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<document id="8E427616D14E44A8E7378A889DF32EF3" ID="10.11646/zootaxa.4144.1.11" ID-DOI="10.11646/zootaxa.4144.1.11" ID-GBIF-Dataset="a9c69f07-cad1-4d84-9d2d-884dd8666b25" ID-ISSN="1175-5326" ID-Zenodo-Dep="258383" ID-ZooBank="D5A12B91-7EE7-4A92-89CE-432D150D8C3F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469516089951" checkinUser="plazi" docAuthor="Takahashi, Naoki, Bocak, Ladislav &amp; Abd, Idris" docDate="2016" docId="785D87FFFFC4FFE7FFE28EF746A2FE67" docLanguage="en" docName="zootaxa.4144.1.11.pdf" docOrigin="Zootaxa 4144 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alyculus malaypeninsularis Takahashi, Bocak &amp; Abd, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="148" masterDocId="8464FF87FFC5FFE4FF758D19422DFF9F" masterDocTitle="Discovery of a new species of the brachyelytrous net-winged beetle genus Alyculus (Coleoptera: Lycidae) from Peninsular Malaysia" masterLastPageNumber="150" masterPageNumber="145" pageNumber="146" updateTime="1698366193869" updateUser="plazi">
<mods:mods id="48A9BD2C8855C5942018D3FE38C91C49" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90867A3BD97D3E732A883586E4399A6C">
<mods:title id="ECA5510F12C63A12653E8D399E8F8DE7">Discovery of a new species of the brachyelytrous net-winged beetle genus Alyculus (Coleoptera: Lycidae) from Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="101C95898884E8315562EAFB4D55E834" type="personal">
<mods:role id="7AF9227DEB1789320E0F9E765FB2F7AD">
<mods:roleTerm id="C50E9B0F2A1BC7602CD0D7F3EEBD584F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0304FB5630D9BED5650C19D45A271472">Takahashi, Naoki</mods:namePart>
</mods:name>
<mods:name id="C3388119B6FBD3433564E0CC02F486E1" type="personal">
<mods:role id="951C61F2F7D9591DEB71D927911C99E6">
<mods:roleTerm id="EB15783E61DF85E48B70AFAB2445113B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C0B0C8C36D911A18ABB0CEFEB09A6F6">Bocak, Ladislav</mods:namePart>
</mods:name>
<mods:name id="772C2579A53771BC8D239237DF63E5D7" type="personal">
<mods:role id="B19E543C19819BC97E8DBF19D236D59A">
<mods:roleTerm id="F309C83B6695CCFA736A7F9D8312C030">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB4E9071EA7BE945B3DEC2803CAD90FC">Abd, Idris</mods:namePart>
</mods:name>
<mods:typeOfResource id="28CB558904C4A37436B3DFD351AD9180">text</mods:typeOfResource>
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<mods:title id="67B0443CAB79DA4FA5FE937E9A7E9C77">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1ADE6D80587B9FB98AB0C8EE7A40216E">
<mods:date id="469944D63E016B05FAC985C348D2F1FA">2016</mods:date>
<mods:detail id="3E813C44E2977BC1FDF29CCDD8734676" type="volume">
<mods:number id="53E16E8BD19B5D5204BAFAFDA481FA27">4144</mods:number>
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<mods:number id="85CC6C9DD1315A89F3EA3021D8CA2280">1</mods:number>
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<mods:classification id="CFC42C0C9006413C78FF6532F0B12EFA">journal article</mods:classification>
<mods:identifier id="3B6E8CD828ABE693F55D34FA0B861112" type="DOI">10.11646/zootaxa.4144.1.11</mods:identifier>
<mods:identifier id="372CA6D8D4031424C47A199F6B36527B" type="GBIF-Dataset">a9c69f07-cad1-4d84-9d2d-884dd8666b25</mods:identifier>
<mods:identifier id="3C524A4FE6F7C2BD2A970553AC135478" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="418D8996822F5729F65898002FC77CE6" type="Zenodo-Dep">258383</mods:identifier>
<mods:identifier id="9366B4F1A3E30F48F983D42BBBBF13CD" type="ZooBank">D5A12B91-7EE7-4A92-89CE-432D150D8C3F</mods:identifier>
