208 lines
18 KiB
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208 lines
18 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.508.10005" ID-GBIF-Dataset="a9b3c019-c048-4d75-9310-4f15ee1e59e7" ID-PMC="PMC4492206" ID-Pensoft-Pub="1313-2970-508-69" ID-PubMed="26167121" ID-ZBK="03EB8C1E01F64E4A88A8D81EF7EAE503" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-508-69" ModsDocOrigin="ZooKeys 508" ModsDocTitle="The genus Scapheremaeus (Acari, Oribatida, Cymbaeremaeidae) in the oribatid mite fauna of New Zealand, with description of two new species" checkinTime="1451244249007" checkinUser="pensoft" docAuthor="Ermilov, Sergey G. & Minor, Maria A." docDate="2015" docId="21DBC5696387DAFD01D3B93A7228541C" docLanguage="en" docName="ZooKeys 508: 69-83" docOrigin="ZooKeys 508" docSource="http://dx.doi.org/10.3897/zookeys.508.10005" docTitle="Scapheremaeus luxtoni Ermilov & Minor, 2015, sp. n." docType="treatment" docUuid="8F13864C-3F7C-44A9-B831-2C4892BB7F89" docUuidSource="ZooBank" docVersion="4" lastPageNumber="75" masterDocId="FFC7FF924E776B09E77CB477FF82FF99" masterDocTitle="The genus Scapheremaeus (Acari, Oribatida, Cymbaeremaeidae) in the oribatid mite fauna of New Zealand, with description of two new species" masterLastPageNumber="83" masterPageNumber="69" pageNumber="71" updateTime="1668160414234" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Scapheremaeus (Acari, Oribatida, Cymbaeremaeidae) in the oribatid mite fauna of New Zealand, with description of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ermilov, Sergey G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Minor, Maria A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>508</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.508.10005</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.508.10005</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-508-69</mods:identifier>
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<mods:identifier type="ZBK">03EB8C1E01F64E4A88A8D81EF7EAE503</mods:identifier>
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<mods:identifier type="ZooBank">03EB8C1E01F64E4A88A8D81EF7EAE503</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152060388" LSID="urn:lsid:zoobank.org:act:8F13864C-3F7C-44A9-B831-2C4892BB7F89" httpUri="http://treatment.plazi.org/id/21DBC5696387DAFD01D3B93A7228541C" lastPageId="7" lastPageNumber="75" pageId="2" pageNumber="71">
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<subSubSection pageId="2" pageNumber="71" type="multiple">
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<paragraph pageId="2" pageNumber="71">Taxon classification Animalia Oribatida Cymbaeremaeidae</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="71" type="nomenclature">
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<paragraph pageId="2" pageNumber="71">
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<taxonomicName LSID="http://zoobank.org/8F13864C-3F7C-44A9-B831-2C4892BB7F89" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="luxtoni">Scapheremaeus luxtoni</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="71">sp. n.</taxonomicNameLabel>
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Figs 23-25, 26-35, 36-42, 43-47
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="72" type="diagnosis">
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<paragraph pageId="3" pageNumber="72">
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<pageBreakToken pageId="3" pageNumber="72" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="3" pageNumber="72">
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Body size: 381-415
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<normalizedToken originalValue="×">x</normalizedToken>
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199-232. Centro-dorsal part of notogaster areolate. Dorso-lateral parts of notogaster and ventral plate tuberculate. Costulae and transcostula strong. Rostral setae thin, straight. Lamellar setae minute. Bothridial setae globular. Humeral processes small, rounded. Circumdorsal scissure present. Ten pairs of short, simple notogastral setae. Palp femora with two setae. Six pairs of genital setae. Lyrifissures iad longitudinally oriented. Tridactylous.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="74" pageId="3" pageNumber="72" type="description">
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<paragraph pageId="3" pageNumber="72">Description.</paragraph>
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<paragraph pageId="3" pageNumber="72">Measurements. Body length: 381 (holotype: male), 381-415 (three paratypes: all females); notogaster width: 199 (holotype), 232 (same for three paratypes).</paragraph>
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<paragraph pageId="3" pageNumber="72">Integument. Body color light yellow-brownish. Anterior part of prodorsum and centro-dorsal part of notogaster with areolate sculpturing. Dorso-lateral parts of notogaster and ventral plate with elongated ridge-like tubercles.</paragraph>
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<paragraph pageId="4" pageNumber="73">
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<pageBreakToken pageId="4" pageNumber="73" start="start">Prodorsum</pageBreakToken>
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. Rostrum broadly rounded. Costulae (cos) distinct, forming slightly visible X-structure, terminated by large tubercles, which connected by thick transcostula (tcos). Rostral setae (6) thin, straight, inserted on transverse fold. Lamellar setae (4) minute, inserted nearer to rostral setae than to bothridia. Interlamellar and exobothridial setae and their alveoli absent. Bothridial setae (22-24) globular, pigmented, with short stalk (6) and longer (16-18) head, having longitudinal ridges.
