,234]" italics="true" pageId="14" pageNumber="299">β</emphasis>
, posterior longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FE6B98C68F97FEF1" box="[431,455,243,269]" italics="true" pageId="14" pageNumber="299">x2</emphasis>
(3‒4) above the opercular edge, transversal row 
<emphasis id="B912EAC4FFF5FFA9FE23992A8FA3FECD" box="[487,499,280,305]" italics="true" pageId="14" pageNumber="299">z</emphasis>
(5‒7) below the posterior end of anterior oculoscapular canal, ends close to row 
<emphasis id="B912EAC4FFF5FFA9FE5F99528FFFFE85" box="[411,431,351,377]" italics="true" pageId="14" pageNumber="299">x1</emphasis>
, transversal row 
<emphasis id="B912EAC4FFF5FFA9FDB699528CD0FE85" box="[626,640,352,377]" italics="true" pageId="14" pageNumber="299">q</emphasis>
(2‒4) behind pore 
<emphasis id="B912EAC4FFF5FFA9FE9A99B68F39FE61" box="[350,361,388,413]" italics="true" pageId="14" pageNumber="299">Ρ</emphasis>
and below longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FF74999A8EEEFE3D" box="[176,190,424,449]" italics="true" pageId="14" pageNumber="299">u</emphasis>
(2), transversal row 
<emphasis id="B912EAC4FFF5FFA9FE5E999A8FEAFE3D" box="[410,442,424,449]" italics="true" pageId="14" pageNumber="299">trp</emphasis>
(3) behind row 
<emphasis id="B912EAC4FFF5FFA9FDA3999A8C2AFE4E" box="[615,634,415,449]" italics="true" pageId="14" pageNumber="299">
x 
<superScript id="7C139B9EFFF5FFA9FDB199AD8C2AFE4E" attach="right" box="[629,634,415,434]" fontSize="8" pageId="14" pageNumber="299">1</superScript>
</emphasis>
, transversal row 
<emphasis id="B912EAC4FFF5FFA9FEF699F98F6EFE18" box="[306,318,459,484]" italics="true" pageId="14" pageNumber="299">y</emphasis>
(2‒3) below row 
<emphasis id="B912EAC4FFF5FFA9FDC099F98C4CFE19" box="[516,540,459,485]" italics="true" pageId="14" pageNumber="299">x2</emphasis>
. Axillary vertical rows 
<emphasis id="B912EAC4FFF5FFA9FED799DD8F62FDF5" box="[275,306,495,521]" italics="true" pageId="14" pageNumber="299">as1</emphasis>
(5‒8), 
<emphasis id="B912EAC4FFF5FFA9FEBB99DD8FF1FDF5" box="[383,417,495,521]" italics="true" pageId="14" pageNumber="299">as2</emphasis>
(5‒8), and 
<emphasis id="B912EAC4FFF5FFA9FDDA99DD8C10FDF5" box="[542,576,495,521]" italics="true" pageId="14" pageNumber="299">as3</emphasis>
(3‒5) and horizontal rows 
<emphasis id="B912EAC4FFF5FFA9FEBC9A208FC4FDD1" box="[376,404,530,557]" italics="true" pageId="14" pageNumber="299">la1</emphasis>
(2‒3) and 
<emphasis id="B912EAC4FFF5FFA9FDDA9A208C6DFDD1" box="[542,573,530,557]" italics="true" pageId="14" pageNumber="299">la2</emphasis>
(1‒3) present, row 
<emphasis id="B912EAC4FFF5FFA9FECB9A048F7EFDAD" box="[271,302,566,593]" italics="true" pageId="14" pageNumber="299">la2</emphasis>
not visible in female 
<typeStatus id="54DD8874FFF5FFA9FDDC9A058C2DFDAD" box="[536,637,567,593]" pageId="14" pageNumber="299">paratype</typeStatus>
. 
<emphasis id="B912EAC4FFF5FFA9FF449A688F7FFD88" box="[128,303,602,628]" italics="true" pageId="14" pageNumber="299">Opercular rows</emphasis>
: transverse row 
<emphasis id="B912EAC4FFF5FFA9FDC69A698C47FD88" box="[514,535,603,628]" italics="true" pageId="14" pageNumber="299">ot</emphasis>
(16–20), superior longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FE589A4D8FE3FD64" box="[412,435,639,664]" italics="true" pageId="14" pageNumber="299">os</emphasis>
(6‒8), and inferior longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FEFB9A918F03FD40" box="[319,339,675,700]" italics="true" pageId="14" pageNumber="299">oi</emphasis>
(3‒5). 
