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178 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.188.2976" ID-GBIF-Dataset="92260542-ac3b-49de-bfe2-3d042357372e" ID-PMC="PMC3345898" ID-Pensoft-Pub="1313-2970-188-1" ID-PubMed="22611332" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-188-1" ModsDocOrigin="ZooKeys 188" ModsDocTitle="Review of the Eustrophinae (Coleoptera, Tetratomidae) of America north of Mexico" checkinTime="1451249074049" checkinUser="pensoft" docAuthor="Pollock, Darren A." docDate="2012" docId="D46B547DF2F442E1E6B945D97C36ED6C" docLanguage="en" docName="ZooKeys 188: 1-153" docOrigin="ZooKeys 188" docSource="http://dx.doi.org/10.3897/zookeys.188.2976" docTitle="Eustrophus tomentosus Say 1826" docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="F26BFFACB84EC224FF90FFB4FC46274C" masterDocTitle="Review of the Eustrophinae (Coleoptera, Tetratomidae) of America north of Mexico" masterLastPageNumber="153" masterPageNumber="1" pageNumber="12" updateTime="1668153814780" updateUser="ExternalLinkService">
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<mods:title>Review of the Eustrophinae (Coleoptera, Tetratomidae) of America north of Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pollock, Darren A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>188</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>153</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.188.2976</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.188.2976</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-188-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152035370" LSID="urn:lsid:plazi:treatment:D46B547DF2F442E1E6B945D97C36ED6C" httpUri="http://treatment.plazi.org/id/D46B547DF2F442E1E6B945D97C36ED6C" lastPageId="13" lastPageNumber="13" pageId="11" pageNumber="12">
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<subSubSection pageId="11" pageNumber="12" type="nomenclature">
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName LSID="http://species-id.net/wiki/Eustrophus_tomentosus" authority="Say 1826" authorityYear="1826" class="Insecta" family="Tetratomidae" genus="Eustrophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrophus tomentosus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tomentosus">Eustrophus tomentosus Say 1826</taxonomicName>
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Figures 41729415067
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="reference_group">
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Insecta" family="Mycetophagidae" genus="Mycetophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mycetophagus niger" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niger">Mycetophagus niger</taxonomicName>
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<bibRefCitation pageId="11" pageNumber="12">Melsheimer 1806</bibRefCitation>
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: 14 [Catalogue];
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<bibRefCitation author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="39" pageNumber="40" pagination="53 - 66" title="Descriptions of new species of Coleoptera of the United States." volume="3" year="1846">Melsheimer 1846</bibRefCitation>
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: 58.
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</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Insecta" family="Mycetophagidae" genus="Mycetophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mycetophagus tomentosus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tomentosus">Mycetophagus tomentosus</taxonomicName>
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<bibRefCitation pageId="11" pageNumber="12">Melsheimer 1806</bibRefCitation>
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: 14 [catalogue].-(
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<normalizedToken originalValue="“Pennsylvania”">"Pennsylvania"</normalizedToken>
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);
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<bibRefCitation author="Say, T" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="40" pageNumber="41" pagination="237 - 284" title="Descriptions of new species of coleopterous insects inhabiting the United States (continued)." volume="5" year="1826">Say 1826</bibRefCitation>
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: 239.
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</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Insecta" family="Tetratomidae" genus="Eustrophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrophus niger" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niger">Eustrophus niger</taxonomicName>
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<bibRefCitation author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="39" pageNumber="40" pagination="53 - 66" title="Descriptions of new species of Coleoptera of the United States." volume="3" year="1846">Melsheimer 1846</bibRefCitation>
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: 58.-
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<bibRefCitation pageId="11" pageNumber="12">Melsheimer 1853</bibRefCitation>
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: 143 (syn.);
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<bibRefCitation author="Leng, CW" journalOrPublisher="John D. Sherman, Jr., Mt. Vernon" pageId="39" pageNumber="40" title="Catalogue of the Coleoptera of America, north of Mexico." year="1920">Leng 1920</bibRefCitation>
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: 238;
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<bibRefCitation pageId="11" pageNumber="12">LeSage 1991</bibRefCitation>
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: 246;
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<bibRefCitation pageId="11" pageNumber="12">Poole and Gentili 1996</bibRefCitation>
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: 299.
