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<document id="48F4C6065212A258538F8F9A318CDC52" ID-CLB-Dataset="56043" ID-DOI="10.5852/ejt.2022.813.1729" ID-GBIF-Dataset="ac406bb4-11b2-425e-b793-599560c503d9" ID-ISSN="2118-9773" ID-Zenodo-Dep="6448751" ID-ZooBank="687DE2C9-28A4-43E6-A47D-DE2D2839AB60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649696887367" checkinUser="felipe" docAuthor="Cole, Mary L. &amp; Herbert, David G." docDate="2022" docId="03A387ECFB607A58CDFE3523026EFD16" docLanguage="en" docName="ejt-813_1-32.pdf" docOrigin="European Journal of Taxonomy 813" docSource="http://dx.doi.org/10.5852/ejt.2022.813.1729" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella fordycei Cole &amp; Herbert 2022, sp. nov." docType="treatment" docUuid="8E77FF84-C946-433A-8070-A4CB4036053D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="10" masterDocId="FF9AFF94FB647A51CF7633470125FFEF" masterDocTitle="Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="5" updateTime="1699319808271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="BDFD03C5F7CBE30B075D6C9E941739E9">
<mods:title id="A5B88A3581FF5DABDC90645ED9B6EE52">Eight new species of Gulella Pfeiffer, 1856 from the south-east coast of South Africa (Gastropoda: Streptaxidae)</mods:title>
</mods:titleInfo>
<mods:name id="E3541A6906B40406E3AFC0FBD506856D" type="personal">
<mods:role id="3F359A535D0C3809F755AA461EE4DC2F">
<mods:roleTerm id="9E15483FB16DC28510218FB3F077346C">Author</mods:roleTerm>
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<mods:namePart id="9EF36C519A45C71D26D54FE91A613993">Cole, Mary L.</mods:namePart>
<mods:nameIdentifier id="96604DB5A725FAC495EE9EA96C8587A1" type="LSID">FDA80F4A-4E08-401E-8FF6-778E033BBBA7</mods:nameIdentifier>
<mods:affiliation id="94074639D9BECA0471AC535A2516CA2F">East London Museum, 319 Oxford St, East London, 5201, South Africa and Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa.</mods:affiliation>
<mods:nameIdentifier id="3557C80C18A0D76E879A97B261E2B890" type="email">marybursey@elmuseum.za.org</mods:nameIdentifier>
</mods:name>
<mods:name id="A6A30AD213BA1DE2F907A3293B060B5A" type="personal">
<mods:role id="5A38DE5F42FF0D8FAE04CD3E227E06D4">
<mods:roleTerm id="AB23D2AF9FB42DEB30D245BB604E5D22">Author</mods:roleTerm>
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<mods:namePart id="D7AA7DFF794529F0DBDEAF880723FA2A">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="7DC92B164EA9AF5FA6B62FA689ACC289" type="LSID">0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="0A1CA322E5AC34EA48833F6A2D1D6E21">Department of Natural Sciences, National Museum Wales, Cardiff, United Kingdom and University of KwaZulu-Natal, P. Bag. X 01, Scottsville, 3209, South Africa.</mods:affiliation>
<mods:nameIdentifier id="581417B8F1AE8E70D90AB626F1856DF6" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="18596AE27BB331B093350BB52CED3C8C">text</mods:typeOfResource>
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<mods:title id="D76692A57DDE7544E0766C79406D1593">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="49551088EACB57783D71BD01231784CB">
<mods:date id="BCC2337F64640DEDA651FD4CD926626A">2022</mods:date>
<mods:detail id="5445F5D9FA9B0AF189A0C603309CB937" type="pubDate">
<mods:number id="98A134EF4CB02687B7376630C56114E4">2022-04-11</mods:number>
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<mods:number id="DE68158D8E026F58CECB394EA3717034">813</mods:number>
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<mods:classification id="F8F9866B67202C76F515D7B308E9F1A0">journal article</mods:classification>
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<mods:identifier id="07EA5D34AAF1885F8A1A0124B9941D4C" type="DOI">10.5852/ejt.2022.813.1729</mods:identifier>
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<treatment id="03A387ECFB607A58CDFE3523026EFD16" ID-DOI="http://doi.org/10.5281/zenodo.6452993" ID-GBIF-Taxon="194587706" ID-Zenodo-Dep="6452993" LSID="urn:lsid:zoobank.org:act:8E77FF84-C946-433A-8070-A4CB4036053D" httpUri="http://treatment.plazi.org/id/03A387ECFB607A58CDFE3523026EFD16" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5">
<subSubSection id="C3106571FB607A55CDFE35230289F96F" box="[648,940,1636,1664]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BB536FAFB607A55CDFE35230289F96F" blockId="4.[372,1215,1636,1734]" box="[648,940,1636,1664]" pageId="4" pageNumber="5">
<heading id="D0FD8196FB607A55CDFE35230289F96F" box="[648,940,1636,1664]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C0A4D79FB607A55CDFE3523026EF990" ID-CoL="8ZHN3" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[648,843,1636,1663]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB607A55CDFE3523026EF990" bold="true" box="[648,843,1636,1663]" italics="true" pageId="4" pageNumber="5">Gulella fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB607A55CC2435210289F96F" box="[850,940,1638,1664]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB607A55CE0235CF024CF929" pageId="4" pageNumber="5" type="description">
<paragraph id="8BB536FAFB607A55CE0235CF059AF94C" blockId="4.[372,1215,1636,1734]" box="[372,1215,1672,1699]" pageId="4" pageNumber="5">
<uri id="FF9B3AF8FB607A55CE0235CF059AF94C" box="[372,1215,1672,1699]" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
<uuid id="FFAC0C2FFB607A55CDE435CF059AF94C" box="[658,1215,1672,1699]" pageId="4" pageNumber="5">8E77FF84-C946-433A-8070-A4CB4036053D</uuid>
</uri>
</paragraph>
<paragraph id="8BB536FAFB607A55CDBC35EC024CF929" blockId="4.[372,1215,1636,1734]" box="[714,873,1707,1734]" pageId="4" pageNumber="5">
<figureCitation id="13312A7FFB607A55CDBC35EC0230F929" box="[714,789,1707,1734]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[189,1398,1181,1899]" captionTargetId="figure-327@6.[189,1398,1181,1899]" captionTargetPageId="6" captionText="Fig. 2. Distribution map of Gulella kenbrowni sp. nov. (pink circles), G. fordycei sp. nov. (blue squares), G. mkombeni sp. nov. (green triangles) and G. crookesi sp. nov. (orange square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.6448759" httpUri="https://zenodo.org/record/6448759/files/figure.png" pageId="4" pageNumber="5">Figs 2</figureCitation>
,
<figureCitation id="13312A7FFB607A55CC5735EB024CF929" box="[801,873,1708,1734]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="4" pageNumber="5">3A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB607A55CFCB35AA0285F82C" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BB536FAFB607A55CFCB35AA0013F8E8" blockId="4.[189,310,1773,1799]" box="[189,310,1773,1799]" pageId="4" pageNumber="5">
<heading id="D0FD8196FB607A55CFCB35AA0013F8E8" bold="true" box="[189,310,1773,1799]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B97EEAE8FB607A55CFCB35AA0013F8E8" bold="true" box="[189,310,1773,1799]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB607A55CFCB345B0285F82C" blockId="4.[189,1399,1819,1987]" pageId="4" pageNumber="5">SHELL minute, cylindrical; smooth and glossy, but with weak axial riblets below suture; apertural dentition seven-fold, including a sinuous parietal lamella, a large labral complex extending deeply into aperture with ridge-like upper and lower margins and a groove in centre, a low, deeply inset transverse basal tooth to right of centre, a ridge-like basal tooth to left of centre beginning close to lip edge and a large inset tricuspid columella lamella; umbilicus widely open.</paragraph>
</subSubSection>
<caption id="DF756672FB617A54CFCB34C20309F809" ID-DOI="http://doi.org/10.5281/zenodo.6448755" ID-Zenodo-Dep="6448755" httpUri="https://zenodo.org/record/6448755/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1925,1951]" targetBox="[217,1376,286,1882]" targetPageId="5">
<paragraph id="8BB536FAFB617A54CFCB34C20309F809" blockId="5.[189,1399,1925,2022]" pageId="5" pageNumber="6">
<emphasis id="B97EEAE8FB617A54CFCB34C20029F870" bold="true" box="[189,268,1925,1951]" pageId="5" pageNumber="6">Fig. 1.</emphasis>
<taxonomicName id="4C0A4D79FB617A54CE6334C200D7F870" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[277,498,1925,1951]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB617A54CE6334C200D7F870" box="[277,498,1925,1951]" italics="true" pageId="5" pageNumber="6">Gulella kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB617A54CE8C34C2037CF870" box="[506,601,1925,1951]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMSA P1678/T4524), length 2.20 mm, width 0.95 mm.
