treatments-xml/data/03/EE/87/03EE8796FF9DB50FE3E1B174FE2D51C8.xml

<document id="4A44999A1BBB822C07FF1B9CFD8EC620" ID-CLB-Dataset="7585" ID-DOI="10.11646/zootaxa.4942.4.3" ID-GBIF-Dataset="ddf1c8ab-ddf2-476d-a319-63046f6d99fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4612451" ID-ZooBank="38464B00-0721-4C27-98F9-599D862E4CB1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615992940915" checkinUser="plazi" docAuthor="Strümpher, Werner P. &amp; Stals, Riaan" docDate="2021" docId="03EE8796FF9DB50FE3E1B174FE2D51C8" docLanguage="en" docName="zootaxa.4942.4.3.pdf" docOrigin="Zootaxa 4942 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phoberus fumarius" docType="treatment" docVersion="7" lastPageNumber="555" masterDocId="FFD7FFEEFF9BB503E376B44EFF905649" masterDocTitle="Proper cleaning of keratin beetles reveals a cryptic species: Phoberus fumarius (Haaf, 1953) from southern Africa is reinstated as valid (Coleoptera: Trogidae)" masterLastPageNumber="557" masterPageNumber="543" pageNumber="549" updateTime="1698933609828" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9E9FAA876D4B002599A58426C9454FFB">Proper cleaning of keratin beetles reveals a cryptic species: Phoberus fumarius (Haaf, 1953) from southern Africa is reinstated as valid (Coleoptera: Trogidae)</mods:title>
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<mods:namePart id="B581DE72DBBE45E1020B5B490ADA15ED">Strümpher, Werner P.</mods:namePart>
<mods:affiliation id="50EA8201C67C34559FC9BD181901041A">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
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<mods:namePart id="8FEDF0D70259B90EF4BB4A1B8E891281">Stals, Riaan</mods:namePart>
<mods:affiliation id="BFA0B2062D652522A8A6911EFBBBE076">South African National Collection of Insects, Agricultural Research Council, Private Bag X 134, Queenswood 0121, Pretoria, South Africa.</mods:affiliation>
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<subSubSection id="C35D650BFF9DB505E3E1B174FC43531D" box="[151,979,1338,1365]" pageId="6" pageNumber="549" type="nomenclature">
<paragraph id="8BF83680FF9DB505E3E1B174FC43531D" blockId="6.[151,979,1338,1401]" box="[151,979,1338,1365]" pageId="6" pageNumber="549">
<heading id="D0B081ECFF9DB505E3E1B174FC43531D" bold="true" box="[151,979,1338,1365]" fontSize="11" level="1" pageId="6" pageNumber="549" reason="1">
<emphasis id="B933EA92FF9DB505E3E1B174FC43531D" bold="true" box="[151,979,1338,1365]" pageId="6" pageNumber="549">
<taxonomicName id="4C474D03FF9DB505E3E1B174FDB4531D" ID-CoL="7ZQBM" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[151,548,1338,1365]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E3E1B174FE11531D" bold="true" box="[151,385,1338,1365]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
(
<bibRefCitation id="EFD64B71FF9DB505E2E6B174FD8C531D" author="Haaf, E." box="[400,540,1338,1364]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
, revalidated and new combination
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF9DB505E3E1B111FEFE5330" box="[151,366,1375,1401]" pageId="6" pageNumber="549" type="description">
<paragraph id="8BF83680FF9DB505E3E1B111FEFE5330" blockId="6.[151,979,1338,1401]" box="[151,366,1375,1401]" pageId="6" pageNumber="549">
(
<figureCitation id="137C2A05FF9DB505E3E9B111FE825330" box="[159,274,1375,1401]" captionStart-0="FIGURES 8–10" captionStart-1="FIGURES 11–14" captionStartId-0="8.[151,264,1932,1956]" captionStartId-1="9.[151,264,1943,1967]" captionTargetBox-0="[182,1402,193,1906]" captionTargetBox-1="[203,1379,197,1904]" captionTargetId-0="figure-31@8.[167,1419,181,1908]" captionTargetId-1="figure-17@9.[188,1399,181,1919]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 8–10. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." captionText-1="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi-0="http://doi.org/10.5281/zenodo.4612459" figureDoi-1="http://doi.org/10.5281/zenodo.4612463" httpUri-0="https://zenodo.org/record/4612459/files/figure.png" httpUri-1="https://zenodo.org/record/4612463/files/figure.png" pageId="6" pageNumber="549">Figs 8–14</figureCitation>
, 