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</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Insecta" family="Tetratomidae" genus="Eustrophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrophus tomentosus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tomentosus">Eustrophus tomentosus</taxonomicName>
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<bibRefCitation author="Say, T" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="40" pageNumber="41" pagination="237 - 284" title="Descriptions of new species of coleopterous insects inhabiting the United States (continued)." volume="5" year="1826">Say 1826</bibRefCitation>
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: 239.-U.S.A., Illinois (no specific localities given in description);
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<bibRefCitation pageId="11" pageNumber="12">Melsheimer 1853</bibRefCitation>
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: 143;
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<bibRefCitation pageId="11" pageNumber="12">Crotch 1873</bibRefCitation>
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: 112;
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<bibRefCitation author="Provancher, L" journalOrPublisher="Volume I - les Coleopteres. Darveau, Quebec City" pageId="40" pageNumber="41" title="Petite Faune Entomologique du Canada, precedee d'un traite elementaire d'entomologie." year="1877">Provancher 1877</bibRefCitation>
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: 466;
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<bibRefCitation author="Henshaw, S" journalOrPublisher="The Canadian Entomologist" pageId="39" pageNumber="40" title="List of the Coleoptera of America, north of Mexico. Entomological Society of America, Philadelphia." url="10.5962/bhl.title.13151" year="1885">Henshaw 1885</bibRefCitation>
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: 124;
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<bibRefCitation author="Horn, GH" journalOrPublisher="Transactions of the American Entomological Society" pageId="39" pageNumber="40" pagination="26 - 48" title="Miscellaneous coleopterous studies." volume="15" year="1888">Horn 1888</bibRefCitation>
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: 35;
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<bibRefCitation pageId="11" pageNumber="12">Blatchley 1910</bibRefCitation>
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: 1293;
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<bibRefCitation author="Leng, CW" journalOrPublisher="John D. Sherman, Jr., Mt. Vernon" pageId="39" pageNumber="40" title="Catalogue of the Coleoptera of America, north of Mexico." year="1920">Leng 1920</bibRefCitation>
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: 238;
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<bibRefCitation pageId="11" pageNumber="12">Csiki 1924</bibRefCitation>
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:10;
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<bibRefCitation pageId="11" pageNumber="12">Hatch 1965</bibRefCitation>
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: 66;
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<bibRefCitation pageId="11" pageNumber="12">LeSage 1991</bibRefCitation>
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: 246;
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<bibRefCitation pageId="11" pageNumber="12">Poole and Gentili 1996</bibRefCitation>
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: 299;
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<bibRefCitation pageId="11" pageNumber="12">Young and Pollock 2002</bibRefCitation>
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: 416;
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<bibRefCitation author="Majka, CG" journalOrPublisher="Zootaxa" pageId="39" pageNumber="40" pagination="45 - 68" title="Understanding saproxylic beetles: new records of Tetratomidae, Melandryidae, Synchroidae, and Scraptiidae from the Maritime Provinces of Canada (Coleoptera: Tenebrionoidea)." volume="1248" year="2006">Majka and Pollock 2006</bibRefCitation>
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: 53;
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<bibRefCitation author="Pollock, DA" journalOrPublisher="ZooKeys" pageId="39" pageNumber="40" pagination="261 - 290" title="Review of the Canadian Eustrophinae (Coleoptera, Tetratomidae)." url="10.3897/zookeys.2.30" volume="2" year="2008">Pollock 2008</bibRefCitation>
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: 273, 290;
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<bibRefCitation pageId="11" pageNumber="12">Majka and Pollock 2010</bibRefCitation>
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: 455.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="diagnosis">
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<paragraph pageId="11" pageNumber="12">Diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">The following combination of characters is diagnostic for this species: widely separated eyes; antennal sensilla interrupted, not completely annular; dorsal setae distinctly golden to brown; meso- and metatibiae with oblique ridges.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="description">
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<paragraph pageId="11" pageNumber="12">Description</paragraph>
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<paragraph pageId="11" pageNumber="12">
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(from
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<bibRefCitation author="Pollock, DA" journalOrPublisher="ZooKeys" pageId="39" pageNumber="40" pagination="261 - 290" title="Review of the Canadian Eustrophinae (Coleoptera, Tetratomidae)." url="10.3897/zookeys.2.30" volume="2" year="2008">Pollock 2008</bibRefCitation>