</mods:mods>
<treatment id="785D87FFFFC4FFE7FFE28EF746A2FE67" ID-DOI="http://doi.org/10.5281/zenodo.6091795" ID-GBIF-Taxon="123207667" ID-Zenodo-Dep="6091795" LSID="urn:lsid:plazi:treatment:785D87FFFFC4FFE7FFE28EF746A2FE67" httpUri="http://treatment.plazi.org/id/785D87FFFFC4FFE7FFE28EF746A2FE67" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146">
<subSubSection id="B8EE6562FFC4FFE5FFE28EF7433FFBB5" pageId="1" pageNumber="146" type="nomenclature">
<paragraph id="F04B36E9FFC4FFE5FFE28EF74061FB97" blockId="1.[151,588,1006,1066]" box="[151,588,1006,1032]" pageId="1" pageNumber="146">
<heading id="AB038185FFC4FFE5FFE28EF74061FB97" bold="true" box="[151,588,1006,1032]" fontSize="11" level="1" pageId="1" pageNumber="146" reason="1">
<emphasis id="C280EAFBFFC4FFE5FFE28EF74061FB97" bold="true" box="[151,588,1006,1032]" pageId="1" pageNumber="146">
<taxonomicName id="37F44D6AFFC4FFE5FFE28EF743CAFB97" box="[151,487,1006,1032]" class="Insecta" family="Lycidae" genus="Alyculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="malaypeninsularis" status="sp. nov.">
<emphasis id="C280EAFBFFC4FFE5FFE28EF743CAFB97" bold="true" box="[151,487,1006,1032]" italics="true" pageId="1" pageNumber="146">Alyculus malaypeninsularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9B35780FFC4FFE5FE9B8EF74061FB97" box="[494,588,1006,1032]" pageId="1" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F04B36E9FFC4FFE5FFE28908433FFBB5" blockId="1.[151,588,1006,1066]" box="[151,274,1041,1066]" pageId="1" pageNumber="146">
(
<figureCitation id="68CF2A6CFFC4FFE5FFEA89084325FBB5" box="[159,264,1041,1066]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURE 4" captionStart-2="FIGURES 5 – 6" captionStartId-0="2.[151,264,1078,1101]" captionStartId-1="3.[151,250,1933,1955]" captionStartId-2="4.[151,264,1381,1404]" captionTargetBox-0="[185,1436,200,1077]" captionTargetBox-1="[500,1071,566,1869]" captionTargetBox-2="[151,1435,878,1360]" captionTargetId-0="figure@2.[150,1436,181,1081]" captionTargetId-1="figure@3.[471,1086,559,1876]" captionTargetId-2="figure@4.[151,1435,878,1360]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." captionText-1="FIGURE 4. Aedeagus of Alyculus malaypeninsularis sp. nov., dorsal view. Scale: 0.1 mm." captionText-2="FIGURES 5 – 6. Alyculus malaypeninsularis sp. nov. 5, habitus on a sweeping net in the field; 6, habitat of A. malaypeninsularis sp. nov. in the Genting Highland, Pahang, Malaysia." httpUri-0="https://zenodo.org/record/258384/files/figure.png" httpUri-1="https://zenodo.org/record/258385/files/figure.png" httpUri-2="https://zenodo.org/record/258386/files/figure.png" pageId="1" pageNumber="146">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC4FFE5FFE28940471DFA9D" pageId="1" pageNumber="146" type="materials_examined">
<paragraph id="F04B36E9FFC4FFE5FFE28940471DFA9D" blockId="1.[151,1437,1113,2038]" pageId="1" pageNumber="146">
<materialsCitation id="409C3CB4FFC4FFE5FFE2894047ACFB09" ID-GBIF-Occurrence="1288049049" collectingDate="2003-07-13" collectingDateMax="2005-03-13" collectingDateMin="2003-07-13" collectionCode="UKMB, ELKU" collectorName="N. Takahashi" country="Malaysia" location="Gunung Brinchang" municipality="Cameron Highlands" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="C280EAFBFFC4FFE5FFE289404369FBED" bold="true" box="[151,324,1113,1138]" pageId="1" pageNumber="146">
<typeStatus id="2F4F884BFFC4FFE5FFE2894042FFFBED" box="[151,210,1113,1138]" pageId="1" pageNumber="146">Type</typeStatus>
material.