<emphasis id="B912EAC4FFF5FFA9FE5A9A908C2AFD40" box="[414,634,674,700]" italics="true" pageId="14" pageNumber="299">Anterior dorsal rows</emphasis>
: anterior transverse row 
<emphasis id="B912EAC4FFF5FFA9FE439AF58FC6FD1C" box="[391,406,711,736]" italics="true" pageId="14" pageNumber="299">n</emphasis>
(6‒8) behind pore 
<emphasis id="B912EAC4FFF5FFA9FDAD9AF58CD0FD1C" box="[617,640,711,736]" italics="true" pageId="14" pageNumber="299">ω,</emphasis>
transverse row 
<emphasis id="B912EAC4FFF5FFA9FEEF9AD98F68FCF8" box="[299,312,747,772]" italics="true" pageId="14" pageNumber="299">o</emphasis>
(3‒4) distant to the fellow in the dorsal midline, longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FDE99B3D8C6BFCD4" box="[557,571,783,808]" italics="true" pageId="14" pageNumber="299">g</emphasis>
(3‒4) not extending anteriorly beyond lateral ends row 
<emphasis id="B912EAC4FFF5FFA9FF759B648EEEFC93" box="[177,190,854,879]" italics="true" pageId="14" pageNumber="299">o</emphasis>
, longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FE4E9B648FF0FC93" box="[394,416,854,879]" italics="true" pageId="14" pageNumber="299">m</emphasis>
(2‒3) behind below row 
<emphasis id="B912EAC4FFF5FFA9FF769B488E94FC6F" box="[178,196,890,915]" italics="true" pageId="14" pageNumber="299">g,</emphasis>
longitudinal row 
<emphasis id="B912EAC4FFF5FFA9FE4F9B4B8FC9FC6F" box="[395,409,889,915]" italics="true" pageId="14" pageNumber="299">h</emphasis>
(7‒8) in front of the first dorsal fin anterior origin, continuous in the 
<typeStatus id="54DD8874FFF5FFA9FF6F9BF38F5EFC27" box="[171,270,961,987]" pageId="14" pageNumber="299">holotype</typeStatus>
and female 
<typeStatus id="54DD8874FFF5FFA9FE509BF08FA6FC20" box="[404,502,962,988]" pageId="14" pageNumber="299">paratype</typeStatus>
and divided in the male 
<typeStatus id="54DD8874FFF5FFA9FEE49BD48FD5FBFC" box="[288,389,998,1024]" pageId="14" pageNumber="299">paratype</typeStatus>
. Interorbital papillae absent.
</paragraph>
<paragraph id="8BD936D6FFF5FFA9FF5B9C1F8CA6FA9C" blockId="14.[128,640,171,1706]" lastBlockId="14.[676,1187,166,1376]" pageId="14" pageNumber="299">
<emphasis id="B912EAC4FFF5FFA9FF5B9C1F8F77FBBB" box="[159,295,1069,1095]" italics="true" pageId="14" pageNumber="299">Colouration.</emphasis>
Live colouration not documented. Colour preserved (
<figureCitation id="135D2A53FFF5FFA9FE2E9C638C63FB97" box="[490,563,1105,1131]" captionStart="figure 1" captionStartId="10.[128,195,1556,1578]" captionTargetBox="[223,1092,575,1537]" captionTargetId="figure-3@10.[223,1092,575,1538]" captionTargetPageId="10" captionText="figure 1 Preserved specimens.(A) Zebrus pallaoroi sp.nov., npm P6V144302, holotype, male, 31.81 + 8.51 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro. Photo by M.Kovačić. (B) Zebrus zebrus, nmp P6V 140912,neotype, female, 23.25 + 6.22mm, Îll Gross, Banyuls sur Mer, France (C) Millerigobius macrocephalus, nmp P6V 142686, juvenile of unidentified sex, 14.28 + 3.97 mm, Îll Gross, Banyuls sur Mer, France." figureDoi="http://doi.org/10.5281/zenodo.8343385" httpUri="https://zenodo.org/record/8343385/files/figure.png" pageId="14" pageNumber="299">fig. 1A</figureCitation>
). Head and body pale yellow to white brown (beige to tan), with dotted dark brown markings of larger and smaller melanophores shaping the patterns. Body with ten to eleven vertical dark brown bands present along lateral side, first in front of the first dorsal fin, last at end of the second dorsal fin. The dark vertical bands are at the upper edge about equal or narrower than paler interspaces in-between. Reticulate pattern, formed by dark markings along the scale margins also visible, better in 
<typeStatus id="54DD8874FFF5FFA9FF449E328EA0F9E6" box="[128,240,1536,1562]" pageId="14" pageNumber="299">paratypes</typeStatus>