<figureCitation id="137C2A05FF9DB505E268B111FEAC5330" box="[286,316,1375,1401]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1442,1466]" captionTargetBox="[171,1416,181,1419]" captionTargetId="figure-31@10.[170,1416,181,1419]" captionTargetPageId="10" captionText="FIGURES 15–16. Pronotal disc, dorsal view. 15, Phoberus cyrtus (Haaf, 1953); 16, Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612465" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="6" pageNumber="549">16</figureCitation>
, 
<figureCitation id="137C2A05FF9DB505E23EB111FEF55330" box="[328,357,1375,1401]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="6" pageNumber="549">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF9DB505E3E1B1EAFB6253B0" pageId="6" pageNumber="549" type="reference_group">
<paragraph id="8BF83680FF9DB505E3E1B1EAFB6253B0" blockId="6.[151,1437,1443,1529]" pageId="6" pageNumber="549">
<treatmentCitationGroup id="AB5711AEFF9DB505E3E1B1EAFB6253B0" pageId="6" pageNumber="549">
<treatmentCitation id="0AE61091FF9DB505E3E1B1EAFE5E53F2" author="Haaf, E." box="[151,462,1443,1467]" page="334" pageId="6" pageNumber="549" year="1953">
<taxonomicName id="4C474D03FF9DB505E3E1B1EAFE5E53F2" ID-CoL="7Z6KF" authority="Haaf, 1953: 334" authorityName="Haaf" authorityPageNumber="334" authorityYear="1953" box="[151,462,1443,1467]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E3E1B1EAFEB553F2" box="[151,293,1443,1467]" italics="true" pageId="6" pageNumber="549">Trox fumarius</emphasis>
<bibRefCitation id="EFD64B71FF9DB505E25DB1EDFE5E53F2" author="Haaf, E." box="[299,462,1443,1467]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953: 334</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description); 
<treatmentCitation id="0AE61091FF9DB505E1C1B1EDFCC453F2" author="Haaf, E." box="[695,852,1443,1467]" page="473" pageId="6" pageNumber="549" year="1959">
<bibRefCitation id="EFD64B71FF9DB505E1C1B1EDFCC453F2" author="Haaf, E." box="[695,852,1443,1467]" pageId="6" pageNumber="549" refId="ref7702" refString="Haaf, E. (1959) Coleoptera: Trogidae. In: Hanstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life. Results of the Lund University Expedition in 1950 - 1951. Vol. 5. Almqvist &amp; Wiksell, Uppsala, pp. 473 - 478. [1958]" type="journal volume" year="1959">Haaf 1959: 473</bibRefCitation>
</treatmentCitation>
(checklist, localities); 
<treatmentCitation id="0AE61091FF9DB505E74FB1EDFB7F53F2" author="Scholtz, C. H." box="[1081,1263,1443,1467]" page="97" pageId="6" pageNumber="549" year="1980">
<bibRefCitation id="EFD64B71FF9DB505E74FB1EDFB7F53F2" author="Scholtz, C. H." box="[1081,1263,1443,1467]" pageId="6" pageNumber="549" pagination="1 - 104" refId="ref8073" refString="Scholtz, C. H. (1980 a) Monograph of the genus Trox F. (Coleoptera: Trogidae) of subsaharan Africa. Cimbebasia Memoir, 4, 1 - 104." type="journal article" year="1980">Scholtz 1980a: 97</bibRefCitation>
</treatmentCitation>
(new junior synonym of 
<taxonomicName id="4C474D03FF9DB505E252B18DFE4B5393" ID-CoL="59DJJ" authorityName="Haaf" authorityYear="1953" box="[292,475,1474,1498]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus" subGenus="Trox">
<emphasis id="B933EA92FF9DB505E252B18DFEC25393" box="[292,338,1475,1498]" italics="true" pageId="6" pageNumber="549">Trox</emphasis>
(
<emphasis id="B933EA92FF9DB505E216B18DFE1E5393" box="[352,398,1475,1498]" italics="true" pageId="6" pageNumber="549">Trox</emphasis>
) 
<emphasis id="B933EA92FF9DB505E2EBB18DFE4B5393" box="[413,475,1475,1498]" italics="true" pageId="6" pageNumber="549">cyrtus</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AE61091FF9DB505E287B18CFD015393" author="Scholtz, C. H." box="[497,657,1474,1498]" page="3" pageId="6" pageNumber="549" year="1982">
<bibRefCitation id="EFD64B71FF9DB505E287B18CFD015393" author="Scholtz, C. H." box="[497,657,1474,1498]" pageId="6" pageNumber="549" pagination="1 - 27" refId="ref8146" refString="Scholtz, C. H. (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries, Republic of South Africa, 54, 1 - 27." type="journal article" year="1982">Scholtz 1982: 3</bibRefCitation>
</treatmentCitation>
(catalogue, as synonym of 
<taxonomicName id="4C474D03FF9DB505E0DCB18DFBF15393" ID-CoL="59DJJ" authorityName="Haaf" authorityYear="1953" box="[938,1121,1474,1498]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus" subGenus="Trox">
<emphasis id="B933EA92FF9DB505E0DCB18DFC485393" box="[938,984,1475,1498]" italics="true" pageId="6" pageNumber="549">Trox</emphasis>
(
<emphasis id="B933EA92FF9DB505E090B18DFB845393" box="[998,1044,1475,1498]" italics="true" pageId="6" pageNumber="549">Trox</emphasis>