</emphasis>
<typeStatus id="2F4F884BFFC4FFE5FE3F8940439FFBED" box="[330,434,1113,1138]" pageId="1" pageNumber="146" type="holotype">Holotype</typeStatus>
(deposited in
<collectionCode id="96E5AE2CFFC4FFE5FD2589434085FBED" box="[592,680,1114,1138]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14454" name="Universiti Kebangsaan Malaysia" pageId="1" pageNumber="146">UKMB</collectionCode>
), male,
<collectingCountry id="88E37679FFC4FFE5FD8A89404144FBED" box="[767,873,1113,1138]" name="Malaysia" pageId="1" pageNumber="146">Malaysia</collectingCountry>
,
<collectingRegion id="3230F80BFFC4FFE5FC06894041E5FBED" box="[883,968,1113,1138]" country="Malaysia" name="Pahang" pageId="1" pageNumber="146">Pahang</collectingRegion>
,
<collectingMunicipality id="102FAC93FFC4FFE5FCA78943469BFBED" box="[978,1206,1113,1138]" pageId="1" pageNumber="146">Cameron Highlands</collectingMunicipality>
,
<location id="F52B6032FFC4FFE5FBB5894347B5FBED" LSID="urn:lsid:plazi:treatment:785D87FFFFC4FFE7FFE28EF746A2FE67:F52B6032FFC4FFE5FBB5894347B5FBED" box="[1216,1432,1113,1138]" country="Malaysia" municipality="Cameron Highlands" name="Gunung Brinchang" pageId="1" pageNumber="146" stateProvince="Pahang">Gunung Brinchang</location>
,
<date id="844A1029FFC4FFE5FFE28964433FFB09" box="[151,274,1149,1174]" pageId="1" pageNumber="146" value="2005-03-13">
<collectingDate id="940EE9C1FFC4FFE5FFE28964433FFB09" box="[151,274,1149,1174]" pageId="1" pageNumber="146" value="2005-03-13">13.iii.2005</collectingDate>
</date>
,
<collectorName id="5D01533FFFC4FFE5FE5589674398FB09" box="[288,437,1149,1174]" pageId="1" pageNumber="146">N. Takahashi</collectorName>
leg.
<typeStatus id="2F4F884BFFC4FFE5FE858967404DFB09" box="[496,608,1150,1174]" pageId="1" pageNumber="146" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="E6F2FD60FFC4FFE5FD1889674097FB09" box="[621,698,1149,1174]" pageId="1" pageNumber="146" type="male">1 male</specimenCount>
(
<collectionCode id="96E5AE2CFFC4FFE5FDBE89674136FB09" box="[715,795,1150,1174]" pageId="1" pageNumber="146">ELKU</collectionCode>
), same locality and collector as
<typeStatus id="2F4F884BFFC4FFE5FBF9896446DCFB09" box="[1164,1265,1149,1174]" pageId="1" pageNumber="146" type="holotype">holotype</typeStatus>
,
<date id="844A1029FFC4FFE5FB8B896447ACFB09" box="[1278,1409,1149,1174]" pageId="1" pageNumber="146" value="2003-07-13">
<collectingDate id="940EE9C1FFC4FFE5FB8B896447ACFB09" box="[1278,1409,1149,1174]" pageId="1" pageNumber="146" value="2003-07-13">13.vii.2003</collectingDate>
</date>
</materialsCitation>
; 1 male (LMBC),
<date id="844A1029FFC4FFE5FE3189B84392FB25" box="[324,447,1185,1210]" pageId="1" pageNumber="146" value="2005-03-11">11.iii.2005</date>
.
<materialsCitation id="409C3CB4FFC4FFE5FEBC89BB4372FB41" ID-GBIF-Occurrence="1288049050" collectingDate="2006-01-19" collectionCode="ELKU" collectorName="N. Takahashi" country="Malaysia" location="Gunung Berembun" municipality="Cameron Highlands" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<specimenCount id="E6F2FD60FFC4FFE5FEBC89BB4039FB25" box="[457,532,1185,1210]" pageId="1" pageNumber="146" type="male">1 male</specimenCount>
(
<collectionCode id="96E5AE2CFFC4FFE5FD5689BB405EFB25" box="[547,627,1186,1210]" pageId="1" pageNumber="146">ELKU</collectionCode>
),
<collectingCountry id="88E37679FFC4FFE5FDF389B840DDFB25" box="[646,752,1185,1210]" name="Malaysia" pageId="1" pageNumber="146">Malaysia</collectingCountry>
,
<collectingRegion id="3230F80BFFC4FFE5FD8F89B84162FB25" box="[762,847,1185,1210]" country="Malaysia" name="Pahang" pageId="1" pageNumber="146">Pahang</collectingRegion>
,
<collectingMunicipality id="102FAC93FFC4FFE5FC2F89BB4613FB25" box="[858,1086,1185,1210]" pageId="1" pageNumber="146">Cameron Highlands</collectingMunicipality>
,
<location id="F52B6032FFC4FFE5FB3D89BB470CFB25" LSID="urn:lsid:plazi:treatment:785D87FFFFC4FFE7FFE28EF746A2FE67:F52B6032FFC4FFE5FB3D89BB470CFB25" box="[1096,1313,1185,1210]" country="Malaysia" municipality="Cameron Highlands" name="Gunung Berembun" pageId="1" pageNumber="146" stateProvince="Pahang">Gunung Berembun</location>
,
<date id="844A1029FFC4FFE5FA5989B847B5FB25" box="[1324,1432,1185,1210]" pageId="1" pageNumber="146" value="2006-01-19">
<collectingDate id="940EE9C1FFC4FFE5FA5989B847B5FB25" box="[1324,1432,1185,1210]" pageId="1" pageNumber="146" value="2006-01-19">19.i.2006</collectingDate>
</date>
,
<collectorName id="5D01533FFFC4FFE5FFE289DF4303FB41" box="[151,302,1221,1246]" pageId="1" pageNumber="146">N. Takahashi</collectorName>
leg
</materialsCitation>
.