. Caudal peduncle is more uniformly and less intensively coloured. Dorsal view also striped, no pale saddle shapes on the back. Ventral bodyside, including belly whitish, more dotted in males than in the female. Head variably dotted, with broad pale transverse stripe across anterior nape behind eyes, not including eyes, curving backward and downwards, less recognizable on opercles and well visible again on pectoral fin lobes. Posteriorly to the transverse stripe nape and predorsal area dark dotted, marbled in males, uniformly dotted in the female. The sides of the head marbled in males, uniformly dotted in the female with an anterior oblique dark band from eye to the upper lip. Snout pigmented. Eyes dark, with a grey pupil. The underside of the head and breast pigmented by fine dots in both sexes, with only anterior isthmus paler, especially in the female. Pectoral fin lobe with the broad pale oblique area and dark upper and lower corners. The dark basal pectoral mark short, restricted to the upper six rays, and the pigmented area across the entire base of pectoral rays poorly defined. The rest of the pectoral fin uniformly pigmented in males, only the basal half pigmented in the female. The first dorsal fin entirely dotted, with two hardly recognizable oblique bands of more intensive pigmentation start at the first spine at middle and at the upper part, and going backward and downward from it, the second dorsal and anal fins dotted with no recognizable pattern in both sexes, less intensively at base of rays. Caudal fin more or less uniformly dotted. Pelvic fin uniformly pigmented in males, only basal and inner part and anterior membrane pigmented in the female.
</paragraph>
<paragraph id="8BD936D6FFF5FFA9FD069D428DABFA2E" blockId="14.[676,1187,1392,1706]" pageId="14" pageNumber="299">
<emphasis id="B912EAC4FFF5FFA9FD069D428D62FA76" box="[706,818,1392,1418]" italics="true" pageId="14" pageNumber="299">Osteology.</emphasis>
Vertebrae (including urostyle) 27 (precaudal + caudal vertebrae: 10 + 17); dorsal pterygiophore formula 3-22110.
</paragraph>
<paragraph id="8BD936D6FFF5FFA8FD069DEE8F15FEF4" blockId="14.[676,1187,1392,1706]" lastBlockId="15.[98,610,166,1167]" lastPageId="15" lastPageNumber="300" pageId="14" pageNumber="299">
<emphasis id="B912EAC4FFF5FFA9FD069DEE8D6AFA0A" box="[706,826,1500,1526]" italics="true" pageId="14" pageNumber="299">Etymology.</emphasis>
Named in honour of Armin Pallaoro, a great ichthyologist from the Institute of Oceanography and Fishery Split, 
<collectingCountry id="F3717646FFF5FFA9FD609E758CA8F99D" box="[676,760,1607,1633]" name="Croatia" pageId="14" pageNumber="299">Croatia</collectingCountry>
, who sadly passed away in 
<date id="FFD81016FFF5FFA9FB899E7A8C8DF97A" pageId="14" pageNumber="299" value="2020-01">January 2020</date>
. Armin unselfishly shared his knowledge and his fieldwork and laboratory skills on fishes with generations of younger Croatian colleagues at their beginnings, including one of the authors (mk).
</paragraph>
<paragraph id="8BD936D6FFF4FFA8FF4599258EC1FE19" blockId="15.[98,610,166,1167]" pageId="15" pageNumber="300">
<emphasis id="B912EAC4FFF4FFA8FF4599258E8AFECD" box="[129,218,279,305]" italics="true" pageId="15" pageNumber="300">Ecology.</emphasis>
The 
<typeStatus id="54DD8874FFF4FFA8FEEE992A8F36FECE" box="[298,358,280,306]" pageId="15" pageNumber="300">types</typeStatus>
and the 
<typeStatus id="54DD8874FFF4FFA8FE2A99258E89FEA9" isOtherTypeStatus="true" pageId="15" pageNumber="300">additional specimens</typeStatus>
were found between gravel and small boulders in very shallow infralittoral waters, just by the shore, at a maximum depth of 
<specimenCount id="9D60FD5FFFF4FFA8FF4599958EFDFE3D" box="[129,173,423,449]" pageId="15" pageNumber="300" type="generic">
<quantity id="4C9E9B33FFF4FFA8FF4599958EFAFE3D" box="[129,170,423,449]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="300" unit="m" value="1.0">1 m</quantity>
.