<emphasis id="B97EEAE8FB617A54CFCB34EF01F7F82D" bold="true" box="[189,210,1960,1986]" pageId="5" pageNumber="6">A</emphasis>
. Aperture view.
<emphasis id="B97EEAE8FB617A54CED434EF0093F82D" bold="true" box="[418,438,1960,1986]" pageId="5" pageNumber="6">B</emphasis>
. Side view.
<emphasis id="B97EEAE8FB617A54CD2534EF034DF82D" bold="true" box="[595,616,1960,1986]" pageId="5" pageNumber="6">C</emphasis>
. Oblique view into aperture.
<emphasis id="B97EEAE8FB617A54CCA134EF02C9F82D" bold="true" box="[983,1004,1960,1986]" pageId="5" pageNumber="6">D</emphasis>
. Oblique view of base showing umbilicus. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3106571FB627A57CFCB324E03E5FEBE" box="[189,704,265,338]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BB536FAFB627A57CFCB324E0061FECC" blockId="6.[189,324,265,291]" box="[189,324,265,291]" pageId="6" pageNumber="7">
<heading id="D0FD8196FB627A57CFCB324E0061FECC" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B97EEAE8FB627A57CFCB324E0061FECC" bold="true" box="[189,324,265,291]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB627A57CFCB327003E5FEBE" blockId="6.[189,704,311,338]" box="[189,704,311,338]" pageId="6" pageNumber="7">
Named after the
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locality, Fort Fordyce.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB627A56CFCB323F0392FE5F" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BB536FAFB627A57CFCB323F008FFE7D" blockId="6.[189,426,376,402]" box="[189,426,376,402]" pageId="6" pageNumber="7">
<heading id="D0FD8196FB627A57CFCB323F008FFE7D" bold="true" box="[189,426,376,402]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B97EEAE8FB627A57CFCB323F008FFE7D" bold="true" box="[189,426,376,402]" pageId="6" pageNumber="7">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB627A57CF9C32E0004FFDC4" blockId="6.[189,1398,423,555]" pageId="6" pageNumber="7">
<materialsCitation id="3B623CA7FB627A57CF9C32E0004FFDC4" ID-GBIF-Occurrence="3750850333" collectingDate="2016-03-31" collectionCode="NMSA" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="P1679/T4525" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<emphasis id="B97EEAE8FB627A57CF9C32E00079FE2E" bold="true" box="[234,348,423,449]" pageId="6" pageNumber="7">
<typeStatus id="54B18858FB627A57CF9C32E00079FE2E" box="[234,348,423,449]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F31D766AFB627A57CFCB328D00B4FE0B" box="[189,401,458,484]" name="South Africa" pageId="6" pageNumber="7">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB627A57CEC5328D0356FE0B" bold="true" box="[435,627,458,484]" pageId="6" pageNumber="7">
<collectingRegion id="49CEF818FB627A57CEC5328D037AFE0B" box="[435,607,458,484]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="7">Eastern Cape</collectingRegion>
•
</emphasis>
<location id="8ED56021FB627A57CD0A328C02FFFE0A" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB627A57CD0A328C02FFFE0A" box="[636,986,458,485]" country="South Africa" latitude="-32.6881" longLatPrecision="6" longitude="26.513" name="Fort Fordyce Nature Reserve" pageId="6" pageNumber="7" stateProvince="Eastern Cape">Fort Fordyce Nature Reserve</location>
,
<location id="8ED56021FB627A57CC9E328D0029FDE8" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB627A57CC9E328D0029FDE8" country="South Africa" latitude="-32.6881" longLatPrecision="6" longitude="26.513" name="kloof with watercourse and sheer krantz" pageId="6" pageNumber="7" stateProvince="Eastern Cape">kloof with watercourse and sheer krantz</location>
;
<geoCoordinate id="EE3E503DFB627A57CE6C32AA00B2FDE7" box="[282,407,493,520]" degrees="32.6881" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="5" value="-32.6881">32.6881°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB627A57CED032AA0303FDE7" box="[422,550,493,520]" degrees="26.5130" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="5" value="26.513">26.5130°E</geoCoordinate>
;
<elevation id="0027D1C9FB627A57CD4332AA03E6FDE8" box="[565,707,493,520]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="6" pageNumber="7" unit="m" value="915.0">
<quantity id="4CF29B1FFB627A57CD4332AA03A4FDE7" box="[565,641,493,520]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="6" pageNumber="7" unit="m" value="915.0">915 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB627A57CDA432A90252FDE8" box="[722,887,493,520]" pageId="6" pageNumber="7" value="2016-03-31">
<collectingDate id="EFF0E9D2FB627A57CDA432A90252FDE8" box="[722,887,493,520]" pageId="6" pageNumber="7" value="2016-03-31">31 Mar. 2016</collectingDate>
</date>
;
<collectorName id="26FF532CFB627A57CCF332A902C3FDE8" box="[901,998,493,520]" pageId="6" pageNumber="7">M. Cole</collectorName>
leg.;
<collectionCode id="ED1BAE3FFB627A57CB5B32A905A5FDE7" box="[1069,1152,494,520]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB627A57CBFF32AA0409FDE7" box="[1161,1324,493,520]" pageId="6" pageNumber="7">P1679/T4525</specimenCode>
, prev. ELMD 18109.