<materialsCitation id="409C3CB4FFC4FFE5FE1889DF4712FB41" ID-GBIF-Occurrence="1288049048" box="[365,1343,1221,1246]" collectingDate="2006-01-12" collectionCode="UKMB" collectorName="N. Takahashi" country="Malaysia" location="Genting Highland" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<specimenCount id="E6F2FD60FFC4FFE5FE1889DF4391FB41" box="[365,444,1221,1246]" pageId="1" pageNumber="146" type="male">1 male</specimenCount>
(
<collectionCode id="96E5AE2CFFC4FFE5FEBA89DF4004FB41" box="[463,553,1222,1246]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14454" name="Universiti Kebangsaan Malaysia" pageId="1" pageNumber="146">UKMB</collectionCode>
),
<collectingCountry id="88E37679FFC4FFE5FD4A89DC4085FB41" box="[575,680,1221,1246]" name="Malaysia" pageId="1" pageNumber="146">Malaysia</collectingCountry>
,
<collectingRegion id="3230F80BFFC4FFE5FDCD89DC4120FB41" box="[696,781,1221,1246]" country="Malaysia" name="Pahang" pageId="1" pageNumber="146">Pahang</collectingRegion>
,
<location id="F52B6032FFC4FFE5FC6989DC41C1FB41" LSID="urn:lsid:plazi:treatment:785D87FFFFC4FFE7FFE28EF746A2FE67:F52B6032FFC4FFE5FC6989DC41C1FB41" box="[796,1004,1221,1246]" country="Malaysia" name="Genting Highland" pageId="1" pageNumber="146" stateProvince="Pahang">Genting Highland</location>
,
<date id="844A1029FFC4FFE5FC8E89DC464AFB41" box="[1019,1127,1221,1246]" pageId="1" pageNumber="146" value="2006-01-12">
<collectingDate id="940EE9C1FFC4FFE5FC8E89DC464AFB41" box="[1019,1127,1221,1246]" pageId="1" pageNumber="146" value="2006-01-12">12.i.2006</collectingDate>
</date>
,
<collectorName id="5D01533FFFC4FFE5FB0289DF4723FB41" box="[1143,1294,1221,1246]" pageId="1" pageNumber="146">N. Takahashi</collectorName>
leg
</materialsCitation>
.