) 
<emphasis id="B933EA92FF9DB505E755B18DFBF15393" box="[1059,1121,1475,1498]" italics="true" pageId="6" pageNumber="549">cyrtus</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AE61091FF9DB505E701B18CFA985393" author="Zidek, J." box="[1143,1288,1474,1498]" page="9" pageId="6" pageNumber="549" year="2013">
<bibRefCitation id="EFD64B71FF9DB505E701B18CFA985393" author="Zidek, J." box="[1143,1288,1474,1498]" pageId="6" pageNumber="549" pagination="1 - 38" refId="ref8671" refString="Zidek, J. (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi, 314, 1 - 38." type="journal article" year="2013">Zidek 2013: 9</bibRefCitation>
</treatmentCitation>
, 10 (checklist, as synonym of 
<taxonomicName id="4C474D03FF9DB505E217B1AFFDDB53B1" ID-CoL="59DJJ" authorityName="Haaf" authorityYear="1953" box="[353,587,1504,1529]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus" subGenus="Phoberus">
<emphasis id="B933EA92FF9DB505E217B1AFFE1F53B1" box="[353,399,1505,1528]" italics="true" pageId="6" pageNumber="549">Trox</emphasis>
(
<emphasis id="B933EA92FF9DB505E2E8B1AEFE6F53B1" box="[414,511,1504,1528]" italics="true" pageId="6" pageNumber="549">Phoberus</emphasis>
) 
<emphasis id="B933EA92FF9DB505E17BB1AFFDDB53B1" box="[525,587,1505,1528]" italics="true" pageId="6" pageNumber="549">cyrtus</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0AE61091FF9DB505E117B1AEFD6F53B0" author="Zidek, J." box="[609,767,1504,1529]" page="99" pageId="6" pageNumber="549" year="2017">
<bibRefCitation id="EFD64B71FF9DB505E117B1AEFD6F53B0" author="Zidek, J." box="[609,767,1504,1529]" pageId="6" pageNumber="549" pagination="93 - 127" refId="ref8699" refString="Zidek, J. (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana, Series A, 25 (1), 93 - 127." type="journal article" year="2017">Zídek 2017: 99</bibRefCitation>
</treatmentCitation>
, 101 (checklist, as synonym of 
<taxonomicName id="4C474D03FF9DB505E748B1AEFB7453B1" ID-CoL="4GJWC" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1086,1252,1504,1528]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9DB505E748B1AEFB7453B1" box="[1086,1252,1504,1528]" italics="true" pageId="6" pageNumber="549">Phoberus cyrtus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF9DB504E3E1B269FBF2528C" lastPageId="7" lastPageNumber="550" pageId="6" pageNumber="549" type="materials_examined">
<paragraph id="8BF83680FF9DB505E3E1B269FABF500B" blockId="6.[151,1437,1574,2033]" box="[151,1327,1574,1602]" pageId="6" pageNumber="549">
<materialsCitation id="3B2F3CDDFF9DB505E3E1B269FABF500B" ID-GBIF-Occurrence="3062300339" box="[151,1327,1574,1602]" country="South Africa" county="Sarah Baartman District" location="Willowmore" municipality="Sarah Baartman District Municipality" pageId="6" pageNumber="549" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<emphasis id="B933EA92FF9DB505E3E1B269FEA65009" bold="true" box="[151,310,1574,1601]" pageId="6" pageNumber="549">
<typeStatus id="54FC8822FF9DB505E3E1B269FF425008" box="[151,210,1575,1601]" pageId="6" pageNumber="549">Type</typeStatus>
locality.
</emphasis>
<location id="8E98605BFF9DB505E24AB269FE585008" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9DB505E24AB269FE585008" box="[316,456,1575,1601]" country="South Africa" county="Sarah Baartman District" municipality="Sarah Baartman District Municipality" name="Willowmore" pageId="6" pageNumber="549" stateProvince="Eastern Cape">Willowmore</location>
(
<collectingMunicipality id="6B9CACFAFF9DB505E2A1B269FC105008" box="[471,896,1575,1601]" pageId="6" pageNumber="549">
<collectingCounty id="62994E0CFF9DB505E2A1B269FD775008" box="[471,743,1575,1601]" pageId="6" pageNumber="549">Sarah Baartman District</collectingCounty>
Municipality
</collectingMunicipality>
, 
<collectingRegion id="4983F862FF9DB505E0FDB269FB1C5008" box="[907,1164,1575,1601]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="549">Eastern Cape Province</collectingRegion>
, 
<collectingCountry id="F3507610FF9DB505E7EEB269FAB9500B" box="[1176,1321,1575,1602]" name="South Africa" pageId="6" pageNumber="549">South Africa</collectingCountry>
)
</materialsCitation>
</paragraph>
<paragraph id="8BF83680FF9DB505E3B1B205FD1051B8" blockId="6.[151,1437,1574,2033]" pageId="6" pageNumber="549">
<emphasis id="B933EA92FF9DB505E3B1B205FDFA502A" bold="true" box="[199,618,1610,1637]" pageId="6" pageNumber="549">
Type material examined. 