</specimenCount>
This species is a typical cryptobenthic fish.
</paragraph>
<paragraph id="8BD936D6FFF4FFA8FF4599DC8C40FD89" blockId="15.[98,610,166,1167]" pageId="15" pageNumber="300">
<emphasis id="B912EAC4FFF4FFA8FF4599DC8FF2FDF4" box="[129,418,494,520]" italics="true" pageId="15" pageNumber="300">Geographical distribution.</emphasis>
Recorded in the southern part of the Adriatic Sea, the northern part of the Ionian Sea, and the northern and western part of the 
<collectingRegion id="49A2F834FFF4FFA8FEF99A698FC0FD89" box="[317,400,603,629]" country="Greece" name="Voreio Aigaio" pageId="15" pageNumber="300">Aegean</collectingRegion>
Sea (
<figureCitation id="135D2A53FFF4FFA8FE089A688C50FD88" box="[460,512,602,628]" captionStart="figure 4" captionStartId="15.[98,165,1633,1655]" captionTargetBox="[193,1062,1203,1614]" captionTargetId="figure-412@15.[193,1062,1203,1614]" captionTargetPageId="15" captionText="figure 4 Map of sampling localities of Zebrus pallaoroi sp. nov. (circle) and Z.zebrus (square). The type locality of Z.pallaoroi and locality of neotype of Z.zebrus are marked with hatching.The westernmost record of Z.zebrus is indicated by ■. The first record of Millerigobius macrocephalus from Cyprus is indicated by ▲." figureDoi="http://doi.org/10.5281/zenodo.8343391" httpUri="https://zenodo.org/record/8343391/files/figure.png" pageId="15" pageNumber="300">fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BD936D6FFF4FFB7FF459A4C8F68FC27" blockId="15.[98,610,166,1167]" lastBlockId="16.[129,640,171,988]" lastPageId="16" lastPageNumber="301" pageId="15" pageNumber="300">
<emphasis id="B912EAC4FFF4FFA8FF459A4C8EB7FD64" box="[129,231,638,664]" italics="true" pageId="15" pageNumber="300">Remarks.</emphasis>
At present, the genus 
<taxonomicName id="4C664D55FFF4FFA8FDD29A4C8C31FD64" authorityName="de Buen" authorityYear="1930" box="[534,609,638,664]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="genus">
<emphasis id="B912EAC4FFF4FFA8FDD29A4C8C31FD64" box="[534,609,638,664]" italics="true" pageId="15" pageNumber="300">Zebrus</emphasis>
</taxonomicName>
includes just two species: 
<taxonomicName id="4C664D55FFF4FFA8FE4D9A918C57FD40" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[393,519,674,700]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF4FFA8FE4D9A918C57FD40" box="[393,519,674,700]" italics="true" pageId="15" pageNumber="300">Z. pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF4FFA8FDCA9A918C32FD41" box="[526,610,675,701]" pageId="15" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C664D55FFF4FFA8FF509AF58EA8FD1C" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[148,248,710,736]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FF509AF58EA8FD1C" box="[148,248,710,736]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C664D55FFF4FFA8FEC19AF48FE9FD1C" authority="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener, 2021" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[261,441,710,736]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="pallaoroi" status="sp. nov.">
<emphasis id="B912EAC4FFF4FFA8FEC19AF48FE9FD1C" box="[261,441,710,736]" italics="true" pageId="15" pageNumber="300">Zebrus pallaoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157BFFFF4FFA8FE049AF58C44FD1D" box="[448,532,711,737]" pageId="15" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
differs in various characters from 
<taxonomicName id="4C664D55FFF4FFA8FE679AD98C5BFCF8" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[419,523,746,772]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FE679AD98C5BFCF8" box="[419,523,746,772]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
(based on comparative material and 
<bibRefCitation id="EFF74B27FFF4FFA8FE029B3C8C32FCD4" author="Miller, P. J." box="[454,610,782,808]" pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller (1977))</bibRefCitation>
as followed: snout longer than eye, its length 1.1‒-1.2 of eye diameter vs. snout shorter than eye (