</materialsCitation>
</paragraph>
<paragraph id="8BB536FAFB627A57CF9C31150245FBB6" blockId="6.[187,1399,594,1114]" pageId="6" pageNumber="7">
<materialsCitation id="3B623CA7FB627A57CF9C31150599FD5C" ID-GBIF-Occurrence="3750850337" collectingDate="2009-10-05" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="1072" latitude="-32.6834" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4767" municipality="Fort Fordyce Nature Reserve" pageId="6" pageNumber="7" specimenCode="ELMD 16088/T 218" specimenCount="4" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="B97EEAE8FB627A57CF9C31150042FD83" bold="true" box="[234,359,594,620]" pageId="6" pageNumber="7">
<typeStatus id="54B18858FB627A57CF9C31150042FD83" box="[234,359,594,620]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F31D766AFB627A57CFCB313200B2FD60" box="[189,407,629,655]" name="South Africa" pageId="6" pageNumber="7">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB627A57CEB03132035DFD60" bold="true" box="[454,632,629,655]" pageId="6" pageNumber="7">
<collectingRegion id="49CEF818FB627A57CEB03132035DFD60" box="[454,632,629,655]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="7">Eastern Cape</collectingRegion>
</emphasis>
•
<specimenCount id="9D0CFD73FB627A57CDD531320222FD7F" box="[675,775,629,656]" pageId="6" pageNumber="7">4 specs</specimenCount>
.;
<collectingMunicipality id="6BD1AC80FB627A57CC56313105B7FD7F" box="[800,1170,629,656]" pageId="6" pageNumber="7">Fort Fordyce Nature Reserve</collectingMunicipality>
,
<locationDeviation id="9A072FD8FB627A57CBD131310457FD7F" box="[1191,1394,629,656]" pageId="6" pageNumber="7">near Harris Hut</locationDeviation>
;
<geoCoordinate id="EE3E503DFB627A57CFCB31DF001AFD5C" box="[189,319,664,691]" degrees="32.6834" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="5" value="-32.6834">32.6834° S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB627A57CE3B31DF00EAFD5C" box="[333,463,664,691]" degrees="26.4767" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="5" value="26.4767">26.4767°E</geoCoordinate>
;
<elevation id="0027D1C9FB627A57CEAB31DE035DFD5D" box="[477,632,664,691]" metricMagnitude="3" metricUnit="m" metricValue="1.072" pageId="6" pageNumber="7" unit="m" value="1072.0">
<quantity id="4CF29B1FFB627A57CEAB31DE0312FD5C" box="[477,567,665,691]" metricMagnitude="3" metricUnit="m" metricValue="1.072" pageId="6" pageNumber="7" unit="m" value="1072.0">1072 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB627A57CDF131DF0233FD5C" box="[647,790,664,691]" pageId="6" pageNumber="7" value="2009-10-05">
<collectingDate id="EFF0E9D2FB627A57CDF131DF0233FD5C" box="[647,790,664,691]" pageId="6" pageNumber="7" value="2009-10-05">5 Oct. 2009</collectingDate>
</date>
;
<collectorName id="26FF532CFB627A57CC5531DE02A1FD5D" box="[803,900,664,691]" pageId="6" pageNumber="7">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB627A57CCBF31DE0599FD5C" box="[969,1212,664,691]" pageId="6" pageNumber="7">ELMD 16088/T 218</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CBA131DE03AAFD16" ID-GBIF-Occurrence="3750850309" collectingDate="2010-01-03" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="1120" latitude="-32.6956" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4857" pageId="6" pageNumber="7" specimenCode="ELMD 16317/T 219" specimenCount="2" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CBA131DE0411FD5C" box="[1239,1332,665,691]" pageId="6" pageNumber="7">2 specs</specimenCount>
.;
<location id="8ED56021FB627A57CA3031DE00FFFD39" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB627A57CA3031DE00FFFD39" country="South Africa" latitude="-32.6956" longLatPrecision="6" longitude="26.4857" name="Fort Fordyce Nature Reserve" pageId="6" pageNumber="7" stateProvince="Eastern Cape">Fort Fordyce Nature Reserve</location>
,
<locationDeviation id="9A072FD8FB627A57CE9231FB0254FD39" box="[484,881,700,726]" pageId="6" pageNumber="7">south-facing slope behind lookout</locationDeviation>
;
<geoCoordinate id="EE3E503DFB627A57CC0D31FC02DEFD39" box="[891,1019,699,726]" degrees="32.6956" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="5" value="-32.6956">32.6956°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB627A57CB7331FC05ACFD39" box="[1029,1161,699,726]" degrees="26.4857" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="5" value="26.4857">26.4857°E</geoCoordinate>
;
<elevation id="0027D1C9FB627A57CBE531FB0402FD39" box="[1171,1319,700,726]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="6" pageNumber="7" unit="m" value="1120.0">
<quantity id="4CF29B1FFB627A57CBE531FB05CCFD39" box="[1171,1257,700,726]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="6" pageNumber="7" unit="m" value="1120.0">1120 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB627A57CA4431FB01DCFD16" pageId="6" pageNumber="7" value="2010-01-03">
<collectingDate id="EFF0E9D2FB627A57CA4431FB01DCFD16" pageId="6" pageNumber="7" value="2010-01-03">3 Jan. 2010</collectingDate>
</date>
;
<collectorName id="26FF532CFB627A57CE7231980047FD16" box="[260,354,735,761]" pageId="6" pageNumber="7">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB627A57CED7319803AAFD16" box="[417,655,735,761]" pageId="6" pageNumber="7">ELMD 16317/T 219</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CDD3319800D8FCF3" ID-GBIF-Occurrence="3750850408" collectingDate="2010-01-03" collectionCode="NMSA" collectorName="M. Cole" country="South Africa" elevation="1120" latitude="-32.6956" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4857" pageId="6" pageNumber="7" specimenCode="P1616/ T4498" specimenCount="2" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CDD3319803DAFD16" box="[677,767,735,761]" pageId="6" pageNumber="7">2 specs</specimenCount>
.; same collection data as for preceding;
<collectionCode id="ED1BAE3FFB627A57CBBE3198043EFD16" box="[1224,1307,735,761]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB627A57CA5731980029FCF3" pageId="6" pageNumber="7">P1616/ T4498</specimenCode>
, prev. ELMD 16317
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CD6130450086FCD0" ID-GBIF-Occurrence="3750850346" collectingDate="2010-01-03" collectionCode="NMW" collectorName="M. Cole" country="South Africa" elevation="1120" latitude="-32.6956" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4857" pageId="6" pageNumber="7" specimenCode="NMW.Z.2021.011.00002" specimenCount="2" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CD6130450356FCF2" box="[535,627,770,797]" pageId="6" pageNumber="7">2 specs</specimenCount>
.; same collection data as for preceding;
<specimenCode id="DBAC9E81FB627A57CB3F30450457FCF3" box="[1097,1394,770,796]" pageId="6" pageNumber="7">NMW.Z.2021.011.00002</specimenCode>
, prev. ELMD 16317
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CEC830620591FC8C" ID-GBIF-Occurrence="3750850418" collectingDate="2008-12-29" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="ELMD 16377/T 220" specimenCount="6" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CEC83062033EFCAF" box="[446,539,805,832]" pageId="6" pageNumber="7">6 specs</specimenCount>
.;
<location id="8ED56021FB627A57CD5B306102A2FCAF" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB627A57CD5B306102A2FCAF" box="[557,903,805,832]" country="South Africa" latitude="-32.6881" longLatPrecision="6" longitude="26.513" name="Fort Fordyce Nature Reserve" pageId="6" pageNumber="7" stateProvince="Eastern Cape">Fort Fordyce Nature Reserve</location>
,
<locationDeviation id="9A072FD8FB627A57CCE230620457FCD0" box="[916,1394,805,832]" pageId="6" pageNumber="7">kloof with watercourse and sheer krantz</locationDeviation>
;
<geoCoordinate id="EE3E503DFB627A57CFC8300F0019FC8C" box="[190,316,840,867]" degrees="32.6881" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="5" value="-32.6881">32.6881°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB627A57CE3F300F00E8FC8C" box="[329,461,840,867]" degrees="26.5130" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="5" value="26.513">26.5130°E</geoCoordinate>
;
<elevation id="0027D1C9FB627A57CEAC300F0341FC8D" box="[474,612,840,867]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="6" pageNumber="7" unit="m" value="915.0">
<quantity id="4CF29B1FFB627A57CEAC300F0301FC8C" box="[474,548,840,867]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="6" pageNumber="7" unit="m" value="915.0">915 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB627A57CD04300E0236FC8C" box="[626,787,841,867]" pageId="6" pageNumber="7" value="2008-12-29">
<collectingDate id="EFF0E9D2FB627A57CD04300E0236FC8C" box="[626,787,841,867]" pageId="6" pageNumber="7" value="2008-12-29">29 Dec. 2008</collectingDate>
</date>
;