<typeStatus id="54FC8822FF9DB505E283B205FDFA502A" box="[501,618,1611,1637]" pageId="6" pageNumber="549" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
Ƌ (
<collectionCode id="ED56AE45FF9DB505E1E2B205FD7C502C" box="[660,748,1611,1637]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="6" pageNumber="549">NHMB</collectionCode>
, aedeagus extracted, examined through detailed photographs) (
<figureCitation id="137C2A05FF9DB505E3E9B221FE8750C0" box="[159,279,1647,1673]" captionStart="FIGURES 8–10" captionStartId="8.[151,264,1932,1956]" captionTargetBox="[182,1402,193,1906]" captionTargetId="figure-31@8.[167,1419,181,1908]" captionTargetPageId="8" captionText="FIGURES 8–10. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." figureDoi="http://doi.org/10.5281/zenodo.4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="6" pageNumber="549">Figs 8–10</figureCitation>
): Rectangle, greyish white: “[
<collectingCountry id="F3507610FF9DB505E11DB221FC9450C0" box="[619,772,1647,1673]" name="South Africa" pageId="6" pageNumber="549">South Africa</collectingCountry>
, 
<collectingRegion id="4983F862FF9DB505E064B221FB8A50C0" box="[786,1050,1646,1673]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="549">Eastern Cape Province</collectingRegion>
] Willowmore [
<geoCoordinate id="EE735047FF9DB505E7B1B23EFAB250C2" box="[1223,1314,1648,1675]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="6" pageNumber="549" precision="925" value="-33.283333">33°17′S</geoCoordinate>
<geoCoordinate id="EE735047FF9DB505E65FB23EFA1650C2" box="[1321,1414,1648,1675]" degrees="23" direction="east" minutes="29" orientation="longitude" pageId="6" pageNumber="549" precision="925" value="23.483334">23°29′E</geoCoordinate>
] | 30.10.[19]48.” || Rectangle, white: “Süd-Afrika | leg. C. Koch” || Rectangle, white: “Museum Frey | Tutzing” || Rectangle, white with red bars at top and bottom, preprinted red text interspersed with black handwriting: “HOLOTYPUS | 
<taxonomicName id="4C474D03FF9DB505E273B295FEAA50BC" authorityName="Fabricius" authorityYear="1775" box="[261,314,1755,1781]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="genus">Trox</taxonomicName>
| fumarius 
<taxonomicNameLabel id="A20057E9FF9DB505E2CDB295FE6050BC" box="[443,496,1755,1781]" pageId="6" pageNumber="549" rank="species">sp.n.</taxonomicNameLabel>
| det.Dr.E. 
<bibRefCitation id="EFD64B71FF9DB505E113B295FD6850BC" author="Haaf, E." box="[613,760,1754,1781]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">HAAF 1953</bibRefCitation>
” || Rectangle, white, preprinted black text interspersed with black handwriting: “ 
<taxonomicName id="4C474D03FF9DB505E208B2B1FDB35150" authorityName="Haaf" authorityYear="1953" box="[382,547,1791,1817]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">Trox fumarius</taxonomicName>
| Haaf (= 
<taxonomicName id="4C474D03FF9DB505E1EDB2B1FD405150" authorityName="Fabricius" authorityYear="1775" box="[667,720,1791,1817]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="genus">Trox</taxonomicName>
| 
<taxonomicName id="4C474D03FF9DB505E191B2B1FCE25150" authority="Haaf." authorityName="Haaf." box="[743,882,1791,1817]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="cyrtus">cyrtus Haaf.</taxonomicName>
| det. C.H. SCHOLTZ 1979” || Rectangle, white cardstock with black margin: “ 
<taxonomicName id="4C474D03FF9DB505E286B36DFD575174" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[496,711,1827,1853]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9DB505E286B36DFD575174" box="[496,711,1827,1853]" italics="true" pageId="6" pageNumber="549">Phoberus fumarius</emphasis>
</taxonomicName>
| (
<bibRefCitation id="EFD64B71FF9DB505E194B36DFCF15174" author="Haaf, E." box="[738,865,1826,1853]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
) | stat. rev., comb. nov. | Strümpher &amp; Stals 2021”. 
<emphasis id="B933EA92FF9DB505E3E1B309FE975116" bold="true" box="[151,263,1863,1889]" pageId="6" pageNumber="549">
<typeStatus id="54FC8822FF9DB505E3E1B309FE975116" box="[151,263,1863,1889]" pageId="6" pageNumber="549" type="paratype">PARATYPE</typeStatus>
</emphasis>
1Ƌ (
<collectionCode id="ED56AE45FF9DB505E248B309FE005128" box="[318,400,1863,1889]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="6" pageNumber="549">TMSA</collectionCode>
, aedeagus extracted): Rectangle, yellowing white: “Willowmore, | Capland. | Dr. H. Brauns.” || Narrow rectangle, red: “ 
<typeStatus id="54FC8822FF9DB505E2C9B325FDC151CC" box="[447,593,1899,1925]" pageId="6" pageNumber="549" type="paratype">PARATYPE</typeStatus>
” || Rectangle, white, preprinted text interspersed with handwriting: “ 
<taxonomicName id="4C474D03FF9DB505E612B325FA0C51CC" authorityName="Fabricius" authorityYear="1775" box="[1380,1436,1899,1925]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="genus">Trox</taxonomicName>
| 
<taxonomicName id="4C474D03FF9DB505E3D0B3C1FED951E0" authority="Haaf" authorityName="Haaf" box="[166,329,1935,1961]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">fumarius Haaf</taxonomicName>
| det.Dr.E. 