<bibRefCitation id="EFF74B27FFF4FFA8FF589B4B8F75FC68" author="Miller, P. J." box="[156,293,889,916]" pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
), its length 0.8‒0.9 of eye diameter in 
<taxonomicName id="4C664D55FFF4FFA8FF299BAC8F00FC4B" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[237,336,925,951]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FF299BAC8F00FC4B" box="[237,336,925,951]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
(comparative material); posterior nostril short tube, 4/5–9/10 of anterior nostril vs. posterior nostril about 1/2 of anterior nostril reported by 
<bibRefCitation id="EFF74B27FFF4FFA8FDDA9C3B8EF6FBBB" author="Miller, P. J." pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller (1977)</bibRefCitation>
, 1/4‒2/5 of anterior nostril found in comparative material of 
<taxonomicName id="4C664D55FFF4FFA8FE499C608FA7FB97" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[397,503,1105,1131]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FE499C608FA7FB97" box="[397,503,1105,1131]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A53FFF4FFA8FDC89C638E28FB73" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="15" pageNumber="300">fig. 2A, D</figureCitation>
); eyes moderately small, eye diameter is 
<quantity id="4C9E9B33FFF4FFA8FD41989E8CA8FF39" box="[645,760,171,198]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.1938" metricValueMin="1.0921999999999998" pageId="15" pageNumber="300" unit="in" value="4.5" valueMax="4.7" valueMin="4.3">4.3‒4.7 in</quantity>
head length vs. eye diameter is 
<quantity id="4C9E9B33FFF4FFA8FD4198FD8CA3FF15" box="[645,755,207,233]" metricMagnitude="-2" metricUnit="m" metricValue="9.525" metricValueMax="10.414" metricValueMin="8.636" pageId="15" pageNumber="300" unit="in" value="3.75" valueMax="4.1" valueMin="3.4">3.4‒4.1 in</quantity>
head length (
<bibRefCitation id="EFF74B27FFF4FFA8FC5A98FD8A78FF15" author="Miller, P. J." box="[926,1064,207,233]" pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
), 
<quantity id="4C9E9B33FFF4FFA8FBFB98FD8CCCFEF1" metricMagnitude="-2" metricUnit="m" metricValue="9.143999999999998" metricValueMax="10.414" metricValueMin="7.874" pageId="15" pageNumber="300" unit="in" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1‒4.1 in</quantity>
head length (comparative material) in 
<taxonomicName id="4C664D55FFF4FFA8FD41992A8CBFFECD" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[645,751,279,305]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FD41992A8CBFFECD" box="[645,751,279,305]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
; left and right ventrolateral head ridges transversally connected on anterior part by short transversal ridge vs. left and right ventrolateral head ridges disconnected anteriorly by midventral flat area in 
<taxonomicName id="4C664D55FFF4FFA8FBE5999A8AD5FE3D" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[1057,1157,423,449]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FBE5999A8AD5FE3D" box="[1057,1157,423,449]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A53FFF4FFA8FD4999F98D4BFE19" box="[653,795,459,485]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="15" pageNumber="300">fig. 2C and F</figureCitation>
) (comparative material, no data in 
<bibRefCitation id="EFF74B27FFF4FFA8FD6899DC8D64FDF5" author="Miller, P. J." box="[684,820,494,521]" pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
, the transversal connection of ventrolateral head ridges should not be confused with the small mental fold placed more anterior on chin below lower lip that is present in 
<taxonomicName id="4C664D55FFF4FFA8FCE29A4D8DF8FD64" authorityName="Cooper &amp; Yamaguchi &amp; Macdonald &amp; Nanova &amp; Yudin &amp; Dugmore &amp; Kitchener" authorityYear="2021" box="[806,936,638,664]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="pallaoroi">
<emphasis id="B912EAC4FFF4FFA8FCE29A4D8DF8FD64" box="[806,936,638,664]" italics="true" pageId="15" pageNumber="300">Z. pallaoroi</emphasis>
</taxonomicName>
and is of variable occurrence in 