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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<pageBreakToken pageId="5" pageNumber="74" start="start">Notogaster</pageBreakToken>
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flattened. Anterior margin straight. Lenticulus distinct. Humeral processes (Hp) slightly developed, tubercle-like in dorsal view and rounded in lateral view. Centrodorsal zone with longitudinal elongate hump-like structure. Circumdorsal scissure (f) present. Ten pairs of simple notogastral setae, located on small tubercles. Centro-dorsal part with two pairs of setae (lm, lm), both inserted near to scissure. All lyrifissures (except ia) well visible. Opisthonotal gland openings located medially to ih.
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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Gnathosoma. Subcapitulum longer than wide (82-90
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<normalizedToken originalValue="×">x</normalizedToken>
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61-69). Subcapitular setae thin, smooth; a and adoral setae (all 10) longer than m and h (both 6-8). Setae a slightly thicker than other. Palps (53-61) with setation 0
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<normalizedToken originalValue="–2–1–3–">-2-1-3-</normalizedToken>
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9(+ω). Solenidion free, not attached to eupathidium. Chelicerae (82-90) with two simple, barbed setae (both 16-20).
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<normalizedToken originalValue="Trägårdh’s">Traegardh's</normalizedToken>
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organ long, tapered.
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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Epimeral and lateral podosomal regions. Anterior tectum slightly developed. Pedotecta I of medium size, concave in dorsal view and scale-like in lateral view. Pedotecta II elongated, bifurcate distally in ventral view and broadly triangular in lateral view. Apodemes 1, 2, sejugal and 3 distinctly developed. Epimeral setal formula 3
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<normalizedToken originalValue="–1–2–">-1-2-</normalizedToken>
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2. Epimeral setae short (4), thin, smooth. Discidia roundly triangular.
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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Anogenital region. Six pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae similar in length (4), thin, smooth, inserted on small tubercles. Lyrifissures iad longitudinally oriented. Ovipositor elongated (52-56
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<normalizedToken originalValue="×">x</normalizedToken>
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41-45), lobes (32-36) longer than length of distal section (beyond middle fold; 20). Each of three lobes with four straight, smooth setae,
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<normalizedToken originalValue="ψ">ψ</normalizedToken>
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1 ≈
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<normalizedToken originalValue="τ">τ</normalizedToken>
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1 (24-28) longer than
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<normalizedToken originalValue="ψ">ψ</normalizedToken>
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2 ≈
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<normalizedToken originalValue="τa">τa</normalizedToken>
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≈
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<normalizedToken originalValue="τb">τb</normalizedToken>
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≈
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<normalizedToken originalValue="τc">τc</normalizedToken>
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(16). Coronal setae and their alveoli absent.
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</paragraph>
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<paragraph pageId="5" pageNumber="74">
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Legs. Tridactylous. Porose areas slightly visible, oval. Formulas of leg setation and solenidia as in
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus gibbus" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="gibbus">Scapheremaeus gibbus</taxonomicName>
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sp. n.; homology of setae and solenidia as indicated in Table 1. Famuli short, slightly dilated distally. Solenidia (except simple
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<normalizedToken originalValue="ω">ω</normalizedToken>
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on tarsi and
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<normalizedToken originalValue="φ">φ</normalizedToken>
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1, and thin
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<normalizedToken originalValue="σ">σ</normalizedToken>
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on genua I) dilated distally. Setae l on tibiae I setiform, not modified.
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</paragraph>
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<caption pageId="5" pageNumber="74">
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<paragraph pageId="5" pageNumber="74">
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Figures 23-25.
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="luxtoni">Scapheremaeus luxtoni</taxonomicName>
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sp. n., adult: 23 dorsal view 24 ventral view (legs not shown) 25 lateral view (gnathosoma and legs except basal parts not shown). Scale bar 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="74">
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<paragraph pageId="5" pageNumber="74">
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Figures 26-35.
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="luxtoni">Scapheremaeus luxtoni</taxonomicName>
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sp. n., adult: 26 frontal view of prodorsum 27 posterior view 28 subcapitulum 29 palp 30 chelicera, antiaxial view 31 ovipositor 32 leg I, right, antiaxial view 33 femur of leg II, left, paraxial view 34 trochanter, femur and genu of leg III, left, antiaxial view 35 leg IV, left, antiaxial view. Scale bars 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(26, 27), 20
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(28-35).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="74">
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<paragraph pageId="5" pageNumber="74">
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Figures 36-42.
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="luxtoni">Scapheremaeus luxtoni</taxonomicName>
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sp. n., dissected adult, microscope images: 36 bothridial seta and sculpture of latero-basal part of prodorsum 37 costulae and transcostula 38 lenticulus and sculpture on latero-anterior part of notogaster 39 humeral process, right, dorsal view 40 sculpture on centro-dorsal part of notogaster 41 notogastral seta p342 pedotecta I and II. Scale bar 20
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="74">
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<paragraph pageId="5" pageNumber="74">
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Figures 43-47.