<bibRefCitation id="EFD64B71FF9DB505E2B7B3C1FDAF51E0" author="Haaf, E." box="[449,575,1934,1961]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf 1953</bibRefCitation>
” || Rectangle, white cardstock with orange margin: “ 
<typeStatus id="54FC8822FF9DB505E7EBB3C1FA9E51E0" box="[1181,1294,1935,1961]" pageId="6" pageNumber="549" type="paratype">Paratypus</typeStatus>
1953 | 
<taxonomicName id="4C474D03FF9DB505E611B3C1FF6B5184" authority="Strümpher &amp; Stals, 2021" authorityName="Strümpher &amp; Stals" authorityYear="2021" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius" status="sp. nov.">Trox fumarius</taxonomicName>
<taxonomicNameLabel id="A20057E9FF9DB505E274B3FDFEA05184" box="[258,304,1971,1997]" pageId="6" pageNumber="549" rank="species">sp.n</taxonomicNameLabel>
| E. Haaf” || Rectangle, white cardstock with black margin: “ 
<emphasis id="B933EA92FF9DB505E0A8B3FDFB545184" box="[990,1220,1971,1997]" italics="true" pageId="6" pageNumber="549">
<taxonomicName id="4C474D03FF9DB505E0A8B3FDFB255184" box="[990,1205,1971,1997]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fumarius">Phoberus fumarius</taxonomicName>
|
</emphasis>
(
<bibRefCitation id="EFD64B71FF9DB505E7A4B3FDFAC25184" author="Haaf, E." box="[1234,1362,1970,1997]" pageId="6" pageNumber="549" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
) | stat. rev., comb. nov. | Strümpher &amp; Stals 2021”.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B4D9FA8856B0" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<bibRefCitation id="EFD64B71FF9CB504E3B1B4D9FEC056F8" author="Haaf, E." box="[199,336,150,177]" pageId="7" pageNumber="550" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf (1953)</bibRefCitation>
described 
<taxonomicName id="4C474D03FF9CB504E2BCB4D9FDF856F8" authorityName="Strümpher &amp; Stals" authorityYear="2021" box="[458,616,151,177]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF9CB504E2BCB4D9FDF856F8" box="[458,616,151,177]" italics="true" pageId="7" pageNumber="550">Trox fumarius</emphasis>
</taxonomicName>
from three males and one female, deposited in 
<collectionCode id="ED56AE45FF9CB504E7F4B4D9FB4656F8" box="[1154,1238,151,177]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="550">NHMB</collectionCode>
(among these we studied the male 
<typeStatus id="54FC8822FF9CB504E225B4F5FE2A569F" box="[339,442,187,214]" pageId="7" pageNumber="550" type="holotype">holotype</typeStatus>
); one male in the Hungarian Natural History Museum, Budapest (not seen by us); and one male and one female in 
<collectionCode id="ED56AE45FF9CB504E2D5B491FE6456B0" box="[419,500,223,249]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
, of which we studied the male 
<typeStatus id="54FC8822FF9CB504E039B491FC3F56B0" box="[847,943,223,249]" pageId="7" pageNumber="550" type="paratype">paratype</typeStatus>
but could not locate the female.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B54CFBF2528C" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B54CFDA05755" bold="true" box="[199,560,258,284]" pageId="7" pageNumber="550">Additional material examined.</emphasis>
<materialsCitation id="3B2F3CDDFF9CB504E143B54CFB475708" ID-GBIF-Occurrence="3062300306" collectingDate="2001-09-06" collectionCode="TMSA" country="South Africa" latitude="-28.316668" location="Paradieskloof" longLatPrecision="1232" longitude="17.0" municipality="Richtersveld" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Northern Cape">
<emphasis id="B933EA92FF9CB504E143B54CFBAC5755" bold="true" box="[565,1084,258,285]" pageId="7" pageNumber="550">
<collectingCountry id="F3507610FF9CB504E143B54CFC995755" box="[565,777,258,284]" name="South Africa" pageId="7" pageNumber="550">SOUTH AFRICA</collectingCountry>
. 
<collectingRegion id="4983F862FF9CB504E065B54DFBA75755" box="[787,1079,258,285]" country="South Africa" name="Northern Cape" pageId="7" pageNumber="550">Northern Cape Province</collectingRegion>
.