<taxonomicName id="4C664D55FFF4FFA8FCE79A918DD4FD40" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[803,900,674,700]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FCE79A918DD4FD40" box="[803,900,674,700]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
; both structures visible in 
<figureCitation id="135D2A53FFF4FFA8FD6F9AF48CABFD1C" box="[683,763,710,736]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="15" pageNumber="300">fig. 2C</figureCitation>
); anterior membrane in midline depth about 2/3 of spinous ray vs. about 1/2 of spinous ray reported in 
<bibRefCitation id="EFF74B27FFF4FFA8FC049B3C8A1FFCD4" author="Miller, P. J." box="[960,1103,782,808]" pageId="15" pageNumber="300" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller (1977)</bibRefCitation>
and 1/4-1/2 of spinous ray found on comparative material of 
<taxonomicName id="4C664D55FFF4FFA8FCC09B648D35FC93" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[772,869,853,879]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FCC09B648D35FC93" box="[772,869,853,879]" italics="true" pageId="15" pageNumber="300">Z.zebrus</emphasis>
</taxonomicName>
(
<figureCitation id="135D2A53FFF4FFA8FCB59B678DABFC8C" box="[881,1019,853,880]" captionStart="figure 2" captionStartId="11.[98,165,1515,1537]" captionTargetBox="[193,1062,520,1497]" captionTargetId="figure-158@11.[193,1062,520,1497]" captionTargetPageId="11" captionText="figure 2 Zebrus pallaoroi sp. nov.nmp P6V 144300, paratype, male, 27.72 + 6.83 mm, Kostanjica, Boka Kotorska, Adriatic Sea, Montenegro: (A) posterior and anterior nostrils; (B) pelvic fin with anterior membrane; (C) ventrolateral head ridges marked with black arrows and transversal connection marked with a grey arrow, small mental fold visible at the lower lip, anteriorly from the transversal connection of ventrolateral ridges. Zebrus zebrus: nmp P6V 142593,male, 21.84 + 5.6 mm, Selce, Kvarner area, Croatia: (D) posterior and anterior nostrils; (E) pelvic fin with anterior membrane; (F) ventrolateral head ridges marked with black arrows. photos by m. kovačiĆ." figureDoi="http://doi.org/10.5281/zenodo.8343387" httpUri="https://zenodo.org/record/8343387/files/figure.png" pageId="15" pageNumber="300">fig. 2B and E</figureCitation>
); head canal pores large, pore 
<emphasis id="B912EAC4FFF4FFA8FC839B488D04FC6F" box="[839,852,890,915]" italics="true" pageId="15" pageNumber="300">α</emphasis>
diameter about half of the distance between pore 
<emphasis id="B912EAC4FFF4FFA8FC4F9BAC8DC6FC4B" box="[907,918,926,951]" italics="true" pageId="15" pageNumber="300">Ρ</emphasis>
and 
<emphasis id="B912EAC4FFF4FFA8FC0A9BAC8DB1FC4B" box="[974,993,925,951]" italics="true" pageId="15" pageNumber="300">Ρ1</emphasis>
vs. head canal pores of moderate size, distance between pore 
<emphasis id="B912EAC4FFF4FFA8FD7A9BD48C99FC03" box="[702,713,998,1023]" italics="true" pageId="15" pageNumber="300">Ρ</emphasis>
and 
<emphasis id="B912EAC4FFF4FFA8FD3A9BD48D41FC03" box="[766,785,997,1023]" italics="true" pageId="15" pageNumber="300">Ρ1</emphasis>
about three times or more longer than pore 
<emphasis id="B912EAC4FFF4FFA8FD3A9C388D5BFBDF" box="[766,779,1034,1059]" italics="true" pageId="15" pageNumber="300">α</emphasis>
diameter (comparative material, no data in 
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) in 
<taxonomicName id="4C664D55FFF4FFA8FC609C1C8A57FBBB" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[932,1031,1069,1095]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="300" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFF4FFA8FC609C1C8A57FBBB" box="[932,1031,1069,1095]" italics="true" pageId="15" pageNumber="300">Z. zebrus</emphasis>
</taxonomicName>
; suborbital sensory papillae row 
<emphasis id="B912EAC4FFF4FFA8FCBA9C638DC1FB97" box="[894,913,1105,1131]" italics="true" pageId="15" pageNumber="300">5i</emphasis>
going downwards to or near the level of row 
<emphasis id="B912EAC4FFF4FFA8FC649C478DFEFB73" box="[928,942,1141,1167]" italics="true" pageId="15" pageNumber="300">d</emphasis>