<collectorName id="26FF532CFB627A57CC56300E02A5FC8D" box="[800,896,840,867]" pageId="6" pageNumber="7">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB627A57CCB5300E0591FC8C" box="[963,1204,840,867]" pageId="6" pageNumber="7">ELMD 16377/T 220</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CBBB300E0513FC6A" ID-GBIF-Occurrence="3750850321" collectingDate="2008-12-29" collectionCode="RMNH" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="RMNH.MOL.452585" specimenCount="2" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CBBB300E040FFC8C" box="[1229,1322,841,867]" pageId="6" pageNumber="7">2 specs</specimenCount>
.; same collection data as for preceding;
<specimenCode id="DBAC9E81FB627A57CD37302C0266FC69" box="[577,835,875,902]" pageId="6" pageNumber="7">RMNH.MOL.452585</specimenCode>
, prev. ELMD 16377
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CB27302B0262FC46" ID-GBIF-Occurrence="3750850311" collectingDate="2009-10-06" collectionCode="RMNH" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="ELMD 16073/T 221" specimenCount="3" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CB27302B058BFC69" box="[1105,1198,876,902]" pageId="6" pageNumber="7">3 specs</specimenCount>
.; same collection data as for preceding;
<date id="FFB4103AFB627A57CEB630C80369FC46" box="[448,588,911,937]" pageId="6" pageNumber="7" value="2009-10-06">
<collectingDate id="EFF0E9D2FB627A57CEB630C80369FC46" box="[448,588,911,937]" pageId="6" pageNumber="7" value="2009-10-06">6 Oct. 2009</collectingDate>
</date>
;
<specimenCode id="DBAC9E81FB627A57CD2130C80262FC46" box="[599,839,911,937]" pageId="6" pageNumber="7">ELMD 16073/T 221</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CC2930C80389FC23" ID-GBIF-Occurrence="3750850349" collectingDate="2009-10-06" collectionCode="NHMUK" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="NHMUK 20210067" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CC2930C80296FC46" box="[863,947,911,937]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<specimenCode id="DBAC9E81FB627A57CFCA30F4008AFC23" box="[188,431,946,973]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="7" type="Museum">NHMUK 20210067</specimenCode>
, prev. ELMD 16073
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CDBB30F500D0FC00" ID-GBIF-Occurrence="3750850397" collectingDate="2015-01-17" collectionCode="NHMUK" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="ELMD 17842/T 222" specimenCount="3" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CDBB30F50208FC22" box="[717,813,946,973]" pageId="6" pageNumber="7">3 specs</specimenCount>
.; same collection data as for preceding;
<date id="FFB4103AFB627A57CA6830F501DEFC1F" pageId="6" pageNumber="7" value="2015-01-17">
<collectingDate id="EFF0E9D2FB627A57CA6830F501DEFC1F" pageId="6" pageNumber="7" value="2015-01-17">17 Jan. 2015</collectingDate>
</date>
;
<specimenCode id="DBAC9E81FB627A57CE70309100D0FC00" box="[262,501,981,1008]" pageId="6" pageNumber="7">ELMD 17842/T 222</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CD7B3092007AFBFC" ID-GBIF-Occurrence="3750850307" collectingDate="2015-01-17" collectionCode="NMSA" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="P1615/T4497" specimenCount="3" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CD7B3092034DFC1F" box="[525,616,981,1008]" pageId="6" pageNumber="7">3 specs</specimenCount>
.; same collection data as for preceding;
<collectionCode id="ED1BAE3FFB627A57CB40309205ACFC00" box="[1078,1161,981,1007]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
<specimenCode id="DBAC9E81FB627A57CBF930920417FC1F" box="[1167,1330,981,1008]" pageId="6" pageNumber="7">P1615/T4497</specimenCode>
, prev. ELMD 17842
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CE0E30BE0450FBFC" ID-GBIF-Occurrence="3750850366" box="[376,1397,1016,1043]" collectingDate="2015-01-17" collectionCode="NHMUK" collectorName="M. Cole" country="South Africa" elevation="915" latitude="-32.6881" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.513" pageId="6" pageNumber="7" specimenCode="NHMUK 20210068" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CE0E30BE00E9FBFC" box="[376,460,1017,1043]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
; same collection data as for preceding;
<specimenCode id="DBAC9E81FB627A57CCEE30BE05A3FBFD" box="[920,1158,1016,1043]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="7" type="Museum">NHMUK 20210068</specimenCode>
, prev. ELMD 17842
</materialsCitation>
•
<materialsCitation id="3B623CA7FB627A57CFA6375B0278FBB6" ID-GBIF-Occurrence="3750850320" collectingDate="2015-01-18" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="1117" latitude="-32.6825" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4794" municipality="Fort Fordyce Nature Reserve" pageId="6" pageNumber="7" specimenCode="ELMD 17864/T 223" specimenCount="2" stateProvince="Eastern Cape" typeStatus="paratype">
<specimenCount id="9D0CFD73FB627A57CFA6375B000AFBD9" box="[208,303,1052,1078]" pageId="6" pageNumber="7">2 specs</specimenCount>
.;
<collectingMunicipality id="6BD1AC80FB627A57CE37375B0385FBD9" box="[321,672,1052,1078]" pageId="6" pageNumber="7">Fort Fordyce Nature Reserve</collectingMunicipality>
,
<locationDeviation id="9A072FD8FB627A57CDD9375B0529FBD9" box="[687,1036,1052,1078]" pageId="6" pageNumber="7">between Harris Hut and dam</locationDeviation>
;
<geoCoordinate id="EE3E503DFB627A57CB6D375C05BFFBD9" box="[1051,1178,1051,1078]" degrees="32.6825" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="5" value="-32.6825">32.6825°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB627A57CBDC375C040AFBD9" box="[1194,1327,1051,1078]" degrees="26.4794" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="5" value="26.4794">26.4794° E</geoCoordinate>
;
<elevation id="0027D1C9FB627A57CA48375B0031FBB6" metricMagnitude="3" metricUnit="m" metricValue="1.117" pageId="6" pageNumber="7" unit="m" value="1117.0">
<quantity id="4CF29B1FFB627A57CA48375B01F1FBB6" metricMagnitude="3" metricUnit="m" metricValue="1.117" pageId="6" pageNumber="7" unit="m" value="1117.0">1117 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB627A57CE573778009FFBB5" box="[289,442,1087,1114]" pageId="6" pageNumber="7" value="2015-01-18">
<collectingDate id="EFF0E9D2FB627A57CE573778009FFBB5" box="[289,442,1087,1114]" pageId="6" pageNumber="7" value="2015-01-18">18 Jan. 2015</collectingDate>
</date>
;
<collectorName id="26FF532CFB627A57CEB037780303FBB6" box="[454,550,1087,1113]" pageId="6" pageNumber="7">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB627A57CD1E37780278FBB6" box="[616,861,1087,1113]" pageId="6" pageNumber="7">ELMD 17864/T 223</specimenCode>
</materialsCitation>
.
</paragraph>
<caption id="DF756672FB627A57CFCB34EC0470F806" ID-DOI="http://doi.org/10.5281/zenodo.6448759" ID-Zenodo-Dep="6448759" httpUri="https://zenodo.org/record/6448759/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1963,1989]" targetBox="[189,1398,1181,1899]" targetPageId="6">
<paragraph id="8BB536FAFB627A57CFCB34EC0470F806" blockId="6.[188,1398,1963,2025]" pageId="6" pageNumber="7">
<emphasis id="B97EEAE8FB627A57CFCB34EC002FF82A" bold="true" box="[189,266,1963,1989]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
Distribution map of
<taxonomicName id="4C0A4D79FB627A57CE8D34EC03F1F82A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[507,724,1963,1989]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB627A57CE8D34EC03F1F82A" box="[507,724,1963,1989]" italics="true" pageId="6" pageNumber="7">Gulella kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB627A57CDAD34EB0210F829" box="[731,821,1964,1990]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(pink circles),
<taxonomicName id="4C0A4D79FB627A57CC9434EB0541F82A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[994,1124,1963,1990]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB627A57CC9434EB0541F82A" box="[994,1124,1963,1990]" italics="true" pageId="6" pageNumber="7">G. fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB627A57CB1D34EB05E0F829" box="[1131,1221,1964,1990]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(blue squares),
<taxonomicName id="4C0A4D79FB627A57CFCA3488007CF807" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[188,345,1998,2025]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="mkombeni" status="sp. nov.">
<emphasis id="B97EEAE8FB627A57CFCA3488007CF807" box="[188,345,1998,2025]" italics="true" pageId="6" pageNumber="7">G. mkombeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB627A57CE173488009EF806" box="[353,443,1999,2025]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(green triangles) and
<taxonomicName id="4C0A4D79FB627A57CDCA34880267F807" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[700,834,1998,2025]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="crookesi" status="sp. nov.">
<emphasis id="B97EEAE8FB627A57CDCA34880267F807" box="[700,834,1998,2025]" italics="true" pageId="6" pageNumber="7">G. crookesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB627A57CC3C34880281F806" box="[842,932,1999,2025]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(orange square). Contour at 1000 m.