</emphasis>
1♁ 
<collectingMunicipality id="6B9CACFAFF9CB504E71FB54DFB695754" box="[1129,1273,259,285]" pageId="7" pageNumber="550">Richtersveld</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E674B54DFA0C5754" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E674B54DFA0C5754" box="[1282,1436,259,285]" country="South Africa" latitude="-28.316668" longLatPrecision="1232" longitude="17.0" municipality="Richtersveld" name="Paradieskloof" pageId="7" pageNumber="550" stateProvince="Northern Cape">Paradieskloof</location>
[
<emphasis id="B933EA92FF9CB504E3D7B569FF485709" box="[161,216,295,320]" italics="true" pageId="7" pageNumber="550">recte</emphasis>
Paradyskloof], 
<geoCoordinate id="EE735047FF9CB504E2E6B566FE7A570A" box="[400,490,296,323]" degrees="28" direction="south" minutes="19" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-28.316668">28°19′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E282B566FDC1570A" box="[500,593,296,323]" degrees="17" direction="east" minutes="00" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.0">17°00′E</geoCoordinate>
, 
<date id="FFF91040FF9CB504E116B569FD4A5708" box="[608,730,295,321]" pageId="7" pageNumber="550" value="2001-09-06">
<collectingDate id="EFBDE9A8FF9CB504E116B569FD4A5708" box="[608,730,295,321]" pageId="7" pageNumber="550" value="2001-09-06">06.ix.2001</collectingDate>
</date>
, E-Y:3412, 
<collectingMethod id="52064E97FF9CB504E010B569FC425708" box="[870,978,295,321]" pageId="7" pageNumber="550">light trap</collectingMethod>
, 
<collectionCode id="ED56AE45FF9CB504E097B569FBBD5708" box="[993,1069,295,321]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
staff (
<collectionCode id="ED56AE45FF9CB504E70DB569FB5F5708" box="[1147,1231,295,321]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E793B569FDAC572C" ID-GBIF-Occurrence="3062300328" collectingDate="2001-09-06" collectionCode="TMSA" country="South Africa" latitude="-28.316668" location="Paradieskloof" longLatPrecision="1232" longitude="17.0" municipality="Richtersveld" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="9D41FD09FF9CB504E793B569FA985708" box="[1253,1288,295,321]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<emphasis id="B933EA92FF9CB504E665B569FAD95708" box="[1299,1353,295,321]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but E- Y:3413, on vegetation [
<emphasis id="B933EA92FF9CB504E2D0B505FE55572D" box="[422,453,331,356]" italics="true" pageId="7" pageNumber="550">sic</emphasis>
] (
<collectionCode id="ED56AE45FF9CB504E296B505FDA4572C" box="[480,564,331,357]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E13FB505FED157C0" ID-GBIF-Occurrence="3062300329" collectingDate="1955-10" collectionCode="TMSA" collectorName="C Koch" country="South Africa" latitude="-28.366667" location="Gelykswerf" longLatPrecision="1232" longitude="17.133333" municipality="Richtersveld" pageId="7" pageNumber="550" specimenCount="2" specimenCount-female="2" stateProvince="Northern Cape">
<specimenCount id="9D41FD09FF9CB504E13FB505FDFC572C" box="[585,620,331,357]" pageId="7" pageNumber="550" type="female">2♀</specimenCount>
<collectingMunicipality id="6B9CACFAFF9CB504E103B505FC95572C" box="[629,773,331,357]" pageId="7" pageNumber="550">Richtersveld</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E065B505FC06572C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E065B505FC06572C" box="[787,918,331,357]" country="South Africa" latitude="-28.366667" longLatPrecision="1232" longitude="17.133333" municipality="Richtersveld" name="Gelykswerf" pageId="7" pageNumber="550" stateProvince="Northern Cape">Gelykswerf</location>
[Mountain], [c. 
<geoCoordinate id="EE735047FF9CB504E723B502FB3E572E" box="[1109,1198,332,359]" degrees="28" direction="south" minutes="22" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-28.366667">28°22′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E7CEB502FA87572E" box="[1208,1303,332,359]" degrees="17" direction="east" minutes="08" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.133333">17°08′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E65AB504FAEB572C" box="[1324,1403,330,357]" pageId="7" pageNumber="550" value="1955-10">
<collectingDate id="EFBDE9A8FF9CB504E65AB504FAEB572C" box="[1324,1403,330,357]" pageId="7" pageNumber="550" value="1955-10">x.1955</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E6FCB505FF4457C0" pageId="7" pageNumber="550">C Koch</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E392B521FEA857C0" box="[228,312,367,393]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E23BB53EFEFB57E4" ID-GBIF-Occurrence="3062300325" collectingDate="1966-10-02" collectingDateMax="1966-10-03" collectingDateMin="1966-10-02" collectionCode="TMSA" country="South Africa" latitude="-30.8" location="Swart Doring River" longLatPrecision="1220" longitude="17.9" municipality="Namaqualand" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Northern Cape">
1♁ 
<collectingMunicipality id="6B9CACFAFF9CB504E20EB521FD8657C0" box="[376,534,367,393]" pageId="7" pageNumber="550">Namaqualand</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E152B521FC9657C0" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E152B521FC9657C0" box="[548,774,367,393]" country="South Africa" latitude="-30.8" longLatPrecision="1220" longitude="17.9" municipality="Namaqualand" name="Swart Doring River" pageId="7" pageNumber="550" stateProvince="Northern Cape">Swart Doring River</location>