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="luxtoni">Scapheremaeus luxtoni</taxonomicName>
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sp. n., dissected adult, microscope images: 43 sculpture on anterior part of epimeral region 44 anterior part of right anal plate 45 sculpture between genital and anal apertures 46 femur I, left, paraxial view 47 tarsus IV, left, antiaxial view. Scale bar 20
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="74" type="material examined">
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<paragraph pageId="5" pageNumber="74">Material examined.</paragraph>
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<paragraph pageId="5" pageNumber="74">
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Holotype (male) and three paratypes (all females): New Zealand, South Island, Central Otago, Pisa Range,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-44.871944">44°52'19"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="169.175">169°10'30"E</geoCoordinate>
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, 1880 m a.s.l., in soil and debris under
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<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Dracophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dracophyllum muscoides" order="Ericales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="muscoides">Dracophyllum muscoides</taxonomicName>
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cushion and in the soil outside of
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<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Dracophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dracophyllum muscoides" order="Ericales" pageId="5" pageNumber="74" phylum="Tracheophyta" rank="species" species="muscoides">Dracophyllum muscoides</taxonomicName>
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cushion, 18 February 2014, collected by M. Minor.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="74" type="type deposition">
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<paragraph pageId="5" pageNumber="74">Type deposition.</paragraph>
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<paragraph pageId="5" pageNumber="74">The holotype and one paratype are deposited in the New Zealand National Arthropod Collection, Auckland, New Zealand; one paratype is deposited in the collection of the Senckenberg Institution, Frankfurt, Germany; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="74" type="etymology">
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<paragraph pageId="5" pageNumber="74">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="74">The specific name is dedicated to the well-known acarologist Malcolm Luxton, for his extensive contributions to our knowledge of New Zealand oribatid mite fauna.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="76" pageId="5" pageNumber="74" type="remarks">
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<paragraph pageId="5" pageNumber="74">Remarks.</paragraph>
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<paragraph pageId="5" pageNumber="74">
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The new species is similar to
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus yamashitai" order="Sarcoptiformes" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yamashitai">Scapheremaeus yamashitai</taxonomicName>
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Aoki, 1970 from Japan (see
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<bibRefCitation author="Aoki, J" journalOrPublisher="Bul. Nat. Sci. Mus., Tokyo" pageId="8" pageNumber="77" pagination="585 - 602" title="Descriptions of oribatid mites collected by smoking of trees with insecticides." volume="13" year="1970">Aoki 1970</bibRefCitation>
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;
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<bibRefCitation author="Fujikawa, T" journalOrPublisher="Edaphologia" pageId="9" pageNumber="78" pagination="13 - 23" title="Three new species of Cepheidae and Cymbaeremaeidae (Acari: Oribatida) from Nippon." volume="69" year="2002">Fujikawa 2002</bibRefCitation>
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) in having circumdorsal furrow, tridactylous legs, costulae and transcostula, ten pairs of minute notogastral setae and areolate centrodorsal region of notogaster. However, it differs from the latter by the presence of thin notogastral setae (versus thickened), straight rostral setae (versus curved medially), short and dilated distally leg solenidia
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<normalizedToken originalValue="φ">φ</normalizedToken>
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2 (versus long and simple) and slightly developed humeral processes (versus well developed).
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</paragraph>
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<paragraph lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75">
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<pageBreakToken pageId="6" pageNumber="75" start="start">Also</pageBreakToken>
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, in having circumdorsal furrow, tridactylous legs, costulae, minute notogastral setae, straight rostral setae and areolate centrodorsal region of notogaster,
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus luxtoni" order="Sarcoptiformes" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="luxtoni">
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<pageBreakToken pageId="7" pageNumber="76" start="start">Scapheremaeus</pageBreakToken>
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luxtoni
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</taxonomicName>
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sp. n. is similar to
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<taxonomicName class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scapheremaeus zephyrus" order="Sarcoptiformes" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="zephyrus">Scapheremaeus zephyrus</taxonomicName>
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Colloff, 2010 from Australia (see
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<bibRefCitation author="Colloff, MJ" journalOrPublisher="Zootaxa" pageId="9" pageNumber="78" pagination="1 - 38" title="The hyperdiverse oribatid mite genus Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) in Australia, with descriptions of new species and consideration of biogeographical affinities." volume="2475" year="2010">Colloff 2010</bibRefCitation>
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) and New Zealand (our data). However, it differs from the latter by the presence of large tubercle-like distal parts of costulae (versus small), strong transcostula (versus absent), ten pairs of thin notogastral setae (versus nine pairs and thickened) and three pairs of adanal setae (versus two pairs).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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