</paragraph>
</caption>
<paragraph id="8BB536FAFB637A56CF9C324F0083FECD" blockId="7.[188,1399,264,432]" box="[234,422,264,290]" pageId="7" pageNumber="8">
<emphasis id="B97EEAE8FB637A56CF9C324F0083FECD" bold="true" box="[234,422,264,290]" pageId="7" pageNumber="8">Other material</emphasis>
</paragraph>
<paragraph id="8BB536FAFB637A56CFCB326B0392FE5F" blockId="7.[188,1399,264,432]" pageId="7" pageNumber="8">
<materialsCitation id="3B623CA7FB637A56CFCB326B05DFFE86" ID-GBIF-Occurrence="3750850409" collectingDate="2008-12-29" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="1137" latitude="-32.6704" location="Fort Fordyce Nature Reserve" longLatPrecision="6" longitude="26.4851" pageId="7" pageNumber="8" specimenCode="ELMD 16083" specimenCount="2" stateProvince="Eastern Cape">
<collectingCountry id="F31D766AFB637A56CFCB326B00B7FEA9" box="[189,402,300,326]" name="South Africa" pageId="7" pageNumber="8">SOUTH AFRICA</collectingCountry>
–
<emphasis id="B97EEAE8FB637A56CEDB326B034AFEA9" bold="true" box="[429,623,299,326]" pageId="7" pageNumber="8">
<collectingRegion id="49CEF818FB637A56CEDB326B037EFEAA" box="[429,603,299,326]" country="South Africa" name="Eastern Cape" pageId="7" pageNumber="8">Eastern Cape</collectingRegion>
•
</emphasis>
<specimenCount id="9D0CFD73FB637A56CD0E326B03F2FEA9" box="[632,727,300,326]" pageId="7" pageNumber="8">2 specs</specimenCount>
.;
<location id="8ED56021FB637A56CD9D326B056EFEA9" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB637A56CD9D326B056EFEA9" box="[747,1099,300,326]" country="South Africa" latitude="-32.6704" longLatPrecision="6" longitude="26.4851" name="Fort Fordyce Nature Reserve" pageId="7" pageNumber="8" stateProvince="Eastern Cape">Fort Fordyce Nature Reserve</location>
,
<locationDeviation id="9A072FD8FB637A56CB2D326B0457FEA9" box="[1115,1394,300,326]" pageId="7" pageNumber="8">kloof with watercourse</locationDeviation>
,
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;
<geoCoordinate id="EE3E503DFB637A56CE3B320800E8FE86" box="[333,461,335,361]" degrees="32.6704" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="-32.6704">32.6704°S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB637A56CEAC32080379FE86" box="[474,604,335,362]" degrees="26.4851" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="26.4851">26.4851°E</geoCoordinate>
;
<elevation id="0027D1C9FB637A56CD1E320803DBFE86" box="[616,766,335,361]" metricMagnitude="3" metricUnit="m" metricValue="1.137" pageId="7" pageNumber="8" unit="m" value="1137.0">
<quantity id="4CF29B1FFB637A56CD1E3208039AFE86" box="[616,703,335,361]" metricMagnitude="3" metricUnit="m" metricValue="1.137" pageId="7" pageNumber="8" unit="m" value="1137.0">1137 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB637A56CC7D3208028EFE86" box="[779,939,335,361]" pageId="7" pageNumber="8" value="2008-12-29">
<collectingDate id="EFF0E9D2FB637A56CC7D3208028EFE86" box="[779,939,335,361]" pageId="7" pageNumber="8" value="2008-12-29">29 Dec. 2008</collectingDate>
</date>
;
<collectorName id="26FF532CFB637A56CCC032080533FE86" box="[950,1046,335,361]" pageId="7" pageNumber="8">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB637A56CB21320805DFFE86" box="[1111,1274,335,361]" pageId="7" pageNumber="8">ELMD 16083</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CA7FB637A56CA6432080396FE5F" ID-GBIF-Occurrence="3750850364" collectingDate="2008-12-30" collectionCode="ELMD" collectorName="M. Cole" country="South Africa" elevation="1120" latitude="-32.6956" location="Fort Ford yce Nature Reserve" longLatPrecision="6" longitude="26.4857" municipality="Fort Ford" pageId="7" pageNumber="8" specimenCode="ELMD 16085" specimenCount="2" stateProvince="Eastern Cape">
<specimenCount id="9D0CFD73FB637A56CA6432080448FE86" box="[1298,1389,335,361]" pageId="7" pageNumber="8">2 specs</specimenCount>
.;
<location id="8ED56021FB637A56CFCA32340332FE62" LSID="urn:lsid:plazi:treatment:03A387ECFB607A58CDFE3523026EFD16:8ED56021FB637A56CFCA32340332FE62" box="[188,535,370,397]" country="South Africa" latitude="-32.6956" longLatPrecision="6" longitude="26.4857" municipality="Fort Ford" name="Fort Ford yce Nature Reserve" pageId="7" pageNumber="8" stateProvince="Eastern Cape">Fort Ford yce Nature Reserve</location>
,
<locationDeviation id="9A072FD8FB637A56CD5432350296FE63" box="[546,947,370,396]" pageId="7" pageNumber="8">south-facing slope behind lookout</locationDeviation>
;
<geoCoordinate id="EE3E503DFB637A56CCC932350565FE63" box="[959,1088,370,397]" degrees="32.6956" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="-32.6956">32.6956° S</geoCoordinate>
,
<geoCoordinate id="EE3E503DFB637A56CB3A323505EBFE62" box="[1100,1230,370,397]" degrees="26.4857" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="26.4857">26.4857°E</geoCoordinate>
;
<elevation id="0027D1C9FB637A56CBAC32350455FE63" box="[1242,1392,370,397]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="7" pageNumber="8" unit="m" value="1120.0">
<quantity id="4CF29B1FFB637A56CBAC32350414FE62" box="[1242,1329,370,397]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="7" pageNumber="8" unit="m" value="1120.0">1120 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB4103AFB637A56CFCA32D20078FE40" box="[188,349,405,432]" pageId="7" pageNumber="8" value="2008-12-30">
<collectingDate id="EFF0E9D2FB637A56CFCA32D20078FE40" box="[188,349,405,432]" pageId="7" pageNumber="8" value="2008-12-30">30 Dec. 2008</collectingDate>
</date>
;
<collectorName id="26FF532CFB637A56CE1F32D100ECFE40" box="[361,457,405,432]" pageId="7" pageNumber="8">M. Cole</collectorName>
leg.;
<specimenCode id="DBAC9E81FB637A56CD7D32D10396FE5F" box="[523,691,405,432]" pageId="7" pageNumber="8">ELMD 16085</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB637A56CFCB32910440FBBD" pageId="7" pageNumber="8" type="description">
<paragraph id="8BB536FAFB637A56CFCB3291006BFE1F" blockId="7.[189,334,470,496]" box="[189,334,470,496]" pageId="7" pageNumber="8">
<heading id="D0FD8196FB637A56CFCB3291006BFE1F" bold="true" box="[189,334,470,496]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B97EEAE8FB637A56CFCB3291006BFE1F" bold="true" box="[189,334,470,496]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB637A56CFCB31420440FBBD" blockId="7.[187,1400,517,1107]" pageId="7" pageNumber="8">
SHELL (
<figureCitation id="13312A7FFB637A56CE6231420079FDF0" box="[276,348,517,543]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
). Shell minute, cylindrical, length
<quantity id="4CF29B1FFB637A56CD8B314202ACFDCF" box="[765,905,517,544]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="7" pageNumber="8" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5mm</quantity>
, width 1.0–
<quantity id="4CF29B1FFB637A56CB6E3142054AFDCF" box="[1048,1135,517,544]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="8" unit="mm" value="1.1">1.1 mm</quantity>
, L:W 2.1–2.3 (n = 6). Protoconch approx.