, [
<geoCoordinate id="EE735047FF9CB504E06FB53EFCE557C2" box="[793,885,368,395]" degrees="30" direction="south" minutes="48" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-30.8">30°48′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E00BB53EFC4D57C2" box="[893,989,368,395]" degrees="17" direction="east" minutes="54" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.9">17°54′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E087B521FB1D57C0" box="[1009,1165,367,393]" pageId="7" pageNumber="550" value="1966-10-02" valueMax="1966-10-03" valueMin="1966-10-02">
<collectingDate id="EFBDE9A8FF9CB504E087B521FB1D57C0" box="[1009,1165,367,393]" pageId="7" pageNumber="550" value="1966-10-02" valueMax="1966-10-03" valueMin="1966-10-02">02–03.x.1966</collectingDate>
</date>
, S.A.M. [South African Museum] (
<collectionCode id="ED56AE45FF9CB504E279B5DDFEF357E4" box="[271,355,403,429]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E205B5DAFDC45798" ID-GBIF-Occurrence="3062300311" collectingDate="1979-08-24" collectingDateMax="1979-10-26" collectingDateMin="1979-08-24" collectionCode="TMSA" collectorName="S Endrody-Younga" country="South Africa" latitude="-30.783333" location="Dermbergsdraai Farm" longLatPrecision="1220" longitude="17.716667" municipality="Namaqualand" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Northern Cape">
1♁ 
<collectingMunicipality id="6B9CACFAFF9CB504E2EDB5DDFDA957E4" box="[411,569,403,429]" pageId="7" pageNumber="550">Namaqualand</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E134B5DDFCA957E4" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E134B5DDFCA957E4" box="[578,825,403,429]" country="South Africa" latitude="-30.783333" longLatPrecision="1220" longitude="17.716667" municipality="Namaqualand" name="Dermbergsdraai Farm" pageId="7" pageNumber="550" stateProvince="Northern Cape">Dermbergsdraai Farm</location>
, 
<geoCoordinate id="EE735047FF9CB504E034B5DAFC0A57E6" box="[834,922,404,431]" degrees="30" direction="south" minutes="47" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-30.783333">30°47′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E0E9B5DAFC6C57E6" box="[927,1020,404,431]" degrees="17" direction="east" minutes="43" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.716667">17°43′E</geoCoordinate>
, 
<collectingMethod id="52064E97FF9CB504E773B5DDFB1857E4" box="[1029,1160,403,429]" pageId="7" pageNumber="550">pitfall traps</collectingMethod>
: 
<date id="FFF91040FF9CB504E7E6B5DDFA8957E4" box="[1168,1305,403,429]" pageId="7" pageNumber="550" value="1979-08-24">
<collectingDate id="EFBDE9A8FF9CB504E7E6B5DDFA8957E4" box="[1168,1305,403,429]" pageId="7" pageNumber="550" value="1979-08-24">24.viii.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E65FB5DDFA0857E4" box="[1321,1432,403,429]" pageId="7" pageNumber="550" value="1979-10-26">
<collectingDate id="EFBDE9A8FF9CB504E65FB5DDFA0857E4" box="[1321,1432,403,429]" pageId="7" pageNumber="550" value="1979-10-26">26.x.1979</collectingDate>
</date>
, E-Y:1589, 
<collectorName id="26B25356FF9CB504E264B5F9FE795798" box="[274,489,439,465]" pageId="7" pageNumber="550">S Endrödy-Younga</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E28EB5F9FDDC5798" box="[504,588,439,465]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E129B5F6FD8757BF" ID-GBIF-Occurrence="3062300308" collectingDate="2003-09-01" collectingDateMax="2003-09-13" collectingDateMin="2003-09-01" collectionCode="UPSA" collectorName="A Frolov &amp; C Deschodt" country="South Africa" latitude="-30.2" location="Kamieskroon" longLatPrecision="1222" longitude="17.933332" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Northern Cape">
1♁ 
<location id="8E98605BFF9CB504E1FFB5F9FCB35798" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E1FFB5F9FCB35798" box="[649,803,439,465]" country="South Africa" latitude="-30.2" longLatPrecision="1222" longitude="17.933332" name="Kamieskroon" pageId="7" pageNumber="550" stateProvince="Northern Cape">Kamieskroon</location>
, 
<geoCoordinate id="EE735047FF9CB504E059B5F6FC18579A" box="[815,904,440,467]" degrees="30" direction="south" minutes="12" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-30.2">30°12′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E0F9B5F6FC7C579A" box="[911,1004,440,467]" degrees="17" direction="east" minutes="56" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.933332">17°56′E</geoCoordinate>
, 
<date id="FFF91040FF9CB504E08EB5F9FB0C5798" box="[1016,1180,439,465]" pageId="7" pageNumber="550" value="2003-09-01" valueMax="2003-09-13" valueMin="2003-09-01">
<collectingDate id="EFBDE9A8FF9CB504E08EB5F9FB0C5798" box="[1016,1180,439,465]" pageId="7" pageNumber="550" value="2003-09-01" valueMax="2003-09-13" valueMin="2003-09-01">01–13.ix.2003</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E7D1B5F9FA9D5798" box="[1191,1293,439,465]" pageId="7" pageNumber="550">A Frolov</collectorName>
, 
<collectorName id="26B25356FF9CB504E66EB5F9FA0C5798" box="[1304,1436,439,465]" pageId="7" pageNumber="550">C Deschodt</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E3E9B595FF7B57BC" box="[159,235,475,501]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="7" pageNumber="550">UPSA</collectionCode>