<materialsCitation id="409C3CB4FFC4FFE5FA3889DF471DFA9D" ID-GBIF-Occurrence="1288049051" collectingDate="2005-03-15" collectionCode="UKMB" collectorName="N. Takahashi" country="Malaysia" location="Abu Suradi Trail-Mager Trail" municipality="Bukit Fraser" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<specimenCount id="E6F2FD60FFC4FFE5FA3889DF47B1FB41" box="[1357,1436,1221,1246]" pageId="1" pageNumber="146" type="male">1 male</specimenCount>
(
<collectionCode id="96E5AE2CFFC4FFE5FFEA89F342D4FA9D" box="[159,249,1258,1282]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14454" name="Universiti Kebangsaan Malaysia" pageId="1" pageNumber="146">UKMB</collectionCode>
),
<collectingCountry id="88E37679FFC4FFE5FE7989F04358FA9D" box="[268,373,1257,1282]" name="Malaysia" pageId="1" pageNumber="146">Malaysia</collectingCountry>
,
<collectingRegion id="3230F80BFFC4FFE5FEF589F043F8FA9D" box="[384,469,1257,1282]" country="Malaysia" name="Pahang" pageId="1" pageNumber="146">Pahang</collectingRegion>
,
<collectingMunicipality id="102FAC93FFC4FFE5FE9589F04043FA9E" box="[480,622,1257,1282]" pageId="1" pageNumber="146">Bukit Fraser</collectingMunicipality>
,
<location id="F52B6032FFC4FFE5FD0C89F041F4FA9D" LSID="urn:lsid:plazi:treatment:785D87FFFFC4FFE7FFE28EF746A2FE67:F52B6032FFC4FFE5FD0C89F041F4FA9D" box="[633,985,1257,1282]" country="Malaysia" municipality="Bukit Fraser" name="Abu Suradi Trail-Mager Trail" pageId="1" pageNumber="146" stateProvince="Pahang">Abu Suradi Trail—Mager Trail</location>
,
<date id="844A1029FFC4FFE5FC9089F0464CFA9D" box="[997,1121,1257,1282]" pageId="1" pageNumber="146" value="2005-03-15">
<collectingDate id="940EE9C1FFC4FFE5FC9089F0464CFA9D" box="[997,1121,1257,1282]" pageId="1" pageNumber="146" value="2005-03-15">15.iii.2005</collectingDate>
</date>
,
<collectorName id="5D01533FFFC4FFE5FB1989F3472DFA9D" box="[1132,1280,1257,1282]" pageId="1" pageNumber="146">N. Takahashi</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC4FFE5FFB288144374FAF1" pageId="1" pageNumber="146" type="diagnosis">
<paragraph id="F04B36E9FFC4FFE5FFB288144374FAF1" blockId="1.[151,1437,1113,2038]" pageId="1" pageNumber="146">
<emphasis id="C280EAFBFFC4FFE5FFB28814436CFAB9" bold="true" box="[199,321,1293,1318]" pageId="1" pageNumber="146">Diagnosis.</emphasis>
Body dark brown. Antennomere 3 slightly compressed and 1.75–2.30 times longer than wide. Pronotum without longitudinal carina on surface. Parameres of aedeagus widest at about basal 3/8 of length and hooked at apices.
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC4FFE7FFB28860407CFE83" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146" type="description">
<paragraph id="F04B36E9FFC4FFE5FFB2886040DAF9BD" blockId="1.[151,1437,1113,2038]" pageId="1" pageNumber="146">
<emphasis id="C280EAFBFFC4FFE5FFB288604374FA0D" bold="true" box="[199,345,1401,1426]" pageId="1" pageNumber="146">Description.</emphasis>
MALE. Body tiny and elongate (
<figureCitation id="68CF2A6CFFC4FFE5FDA188604138FA0D" box="[724,789,1401,1426]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Fig. 1</figureCitation>
), almost uniform dark brown and covered with brownish pubescence; cranium slightly dark reddish and sometimes faintly lighter around prominent frontal part (
<figureCitation id="68CF2A6CFFC4FFE5FA33888447A0FA29" box="[1350,1421,1437,1462]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Fig. 2</figureCitation>
); eyes black; pronotum and prosternum feebly dark reddish in some cases; antennomere 2 yellowish (
<figureCitation id="68CF2A6CFFC4FFE5FA5088D847A3FA45" box="[1317,1422,1473,1498]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Figs 2–3</figureCitation>
); pterothoracic ventrites and pleura sometimes faintly lighter; hindwings slightly dusky (
<figureCitation id="68CF2A6CFFC4FFE5FB0288FC4697FA61" box="[1143,1210,1509,1534]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Fig. 1</figureCitation>
); apical portions of tarsomere 5 with claws of legs more or less yellowish.