<quantity id="4CF29B1FFB637A56CEC4316E00D5FDAC" box="[434,496,553,579]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="1.0">1mm</quantity>
in diameter, comprising approx. 2.25 whorls, smooth and glossy; junction between protoconch and teleoconch evident. Teleoconch comprising approx. 3.5–4 whorls; first whorl convex, remaining ones weakly so; weak axial riblets extend more or less half-way down each whorl, stronger immediately below sutures (
<figureCitation id="13312A7FFB637A56CDF931D503D6FD43" box="[655,755,658,684]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
). Aperture sub-quadrate, rounded basally, markedly constricted by teeth; peristome thick and reflected, broadly interrupted in parietal region; dentition eightfold (
<figureCitation id="13312A7FFB637A56CF8D319F0073FD1C" box="[251,342,728,755]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
): 1) a strong parietal lamella with inner portion which runs into aperture somewhat sinuous, outer portion curving to right; parietal lamella projects well beyond profile of aperture in side view; 2) a large labral slab extending into aperture beyond columella lamella (in apertural view), with ridges on its upper and lower margins, bordering a V-shaped central groove, upper ridge sharp with a cusp near lip edge in close proximity to parietal lamella; 3) a low, deeply inset transverse basal tooth to right of centre; 4) an in-running basal ridge to left of centre beginning near lip edge; 5–7) a large columella lamella with three teeth, upper two ridge-like and lowest one small, sometimes little more than a low bulge, middle one largest and extending closer to lip edge. Labral slab corresponds with a deep pit behind outer lip (
<figureCitation id="13312A7FFB637A56CFBE30B50007FBE3" box="[200,290,1010,1036]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
); basal ridge with a shallow indentation behind lip. Umbilicus widely open, elongate-oval, with a deep indentation underlying columella lamella, indistinct axial pleats around umbilicus (
<figureCitation id="13312A7FFB637A56CA703752044DFBC0" box="[1286,1384,1045,1071]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[239,1349,282,1880]" captionTargetId="figure-18@8.[235,1352,261,1885]" captionTargetPageId="8" captionText="Fig. 3. Gulella fordycei sp. nov., holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
). Shell almost transparent when fresh, orange-red coloration of dried tissue of animal visible internally.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB637A56CFCB373D03E5FB09" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BB536FAFB637A56CFCB373D0093FB7B" blockId="7.[189,438,1146,1172]" box="[189,438,1146,1172]" pageId="7" pageNumber="8">
<emphasis id="B97EEAE8FB637A56CFCB373D0073FB7B" bold="true" box="[189,342,1146,1172]" pageId="7" pageNumber="8">Distribution</emphasis>
(
<figureCitation id="13312A7FFB637A56CE10373D0088FB7B" box="[358,429,1146,1172]" captionStart="Fig" captionStartId="6.[189,232,1963,1989]" captionTargetBox="[189,1398,1181,1899]" captionTargetId="figure-327@6.[189,1398,1181,1899]" captionTargetPageId="6" captionText="Fig. 2. Distribution map of Gulella kenbrowni sp. nov. (pink circles), G. fordycei sp. nov. (blue squares), G. mkombeni sp. nov. (green triangles) and G. crookesi sp. nov. (orange square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.6448759" httpUri="https://zenodo.org/record/6448759/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
)
</paragraph>
<paragraph id="8BB536FAFB637A56CFCB37EF03E5FB09" blockId="7.[189,1398,1192,1254]" pageId="7" pageNumber="8">
Endemic to the Fort Fordyce Nature Reserve, considered part of the Amathole Mountains; at altitudes between
<quantity id="4CF29B1FFB637A56CE51378B0048FB09" box="[295,365,1228,1254]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="m" value="900.0">900m</quantity>
and
<quantity id="4CF29B1FFB637A56CED0378C00D8FB09" box="[422,509,1227,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="8" unit="m" value="1150.0">1150 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB637A56CFCB364A0064FA96" pageId="7" pageNumber="8" type="biology_ecology">
<paragraph id="8BB536FAFB637A56CFCB364A0038FAC8" blockId="7.[189,285,1293,1319]" box="[189,285,1293,1319]" pageId="7" pageNumber="8">
<heading id="D0FD8196FB637A56CFCB364A0038FAC8" bold="true" box="[189,285,1293,1319]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B97EEAE8FB637A56CFCB364A0038FAC8" bold="true" box="[189,285,1293,1319]" pageId="7" pageNumber="8">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB637A56CFCB367B0064FA96" blockId="7.[189,1398,1340,1401]" pageId="7" pageNumber="8">
Amathole Mistbelt Forest (Southern Mistbelt Forest group) (
<bibRefCitation id="EF9B4B0BFB637A56CCFD367B0584FAB9" author="Von Maltitz G. &amp; Mucina L. &amp; Geldenhuys C. &amp; Lawes M. &amp; Eeley H. &amp; Adie H. &amp; Vink D. &amp; Fleming G. &amp; Bailey C." box="[907,1185,1340,1366]" pageId="7" pageNumber="8" refId="ref16733" refString="Von Maltitz G., Mucina L., Geldenhuys C., Lawes M., Eeley H., Adie H., Vink D., Fleming G. &amp; Bailey C. 2003. Classification System for South African Indigenous Forests. Unpublished Report ENV-P-C 2003 - 017. CSIR, Pretoria." type="book" year="2003">
von Maltitz
<emphasis id="B97EEAE8FB637A56CB69367B0576FAB9" box="[1055,1107,1340,1366]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2003
</bibRefCitation>
); in leaf-litter and under logs.
</paragraph>
</subSubSection>
<subSubSection id="C3106571FB637A58CFCB36E70502FE63" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BB536FAFB637A56CFCB36E7000AFA55" blockId="7.[189,303,1440,1466]" box="[189,303,1440,1466]" pageId="7" pageNumber="8">
<heading id="D0FD8196FB637A56CFCB36E7000AFA55" bold="true" box="[189,303,1440,1466]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B97EEAE8FB637A56CFCB36E7000AFA55" bold="true" box="[189,303,1440,1466]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB637A56CFCB3688033EF8ED" blockId="7.[188,1400,1487,1795]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D79FB637A56CFCB3688005BFA06" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[189,382,1487,1513]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CFCB3688005BFA06" box="[189,382,1487,1513]" italics="true" pageId="7" pageNumber="8">Gulella fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CEFF368800C3FA06" box="[393,486,1487,1513]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C0A4D79FB637A56CDBF3688024EFA06" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[713,875,1487,1513]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CDBF3688024EFA06" box="[713,875,1487,1513]" italics="true" pageId="7" pageNumber="8">G. kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CC03368802F6FA06" box="[885,979,1487,1513]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
except that it has weak subsutural riblets and is larger (mean length
<quantity id="4CF29B1FFB637A56CD3936B50380F9E2" box="[591,677,1522,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="8" unit="mm" value="2.3">2.3mm</quantity>
and
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, respectively). The parietal lamella does not run parallel to the upper ridge of the labral slab, but the lowest point of its edge is in close proximity to the labral slab in the vicinity of the cusp. The labral complex is slightly differently shaped; the ridges of the labral slab have a V-shaped groove between them in
<taxonomicName id="4C0A4D79FB637A56CC16351B02C1F99A" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[864,996,1627,1654]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CC16351B02C1F99A" box="[864,996,1627,1654]" italics="true" pageId="7" pageNumber="8">G. fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CC9A351B0562F999" box="[1004,1095,1628,1654]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
The basal ridge to left of centre commences close to the lip edge and causes a shallow external furrow behind the lip, absent in
<taxonomicName id="4C0A4D79FB637A56CFCB35E50078F953" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[189,349,1698,1724]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CFCB35E50078F953" box="[189,349,1698,1724]" italics="true" pageId="7" pageNumber="8">G. kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CE1335E400E4F952" box="[357,449,1699,1725]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
In addition, the ridge-like middle tooth of the columella lamella is closer to the columella lip. Compared to other species described herein, the protoconch of
<taxonomicName id="4C0A4D79FB637A56CB4D35810598F930" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[1083,1213,1733,1760]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CB4D35810598F930" box="[1083,1213,1733,1760]" italics="true" pageId="7" pageNumber="8">G. fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CBB235810438F90F" box="[1220,1309,1734,1760]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
is large relative to the size of its shell.