, distance between row 
<emphasis id="B912EAC4FFEBFFB7FF75989E8E94FF3A" box="[177,196,172,198]" italics="true" pageId="16" pageNumber="301">5i</emphasis>
and row 
<emphasis id="B912EAC4FFEBFFB7FEEF98998F69FF39" box="[299,313,171,197]" italics="true" pageId="16" pageNumber="301">d</emphasis>
absent or much smaller than length of row 
<emphasis id="B912EAC4FFEBFFB7FEDE98E28F7DFF16" box="[282,301,208,234]" italics="true" pageId="16" pageNumber="301">5i</emphasis>
vs. suborbital sensory papillae row 
<emphasis id="B912EAC4FFEBFFB7FF7298C68E99FEF2" box="[182,201,244,270]" italics="true" pageId="16" pageNumber="301">5i</emphasis>
ends downwards distant from row 
<emphasis id="B912EAC4FFEBFFB7FDA898C18C2AFEF1" box="[620,634,243,269]" italics="true" pageId="16" pageNumber="301">d</emphasis>
, row 
<emphasis id="B912EAC4FFEBFFB7FF70992A8E97FECE" box="[180,199,280,306]" italics="true" pageId="16" pageNumber="301">5i</emphasis>
length equal or smaller than distance between row 
<emphasis id="B912EAC4FFEBFFB7FEEA99098F11FEA9" box="[302,321,315,341]" italics="true" pageId="16" pageNumber="301">5i</emphasis>
and row 
<emphasis id="B912EAC4FFEBFFB7FE0E99098F88FEA9" box="[458,472,315,341]" italics="true" pageId="16" pageNumber="301">d</emphasis>
(comparative material, no data in 
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) in 
<taxonomicName id="4C664D55FFEBFFB7FDD299528C2AFE85" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[534,634,351,377]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFEBFFB7FDD299528C2AFE85" box="[534,634,351,377]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
</taxonomicName>
; body with 10‒11 vertical dark brown bands present along lateral side, first in front of the first dorsal fin, last at end of the second dorsal fin, at upper edge about equal or narrower than pale interspaces inbetween vs. 6‒9 dark vertical stripes at upper edge, broader or equal than pale interspaces in 
<taxonomicName id="4C664D55FFEBFFB7FE719A698C70FD88" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[437,544,602,628]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFEBFFB7FE719A698C70FD88" box="[437,544,602,628]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF74B27FFEBFFB7FDF29A688EE0FD64" author="Miller, P. J." pageId="16" pageNumber="301" pagination="339 - 362" refId="ref16111" refString="Miller, P. J. (1977) Gobies from Rhodes and systematic features of Zebrus zebrus (Teleostei: Gobiidae). Zool. J. Linn. Soc., 60, 339 - 362." type="journal article" year="1977">Miller, 1977</bibRefCitation>
and comparative material). In addition to these differences, the new species has a longer and more slender body which results in body depth at pelvic-fin origin 5.3 (5.4) in sl vs. body depth at pelvic-fin origin 
<quantity id="4C9E9B33FFEBFFB7FDE29B3C8CD0FCD4" box="[550,640,782,808]" metricMagnitude="-1" metricUnit="m" metricValue="1.1049" metricValueMax="1.1938" metricValueMin="1.016" pageId="16" pageNumber="301" unit="in" value="4.35" valueMax="4.7" valueMin="4.0">4‒4.7 in</quantity>
sl in the comparative material of 
<taxonomicName id="4C664D55FFEBFFB7FDD09B018C2AFCB0" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[532,634,818,844]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFEBFFB7FDD09B018C2AFCB0" box="[532,634,818,844]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
</taxonomicName>
. However, 
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reported a different range of body depth at the pelvic-fin origin in sl for 
<taxonomicName id="4C664D55FFEBFFB7FF0E9BAC8F7DFC4B" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[202,301,925,951]" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="301" phylum="Chordata" rank="species" species="zebrus">
<emphasis id="B912EAC4FFEBFFB7FF0E9BAC8F7DFC4B" box="[202,301,925,951]" italics="true" pageId="16" pageNumber="301">Z. zebrus</emphasis>
</taxonomicName>
(4.4‒5.5) which overlaps with the new species.
</paragraph>
</subSubSection>
</treatment>
</document>