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: 273). TL 4.5-6.0 mm; GEW 2.1-3.0 mm. Body oval, parallel-sided (Fig. 4), distinctly convex dorsally (Fig. 17); dorsal color dark brown, with golden sheen due to dense pubescence; venter and antennae uniformly dark rufous, lighter than dorsal color; dorsal pubescence relatively short, but dense, giving distinct sheen (almost iridescent); eyes widely separated (space ~1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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length of first antennomere), inner margins deeply emarginate (Fig. 29); antennomeres 2-11 only slightly but evenly widened to apex, without distinct change in size between any 2 adjacent antennomeres; distal antennomeres subtriangular to nearly quadrate; antennal sensilla not completely annular, present on short sides of antennomeres only; last maxillary palpomere slightly securiform; prosternal process (Fig. 41) triangular, narrowed distally, extended to slightly short of posterior margin of procoxae; prothoracic episternal suture absent (Fig. 50); elytral punctation relatively fine, punctures arranged in longitudinal striae; meso- and metatibiae with oblique ridges present.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="distribution">
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<paragraph pageId="11" pageNumber="12">Distribution</paragraph>
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<paragraph lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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(Fig. 67). The distribution of this species is virtually transcontinental, with a gap in the interior; for example, no specimens are known from Saskatch
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<pageBreakToken pageId="12" pageNumber="13" start="start">ewan</pageBreakToken>
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and Alberta. Most records from the United States are eastern, but specimens are also known from the Pacific Northwest, California, and Arizona. The 983 specimens examined in this study are from the following: CANADA: BRITISH COLUMBIA, MANITOBA, NOVA SCOTIA, ONTARIO, QUEBEC. UNITED STATES: ALABAMA: Lee, Madison, Marion, Mobile. ARIZONA: Coconino, Pima. ARKANSAS: Crawford, Faulkner, Fulton, Logan, Polk, Pulaski. CALIFORNIA: Butte, Calaveras, El Dorado, Placer, Yuba. COLORADO. CONNECTICUT: Litchfield. DELAWARE: Sussex. DISTRICT OF COLUMBIA. FLORIDA: Dixie, Monroe. GEORGIA: Bartow, Clarke, Echols. IDAHO: Boundary, Clearwater, Latah. IOWA: Johnson, Keokuk, Story. ILLINOIS: Bond, Champaign, Clark, Coles, Cook, LaSalle, McHenry, St. Clair, Union, Wabash. INDIANA: Montgomery, Parke, Porter, Tippecanoe, Vigo. KANSAS: Douglas. KENTUCKY: Butler, Henderson. LOUISIANA: Natchitoches. MARYLAND: Anne Arundel, Charles, Dorchester, Montgomery, Prince
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<normalizedToken originalValue="George’s">George's</normalizedToken>
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, Talbot. MASSACHUSETTS. MICHIGAN: Berrien, Charlevoix, Fillmore, Gogebic, Ingham, Kalamazoo, Kent, Lapeer, Leelanau, Livingston, Macomb, Midland, Monroe, Oakland, St. Joseph, Shiawassee, Washtenaw, Wayne. MINNESOTA: Crow Wing, Hennepin, Sherburne, Washington. MISSISSIPPI: George, Pearl River. MISSOURI: Boone, Dent, Franklin, Randolph, St. Charles. MONTANA: Dawson, Lake, Lincoln, Missoula, Ravalli, Rosebud, Sanders. NEBRASKA: Douglas, Sarpy. NEW HAMPSHIRE: Carroll. NEW JERSEY: Burlington, Gloucester. NEW MEXICO: San Miguel. NEW YORK: Erie, Essex, Genesee, Niagara, Onondaga, Orleans, St. Lawrence, Tompkins. NORTH CAROLINA: Durham, Moore, Stokes, Wake. NORTH DAKOTA: Pembina, Richland. OHIO: Athens, Butler, Clinton, Delaware, Hocking, Ottawa, Portage, Scioto, Vinton, Wayne, Wyandot. OKLAHOMA: Latimer, Oklahoma. OREGON: Benton, Crook, Douglas, Harney, Jackson, Lane, Linn, Multnomah, Wallowa, Yamhill. PENNSYLVANIA: Allegheny, Tioga, Westmoreland. SOUTH CAROLINA: Anderson, Florence, Pickens. TENNESSEE: Shelby. TEXAS: Angelina, Brewster, Montgomery, Sabine, Walker. UTAH: Washington. VERMONT: Franklin. VIRGINIA: Alexandria, Hampton, Nelson, Shenandoah. WASHINGTON: Walla Walla. WEST VIRGINIA: Braxton, Doddridge, Greenbrier, Hampshire, Jackson, Kanawha, Lincoln, Mason, Mineral, Nicholas, Pendleton, Randolph, Ritchie, Roane, Taylor, Webster, Wirt, Wood. WISCONSIN: Dane, Racine, Sauk, Washington. (Complete label data given in Appendix 1).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="types">
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<paragraph pageId="12" pageNumber="13">Types.</paragraph>
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<paragraph pageId="12" pageNumber="13">
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<taxonomicName class="Insecta" family="Tetratomidae" genus="Eustrophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrophus tomentosus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tomentosus">Eustrophus tomentosus</taxonomicName>
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Say. NEOTYPE (designated by
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<bibRefCitation author="Pollock, DA" journalOrPublisher="ZooKeys" pageId="39" pageNumber="40" pagination="261 - 290" title="Review of the Canadian Eustrophinae (Coleoptera, Tetratomidae)." url="10.3897/zookeys.2.30" volume="2" year="2008">Pollock 2008</bibRefCitation>
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), sex unknown, labeled "Ill. / E. tomentosus Say. niger Mels.", in MCZ (LeConte collection).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="natural history">
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<paragraph pageId="12" pageNumber="13">Natural history.</paragraph>