</paragraph>
<paragraph id="8BB536FAFB637A58CFCB346D0502FE63" blockId="7.[189,1399,1834,2001]" lastBlockId="9.[188,1398,265,397]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
The labral slabs of
<taxonomicName id="4C0A4D79FB637A56CEC4346D0350F8AB" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[434,629,1834,1860]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CEC4346D0350F8AB" box="[434,629,1834,1860]" italics="true" pageId="7" pageNumber="8">Gulella fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CDF7346D03C4F8AB" box="[641,737,1834,1860]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0A4D79FB637A56CC55346D02E2F8AB" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[803,967,1834,1860]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB637A56CC55346D02E2F8AB" box="[803,967,1834,1860]" italics="true" pageId="7" pageNumber="8">G. kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB637A56CCA5346D0517F8AB" box="[979,1074,1834,1860]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
closely resemble those of
<taxonomicName id="4C0A4D79FB637A56CFCB340A0074F888" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[189,337,1869,1895]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="bomvana">
<emphasis id="B97EEAE8FB637A56CFCB340A0074F888" box="[189,337,1869,1895]" italics="true" pageId="7" pageNumber="8">G. bomvana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D79FB637A56CEE7340A0323F888" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[401,518,1869,1895]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tietzae">
<emphasis id="B97EEAE8FB637A56CEE7340A0323F888" box="[401,518,1869,1895]" italics="true" pageId="7" pageNumber="8">G. tietzae</emphasis>
</taxonomicName>
from the
<collectingRegion id="49CEF818FB637A56CDF234090206F888" box="[644,803,1869,1896]" country="South Africa" name="Eastern Cape" pageId="7" pageNumber="8">Eastern Cape</collectingRegion>
coast and several narrow-range species endemic to Zululand,
<taxonomicName id="4C0A4D79FB637A56CE1634360273F865" authority="(Melvill &amp; Ponsonby, 1903)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1903" box="[352,854,1904,1931]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="genialis">
<emphasis id="B97EEAE8FB637A56CE16343600CDF865" box="[352,488,1904,1931]" italics="true" pageId="7" pageNumber="8">G. genialis</emphasis>
(Melvill &amp; Ponsonby, 1903)
</taxonomicName>
,
<taxonomicName id="4C0A4D79FB637A56CC1E34360588F864" authority="Burnup, 1925" authorityName="Burnup" authorityYear="1925" box="[872,1197,1904,1931]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="laevorsa">
<emphasis id="B97EEAE8FB637A56CC1E343602D2F865" box="[872,1015,1904,1931]" italics="true" pageId="7" pageNumber="8">G. laevorsa</emphasis>
Burnup, 1925
</taxonomicName>
and
<taxonomicName id="4C0A4D79FB637A56CB8534360339F841" authority="(Melvill &amp; Ponsonby, 1907)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1907" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="vallaris">
<emphasis id="B97EEAE8FB637A56CB8534360452F865" box="[1267,1399,1904,1931]" italics="true" pageId="7" pageNumber="8">G. vallaris</emphasis>
(Melvill &amp; Ponsonby, 1907)
</taxonomicName>
(see
<taxonomicName id="4C0A4D79FB637A56CD1E34D403E5F842" authorityName="Pfeiffer" authorityYear="1856" box="[616,704,1939,1965]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="B97EEAE8FB637A56CD1E34D403E5F842" box="[616,704,1939,1965]" italics="true" pageId="7" pageNumber="8">Gulella</emphasis>
</taxonomicName>
Group
<quantity id="4CF29B1FFB637A56CC5134D3027EF842" box="[807,859,1939,1966]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="7" pageNumber="8" unit="in" value="9.0">9 in</quantity>
<bibRefCitation id="EF9B4B0BFB637A56CC1F34D405BAF841" author="Herbert D. &amp; Kilburn D." box="[873,1183,1939,1966]" pageId="7" pageNumber="8" refId="ref15954" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg, South Africa." type="book" year="2004">Herbert &amp; Kilburn 2004</bibRefCitation>
). These Zululand species, however, all possess strong axial riblets running from suture to suture. The tricuspid columella lamella of
<taxonomicName id="4C0A4D79FB6D7A58CEB8324E037CFECC" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[462,601,265,291]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB6D7A58CEB8324E037CFECC" box="[462,601,265,291]" italics="true" pageId="9" pageNumber="10">G. fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB6D7A58CD1E324D03EEFECB" box="[616,715,266,292]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0A4D79FB6D7A58CC62324E029FFECC" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[788,954,265,291]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="kenbrowni" status="sp. nov.">
<emphasis id="B97EEAE8FB6D7A58CC62324E029FFECC" box="[788,954,265,291]" italics="true" pageId="9" pageNumber="10">G. kenbrowni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB6D7A58CCBC324D0509FECB" box="[970,1068,266,292]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
closely resembles that of
<taxonomicName id="4C0A4D79FB6D7A58CFCB326A0077FEA9" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[189,338,300,327]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bomvana">
<emphasis id="B97EEAE8FB6D7A58CFCB326A0077FEA9" box="[189,338,300,327]" italics="true" pageId="9" pageNumber="10">G. bomvana</emphasis>
</taxonomicName>
and several other smooth-shelled
<taxonomicName id="4C0A4D79FB6D7A58CD8C326B0291FEA9" box="[762,948,300,326]" pageId="9" pageNumber="10">
<emphasis id="B97EEAE8FB6D7A58CD8C326B0277FEA9" box="[762,850,300,326]" italics="true" pageId="9" pageNumber="10">Gulella</emphasis>
species
</taxonomicName>
from the
<collectingRegion id="49CEF818FB6D7A58CB42326A05F1FEA9" box="[1076,1236,300,327]" country="South Africa" name="Eastern Cape" pageId="9" pageNumber="10">Eastern Cape</collectingRegion>
coast with a tricuspid columella,
<taxonomicName id="4C0A4D79FB6D7A58CEDC32080271FE85" authority="Connolly, 1939" authorityName="Connolly" authorityYear="1939" box="[426,852,335,362]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="aprosdoketa">
<emphasis id="B97EEAE8FB6D7A58CEDC320803BDFE86" box="[426,664,335,361]" italics="true" pageId="9" pageNumber="10">Gulella aprosdoketa</emphasis>
Connolly, 1939
</taxonomicName>
,
<taxonomicName id="4C0A4D79FB6D7A58CC16321702EAFE86" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[864,975,336,362]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tietzae">
<emphasis id="B97EEAE8FB6D7A58CC16321702EAFE86" box="[864,975,336,362]" italics="true" pageId="9" pageNumber="10">G. tietzae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D79FB6D7A58CB7032170457FE85" authority="Cole &amp; Herbert, 2009" authorityName="Cole &amp; Herbert" authorityYear="2009" box="[1030,1394,335,362]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="ndibo">
<emphasis id="B97EEAE8FB6D7A58CB703217054EFE86" box="[1030,1131,335,362]" italics="true" pageId="9" pageNumber="10">G. ndibo</emphasis>
<bibRefCitation id="EF9B4B0BFB6D7A58CB0532080457FE85" author="Cole M. L. &amp; Herbert D. G." box="[1139,1394,335,362]" pageId="9" pageNumber="10" pagination="547 - 564" refId="ref15568" refString="Cole M. L. &amp; Herbert D. G. 2009. Description of four new species of Gulella Pfeiffer, 1856 from Eastern Cape, South Africa, with additional notes on two poorly known species (Mollusca: Eupulmonata: Streptaxidae). Zoologische Mededelingen 83: 547 - 564." type="journal article" year="2009">Cole &amp; Herbert, 2009</bibRefCitation>
</taxonomicName>
, but in the latter three species the columella lamella extends to the lip edge.