</paragraph>
<paragraph id="F04B36E9FFC4FFE5FFB28B344304F869" blockId="1.[151,1437,1113,2038]" pageId="1" pageNumber="146">
Head (
<figureCitation id="68CF2A6CFFC4FFE5FE638B344372F9D9" box="[278,351,1581,1606]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Fig. 2</figureCitation>
) small, hypognathous, quadrately rounded and widest at eyes in dorsal view; cranium with prominent frontal part, gently convex dorsad though depressed just behind prominent frontal part and slightly flattened behind vertex, lustrous without apparent microstructure and sparsely covered with long pubescence. Eyes with granular facets, large, semiglobularly prominent, shallowly and obtusely emarginate at posterior margin in profile, and diameter of an eye in profile 0.87–1.18 times as large as interocular distance. Antennae (
<figureCitation id="68CF2A6CFFC4FFE5FA588BA447BEF949" box="[1325,1427,1725,1750]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="1" pageNumber="146">Figs 1–3</figureCitation>
) inserted anterolaterally on prominent frontal part, moderately robust, densely covered with pubescence though less so on antennomeres 1–2, extending over elytral apices by antennomeres 10–11, scapus more or less robust, subcylindrical but weakly constricted on laterodistal surface at near the base, pedicel small and slightly broad, antennomeres 3–5 slightly compressed dorsoventrad, antennomere 3 subparallel at sides, but weakly constricted at base and 1.75–2.30 times longer than wide, subsequent antennomeres becoming gradually more subcylindrical and less compressed, antennomere 11 longest. Mouthparts with reduced mandibles, labrum minute, maxillary palpi 4- segmented with lanceolate terminal palpomeres prolonged at apices and more than 1.50 times longer than palpomere 3.
</paragraph>
<caption id="A48B6661FFC7FFE6FFE2892F40C3FBF3" httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="2" pageNumber="147" targetBox="[185,1436,200,1077]" targetPageId="2">
<paragraph id="F04B36E9FFC7FFE6FFE2892F40C3FBF3" blockId="2.[151,1436,1078,1132]" pageId="2" pageNumber="147">
<emphasis id="C280EAFBFFC7FFE6FFE2892F4310FBD3" bold="true" box="[151,317,1078,1101]" pageId="2" pageNumber="147">FIGURES 1–3.</emphasis>
<taxonomicName id="37F44D6AFFC7FFE6FE32892E404EFBD3" box="[327,611,1079,1100]" class="Insecta" family="Lycidae" genus="Alyculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="malaypeninsularis" status="sp. nov.">
<emphasis id="C280EAFBFFC7FFE6FE32892E404EFBD3" box="[327,611,1079,1100]" italics="true" pageId="2" pageNumber="147">Alyculus malaypeninsularis</emphasis>
</taxonomicName>
<emphasis id="C280EAFBFFC7FFE6FD18892E40ECFBD3" bold="true" box="[621,705,1079,1100]" pageId="2" pageNumber="147">
<taxonomicNameLabel id="D9B35780FFC7FFE6FD18892E40ECFBD3" box="[621,705,1079,1100]" pageId="2" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2–3).
</paragraph>
</caption>
<paragraph id="F04B36E9FFC7FFE6FFB2898D46F6FA53" blockId="2.[151,1436,1171,2024]" pageId="2" pageNumber="147">
Pronotum subtrapezoidal (
<figureCitation id="68CF2A6CFFC7FFE6FE9B898A407EFB33" box="[494,595,1171,1196]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="2" pageNumber="147">Figs 1–2</figureCitation>
), broader posteriad, strongly transverse, 1.7 5–1.88 times as wide as long at posterior margin; disc without longitudinal carina, sparsely provided with long erect pubescence, irregularly punctate particularly so on the areas along anterior and posterior margins and midline, finely granulate in basal third, gently convex medio-dorsad, transversely and feebly depressed at basal third, almost straight and feebly marginate at anterior margin, obtuse with blunt tips at front angles, somewhat emarginate mesad in dorsal view at lateral margins, which are weakly carinated in profile due to the continuity of pronotum and epipleura, acutely and apparently prominent laterad at hind angles, gently bisinuate and feebly marginate though sometimes faintly emarginate in the middle of posterior margin. Prothoracic epipleura irregularly punctate on surfaces. Scutellum bifurcate at apex, each distal process slender, protruded lateroposteriad and gently curved laterad.