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<paragraph lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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Label data:
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
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(BC), from fungus (BC), evening flight, 4.ix (BC), ex
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus trichocarpa" order="Malpighiales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="trichocarpa">Populus trichocarpa</taxonomicName>
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(BC), in bark
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Larix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Larix occidentalis" order="Pinales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="occidentalis">Larix occidentalis</taxonomicName>
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(BC), fleshy fungus on tree (ON), under bark of pine (ON), under wet moldy bark on dead tree (ON), elm (QC), uv light trap (QC), cut surface of stump (QC),
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<taxonomicName class="Magnoliopsida" family="Ulmaceae" genus="Ulmus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ulmus americana" order="Rosales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="americana">Ulmus americana</taxonomicName>
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(QC), under bark of pine (AL),
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<taxonomicName class="Agaricomycetes" family="Meruliaceae" genus="Spongipellis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spongipellis unicolor" order="Polyporales" pageId="12" pageNumber="13" phylum="Basidiomycota" rank="species" species="unicolor">Spongipellis unicolor</taxonomicName>
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(AR),
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<taxonomicName genus="Climacodes" lsidName="Climacodes septentrionale" pageId="12" pageNumber="13" rank="species" species="septentrionale">Climacodes septentrionale</taxonomicName>
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(AR),
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<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Trametes" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Trametes versicolor" order="Polyporales" pageId="12" pageNumber="13" phylum="Basidiomycota" rank="species" species="versicolor">Trametes versicolor</taxonomicName>
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(AR), under pine bark (CA), Lindgren trap with turpentine bait (CA), on
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus ponderosa" order="Pinales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="ponderosa">Pinus ponderosa</taxonomicName>
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(CA), ex.
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<taxonomicName genus="Polyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyporus" pageId="12" pageNumber="13" phylum="Platyhelminthes" rank="genus">Polyporus</taxonomicName>
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fungus (FL), uv light in oak-maple forest (IA), fungus on dead pine (ID), under bark of old pine stump (MD), in
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<taxonomicName class="Mammalia" family="Cricetidae" genus="Peromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peromyscus" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
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<pageBreakToken pageId="13" pageNumber="14" start="start">Peromyscus</pageBreakToken>
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</taxonomicName>
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nest debris under bark of dead standing
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<taxonomicName class="Magnoliopsida" family="Magnoliaceae" genus="Liriodendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Liriodendron" order="Magnoliales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Liriodendron</taxonomicName>
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(MD), fungus on oak (MN), funnel traps (MT), at black light, vi.1970 (NC), malaise trap, viii-ix (NE), window trap, 8-14.vi (NH), light trap, vi (NY), Lindgren funnel trap, alpha-pinene and ethanol lure (OR), stale molasses trap (SC), at wound on oak trunk (SC), malaise in mature hardwood forest, vi (SC), under bark of old dead decid. tree (VT), ex dead oak stump (WI), under bark of chestnut oak (WV). According to
|
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<bibRefCitation pageId="13" pageNumber="14">Chantal (1985)</bibRefCitation>
|
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,
|
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<taxonomicName class="Insecta" family="Tetratomidae" genus="Eustrophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrophus tomentosus" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tomentosus">Eustrophus tomentosus</taxonomicName>
|
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are found under bark of dead trees, especially
|
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<taxonomicName class="Insecta" family="Tingidae" genus="Ulmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulmus americanus" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="americanus">Ulmus americanus</taxonomicName>
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, as well as at sap exudations.
|
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</paragraph>
|
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</subSubSection>
|
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</treatment>
|
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</document>
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