</paragraph>
</subSubSection>
<caption id="DF756672FB6C7A59CFCB34C70309F80E" ID-DOI="http://doi.org/10.5281/zenodo.6448763" ID-Zenodo-Dep="6448763" httpUri="https://zenodo.org/record/6448763/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1920,1946]" targetBox="[239,1349,282,1880]" targetPageId="8">
<paragraph id="8BB536FAFB6C7A59CFCB34C70309F80E" blockId="8.[189,1399,1920,2018]" pageId="8" pageNumber="9">
<emphasis id="B97EEAE8FB6C7A59CFCB34C7002DF875" bold="true" box="[189,264,1920,1946]" pageId="8" pageNumber="9">Fig. 3</emphasis>
.
<taxonomicName id="4C0A4D79FB6C7A59CE6C34C700F9F875" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[282,476,1920,1946]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB6C7A59CE6C34C700F9F875" box="[282,476,1920,1946]" italics="true" pageId="8" pageNumber="9">Gulella fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB6C7A59CE9E34C6036CF874" box="[488,585,1921,1947]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMSA P1679/T4525), length 2.34 mm, width 1.06 mm.
<emphasis id="B97EEAE8FB6C7A59CFCB34E401F7F852" bold="true" box="[189,210,1955,1981]" pageId="8" pageNumber="9">A</emphasis>
. Aperture view.
<emphasis id="B97EEAE8FB6C7A59CED434E30093F851" bold="true" box="[418,438,1956,1982]" pageId="8" pageNumber="9">B</emphasis>
. Side view.
<emphasis id="B97EEAE8FB6C7A59CD2534E4034DF852" bold="true" box="[595,616,1955,1981]" pageId="8" pageNumber="9">C</emphasis>
. Oblique view into aperture.
<emphasis id="B97EEAE8FB6C7A59CCA134E302C9F851" bold="true" box="[983,1004,1956,1982]" pageId="8" pageNumber="9">D</emphasis>
. Oblique view of base showing umbilicus. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3106571FB6D7A58CFCB32FD026EFD16" pageId="9" pageNumber="10" type="conservation">
<paragraph id="8BB536FAFB6D7A58CFCB32FD0040FE3B" blockId="9.[189,357,442,468]" box="[189,357,442,468]" pageId="9" pageNumber="10">
<heading id="D0FD8196FB6D7A58CFCB32FD0040FE3B" bold="true" box="[189,357,442,468]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B97EEAE8FB6D7A58CFCB32FD0040FE3B" bold="true" box="[189,357,442,468]" pageId="9" pageNumber="10">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FAFB6D7A58CFCB32AE026EFD16" blockId="9.[187,1399,489,762]" pageId="9" pageNumber="10">
<taxonomicName id="4C0A4D79FB6D7A58CFCB32AE005FFDEC" authority="Cole &amp; Herbert, 2022" authorityName="Cole &amp; Herbert" authorityYear="2022" box="[189,378,489,515]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="fordycei" status="sp. nov.">
<emphasis id="B97EEAE8FB6D7A58CFCB32AE005FFDEC" box="[189,378,489,515]" italics="true" pageId="9" pageNumber="10">Gulella fordycei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5793FB6D7A58CEF632AE00FCFDEC" box="[384,473,489,515]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
has been found only at Fort Fordyce Nature Reserve, an outlier of the Amathole Mountains, despite extensive collecting efforts throughout the region. It therefore appears to be a very narrow-range endemic. Fort Fordyce is a protected area under the jurisdiction of the
<collectingRegion id="49CEF818FB6D7A58CBA131770452FDA6" box="[1239,1399,559,586]" country="South Africa" name="Eastern Cape" pageId="9" pageNumber="10">Eastern Cape</collectingRegion>
Parks and Tourism Agency. It is known for endemicity of other terrestrial molluscs, viz.
<taxonomicName id="4C0A4D79FB6D7A58CBB531150343FD7E" authority="Herbert, 2020" authorityName="Herbert" authorityYear="2020" class="Gastropoda" family="Charopidae" genus="Amatholedonta" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="fordycei">
<emphasis id="B97EEAE8FB6D7A58CBB531150039FD7F" italics="true" pageId="9" pageNumber="10">Amatholedonta fordycei</emphasis>
<bibRefCitation id="EF9B4B0BFB6D7A58CE55313100E0FD7F" author="Herbert D. G." box="[291,453,630,656]" pageId="9" pageNumber="10" pagination="1 - 55" refId="ref15900" refString="Herbert D. G. 2020. Revision of the aperturally dentate Charopidae (Gastropoda: Sylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies. European Journal of Taxonomy 629: 1 - 55." type="journal article" year="2020">Herbert, 2020</bibRefCitation>
(Charopidae)
</taxonomicName>
and an undescribed species of
<taxonomicName id="4C0A4D79FB6D7A58CCB831310456FD7E" authority="" box="[974,1395,630,657]" class="Gastropoda" family="Fauxulidae" genus="Fauxulus" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subGenus" subGenus="Anisoloma">
<emphasis id="B97EEAE8FB6D7A58CCB83131051EFD7F" box="[974,1083,630,656]" italics="true" pageId="9" pageNumber="10">Fauxulus</emphasis>
(
<emphasis id="B97EEAE8FB6D7A58CB3D313105EDFD7F" box="[1099,1224,630,656]" italics="true" pageId="9" pageNumber="10">Anisoloma</emphasis>
) (Fauxulidae)
</taxonomicName>
. Furthermore, specimens of another litter-dwelling invertebrate, the velvet worm
<taxonomicName id="4C0A4D79FB6D7A58CB1131DD004FFD39" authority="(Purcell, 1899)" baseAuthorityName="Purcell" baseAuthorityYear="1899" class="Udeonycophora" family="Peripatopsidae" genus="Peripatopsis" kingdom="Animalia" order="Euonycophora" pageId="9" pageNumber="10" phylum="Onychophora" rank="species" species="sedgwicki">
<emphasis id="B97EEAE8FB6D7A58CB1131DD0453FD5C" box="[1127,1398,665,691]" italics="true" pageId="9" pageNumber="10">Peripatopsis sedgwicki</emphasis>
(Purcell, 1899)
</taxonomicName>
from Fort Fordyce were genetically and morphologically distinct, suggesting the presence of a novel lineage at Fort Fordyce (
<bibRefCitation id="EF9B4B0BFB6D7A58CD213198021AFD16" author="Daniels S. &amp; Dreyer M. &amp; Sharma P." box="[599,831,735,762]" pageId="9" pageNumber="10" pagination="781 - 796" refId="ref15622" refString="Daniels S. R, Dreyer M. &amp; Sharma P. 2017. Contrasting the population genetic structure of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsis) in forest fragments along the south-eastern Cape, South Africa. Invertebrate Systematics 31: 781 - 796. https: // doi. org / 10.1071 / IS 16085" type="journal article" year="2017">
Daniels
<emphasis id="B97EEAE8FB6D7A58CDCB31A703D5FD16" box="[701,752,735,761]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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