</paragraph>
<paragraph id="F04B36E9FFC7FFE6FFB288CE4071F9C3" blockId="2.[151,1436,1171,2024]" pageId="2" pageNumber="147">
Elytra abbreviated (
<figureCitation id="68CF2A6CFFC7FFE6FED388CE43C5FA70" box="[422,488,1495,1520]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="2" pageNumber="147">Fig. 1</figureCitation>
), 1.7–2.0 times as long as wide at humeri, covering two basal abdominal segments, slightly dehiscent in posterior half of sutural line, each gradually narrowed posteriad and rounded at apex; their surface without costae or reticulate cells, rugose, densely covered with minute papillae bearing setae. Hind wings extending beyond elytral apices (
<figureCitation id="68CF2A6CFFC7FFE6FD7C8B5A4066F9C3" box="[521,587,1603,1628]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1078,1101]" captionTargetBox="[185,1436,200,1077]" captionTargetId="figure@2.[150,1436,181,1081]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Alyculus malaypeninsularis sp. nov. 1, habitus, dorsal view; 2, head and pronotum, dorsal view; 3, basal antennomeres. Scales: 0.5 mm (Fig. 1); 0.1 mm (Figs 2 – 3)." httpUri="https://zenodo.org/record/258384/files/figure.png" pageId="2" pageNumber="147">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F04B36E9FFC7FFE6FFB28B714634F957" blockId="2.[151,1436,1171,2024]" pageId="2" pageNumber="147">
Legs slender; femora and tibiae slightly flattened; tibia 1.15–1.30 times as long as femur and tarsomere V 1.33–1.76 times as long as tarsomere IV in fore leg; tarsomeres 2–4 usually faintly shorter than the preceding one, and tarsomere 5 approximately as long as tarsomere
<quantity id="370C9B0CFFC7FFE6FD918BB64122F957" box="[740,783,1711,1736]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="147" unit="in" value="1.0">1 in</quantity>
hind leg; claws simple.
</paragraph>
<paragraph id="F04B36E9FFC7FFE6FFB28BCD466BF8E3" blockId="2.[151,1436,1171,2024]" pageId="2" pageNumber="147">Meso- and metaventrites lustrous, and densely covered with pubescence on their surfaces compared to pubescence of cranium; discrimen traced in posterior half, but sometimes reaching anterior third to anterior margin of metaventrite. Abdomen elongate; venter lustrous though somewhat rugulose, densely covered with darker pubescence; penultimate sternite slightly narrower than the preceding one; ultimate tergite slightly elongate, semicylindrical, narrowed apicad; ultimate sternite leaf-shaped and narrowed basad.</paragraph>
<paragraph id="F04B36E9FFC7FFE6FFB28A9E43D9F877" blockId="2.[151,1436,1171,2024]" pageId="2" pageNumber="147">
Aedeagus (
<figureCitation id="68CF2A6CFFC7FFE6FE3F8A9E43BEF800" box="[330,403,1927,1952]" captionStart="FIGURE 4" captionStartId="3.[151,250,1933,1955]" captionTargetBox="[500,1071,566,1869]" captionTargetId="figure@3.[471,1086,559,1876]" captionTargetPageId="3" captionText="FIGURE 4. Aedeagus of Alyculus malaypeninsularis sp. nov., dorsal view. Scale: 0.1 mm." httpUri="https://zenodo.org/record/258385/files/figure.png" pageId="2" pageNumber="147">Fig. 4</figureCitation>
) trilobate; phallus slender, tapering to apex, slightly shorter than parameres; parameres widest at about basal 3/8 of length and tapering apicad in distal halves, flattened in profile and subparallel in distal third, weakly hooked at apices.
</paragraph>
<paragraph id="F04B36E9FFC6FFE7FFB28D814383FF2F" blockId="3.[151,1437,151,504]" box="[199,430,151,176]" pageId="3" pageNumber="148">FEMALE unknown.</paragraph>
<paragraph id="F04B36E9FFC6FFE7FFB28DA5407CFE83" blockId="3.[151,1437,151,504]" pageId="3" pageNumber="148">
<emphasis id="C280EAFBFFC6FFE7FFB28DA54350FF4A" bold="true" box="[199,381,188,213]" pageId="3" pageNumber="148">Measurements.</emphasis>
Body length
<quantity id="370C9B0CFFC6FFE7FD688DA540E9FF4B" box="[541,708,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.405" metricValueMax="2.7" metricValueMin="2.11" pageId="3" pageNumber="148" unit="mm" value="2.405" valueMax="2.7" valueMin="2.11">2.11–2.70 mm</quantity>
, pronotal length
<quantity id="370C9B0CFFC6FFE7FCFC8DA5461DFF4B" box="[905,1072,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="148" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20–0.30 mm</quantity>
, pronotal width
<quantity id="370C9B0CFFC6FFE7FB858DA547B5FF4B" box="[1264,1432,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.65" metricValueMax="5.5" metricValueMin="3.8" pageId="3" pageNumber="148" unit="mm" value="0.465" valueMax="0.55" valueMin="0.38">0.38–0.55 mm</quantity>
, length of elytra 0.75–1.0 mm, humeral width
<quantity id="370C9B0CFFC6FFE7FDCE8DC64148FF68" box="[699,869,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.3" metricValueMin="4.4" pageId="3" pageNumber="148" unit="mm" value="0.485" valueMax="0.53" valueMin="0.44">0.44–0.53 mm</quantity>
, frontal interocular distance
<quantity id="370C9B0CFFC6FFE7FBCE8DF94749FF68" box="[1211,1380,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="3" pageNumber="148" unit="mm" value="0.14" valueMax="0.17" valueMin="0.11">0.11–0.17 mm</quantity>
, eye diameter in lateral view
<quantity id="370C9B0CFFC6FFE7FED28C1D4060FE83" box="[423,589,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="3" pageNumber="148" unit="mm" value="0.145" valueMax="0.16" valueMin="0.13">0.13–0.16 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC6FFE7FFB28C3E40ADFEDF" box="[199,640,295,320]" pageId="3" pageNumber="148" type="distribution">
<paragraph id="F04B36E9FFC6FFE7FFB28C3E40ADFEDF" blockId="3.[151,1437,151,504]" box="[199,640,295,320]" pageId="3" pageNumber="148">
<emphasis id="C280EAFBFFC6FFE7FFB28C3E434DFEDF" bold="true" box="[199,352,295,320]" pageId="3" pageNumber="148">Distribution.</emphasis>
<collectingRegion id="3230F80BFFC6FFE7FE128C3143B3FEA0" box="[359,414,296,319]" country="Tanzania" name="Mjini Magharibi" pageId="3" pageNumber="148">West</collectingRegion>
<collectingCountry id="88E37679FFC6FFE7FED08C3E4026FEDF" box="[421,523,295,320]" name="Malaysia" pageId="3" pageNumber="148">Malaysia</collectingCountry>
(
<collectingRegion id="3230F80BFFC6FFE7FD6E8C3E4059FEDF" box="[539,628,295,320]" country="Malaysia" name="Pahang" pageId="3" pageNumber="148">Pahang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC6FFE7FFB28C554003FE4B" pageId="3" pageNumber="148" type="discussion">
<paragraph id="F04B36E9FFC6FFE7FFB28C554003FE4B" blockId="3.[151,1437,151,504]" pageId="3" pageNumber="148">
<emphasis id="C280EAFBFFC6FFE7FFB28C554316FEFA" bold="true" box="[199,315,332,357]" pageId="3" pageNumber="148">Remarks.</emphasis>
This new species has the largest in body size in the genus. It resembles
<taxonomicName id="37F44D6AFFC6FFE7FB0D8C5546D8FEFB" box="[1144,1269,332,356]" class="Insecta" family="Lycidae" genus="Alyculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="C280EAFBFFC6FFE7FB0D8C5546A4FEFC" box="[1144,1161,332,355]" italics="true" pageId="3" pageNumber="148">A</emphasis>
.
<emphasis id="C280EAFBFFC6FFE7FBED8C5446D8FEFB" box="[1176,1269,333,356]" italics="true" pageId="3" pageNumber="148">wittmeri</emphasis>
</taxonomicName>
from northern
<collectingRegion id="3230F80BFFC6FFE7FFE28C6942D9FE17" box="[151,244,368,392]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="148">Sumatra</collectingRegion>
in having the parameres of aedeagus with the hooked apices, but the body is much larger and dark brownish in coloration unlike the testaceous color of
<taxonomicName id="37F44D6AFFC6FFE7FD858C8D4141FE33" box="[752,876,404,428]" class="Insecta" family="Lycidae" genus="Alyculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="C280EAFBFFC6FFE7FD858C8D4141FE33" box="[752,876,404,428]" italics="true" pageId="3" pageNumber="148">A. wittmeri</emphasis>
</taxonomicName>
. The pronotum is narrower, and the parameres of aedeagus are widest at the basal 3/8.
</paragraph>
</subSubSection>
<subSubSection id="B8EE6562FFC6FFE7FFB28CC646A2FE67" box="[199,1167,479,504]" pageId="3" pageNumber="148" type="etymology">
<paragraph id="F04B36E9FFC6FFE7FFB28CC646A2FE67" blockId="3.[151,1437,151,504]" box="[199,1167,479,504]" pageId="3" pageNumber="148">
<emphasis id="C280EAFBFFC6FFE7FFB28CC64363FE67" bold="true" box="[199,334,479,504]" pageId="3" pageNumber="148">Etymology.</emphasis>
The specific epithet is derived from the known distribution of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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