, preserved in 99% ethanol)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E168B595FE035450" ID-GBIF-Occurrence="3062300330" collectingDate="1930-06" collectionCode="SAMC" collectorName="Museum" country="South Africa" latitude="-30.558332" location="Garies" longLatPrecision="797" longitude="17.983334" municipality="Namaqualand" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Northern Cape">
<specimenCount id="9D41FD09FF9CB504E168B595FDD157BC" box="[542,577,475,501]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<collectingMunicipality id="6B9CACFAFF9CB504E130B595FD7457BC" box="[582,740,475,501]" pageId="7" pageNumber="550">Namaqualand</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E19BB595FCA957BC" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E19BB595FCA957BC" box="[749,825,475,501]" country="South Africa" latitude="-30.558332" longLatPrecision="797" longitude="17.983334" municipality="Namaqualand" name="Garies" pageId="7" pageNumber="550" stateProvince="Northern Cape">Garies</location>
, [
<geoCoordinate id="EE735047FF9CB504E03EB592FC5B57BE" box="[840,971,476,503]" degrees="30" direction="south" minutes="33" orientation="latitude" pageId="7" pageNumber="550" precision="15" seconds="30" value="-30.558332">30°33′30″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E0A6B592FBBF57BE" box="[976,1071,476,503]" degrees="17" direction="east" minutes="59" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="17.983334">17°59′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E749B595FB0657BC" box="[1087,1174,475,501]" pageId="7" pageNumber="550" value="1930-06">
<collectingDate id="EFBDE9A8FF9CB504E749B595FB0657BC" box="[1087,1174,475,501]" pageId="7" pageNumber="550" value="1930-06">vi.1930</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E7E9B595FA9157BC" box="[1183,1281,475,501]" pageId="7" pageNumber="550">Museum</collectorName>
staff (
<collectionCode id="ED56AE45FF9CB504E632B595FA0957BC" box="[1348,1433,475,501]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="550" type="Museum">SAMC</collectionCode>
: SAM-COL-A082505)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E2E9B5B1FD725474" ID-GBIF-Occurrence="3062300344" collectingDate="2003-11-27" collectionCode="TMSA" country="South Africa" latitude="-32.15" location="Clanwilliam" longLatPrecision="1212" longitude="18.75" municipality="Farm Zeekoeivlei" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<emphasis id="B933EA92FF9CB504E2E9B5B1FD525451" bold="true" box="[415,706,510,537]" pageId="7" pageNumber="550">
<collectingRegion id="4983F862FF9CB504E2E9B5B1FD2C5451" box="[415,700,510,537]" country="South Africa" name="Western Cape" pageId="7" pageNumber="550">Western Cape Province</collectingRegion>
.
</emphasis>
1♁, 
<specimenCount id="9D41FD09FF9CB504E18AB5B1FC8F5450" box="[764,799,511,537]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<collectingMunicipality id="6B9CACFAFF9CB504E051B5B1FC7E5450" box="[807,1006,511,537]" pageId="7" pageNumber="550">Farm Zeekoeivlei</collectingMunicipality>
[
<emphasis id="B933EA92FF9CB504E776B5B1FBA75451" box="[1024,1079,511,536]" italics="true" pageId="7" pageNumber="550">recte</emphasis>
Seekoevlei], near 
<location id="8E98605BFF9CB504E67DB5B1FA085450" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E67DB5B1FA085450" box="[1291,1432,511,537]" country="South Africa" latitude="-32.15" longLatPrecision="1212" longitude="18.75" municipality="Farm Zeekoeivlei" name="Clanwilliam" pageId="7" pageNumber="550" stateProvince="Western Cape">Clanwilliam</location>
, [
<geoCoordinate id="EE735047FF9CB504E3E8B66AFF6A5476" box="[158,250,548,575]" degrees="32" direction="south" minutes="09" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-32.15">32°09′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E272B66AFEF35476" box="[260,355,548,575]" degrees="18" direction="east" minutes="45" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="18.75">18°45′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E20FB66DFE645474" box="[377,500,547,573]" pageId="7" pageNumber="550" value="2003-11-27">
<collectingDate id="EFBDE9A8FF9CB504E20FB66DFE645474" box="[377,500,547,573]" pageId="7" pageNumber="550" value="2003-11-27">27.xi.2003</collectingDate>
</date>
, farm staff (
<collectionCode id="ED56AE45FF9CB504E1F0B66DFD4A5474" box="[646,730,547,573]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E186B66AFE0054CF" ID-GBIF-Occurrence="3062300307" collectionCode="MZLU" collectorName="P Brink &amp; G Rudebeck" country="South Africa" county="Saldanha Bay" latitude="-33.08347" location="South African Expedition" longLatPrecision="13" longitude="17.955278" municipality="Jutten Island" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<collectingCounty id="62994E0CFF9CB504E06BB66DFC2C5474" box="[797,956,547,573]" pageId="7" pageNumber="550">Saldanha Bay</collectingCounty>
, 
<collectingMunicipality id="6B9CACFAFF9CB504E0BDB66DFBCF5474" box="[971,1119,547,573]" pageId="7" pageNumber="550">Jutten Island</collectingMunicipality>
, [
<geoCoordinate id="EE735047FF9CB504E703B66AFB6F5476" box="[1141,1279,548,575]" degrees="33" direction="south" minutes="05" orientation="latitude" pageId="7" pageNumber="550" precision="1" seconds="0.5" value="-33.08347">33°05′0.5″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E67CB66AFA005476" box="[1290,1424,548,575]" degrees="17" direction="east" minutes="57" orientation="longitude" pageId="7" pageNumber="550" precision="15" seconds="19" value="17.955278">17°57′19″E</geoCoordinate>
], 25.i.[19]51, Swedish 
<location id="8E98605BFF9CB504E2F0B609FD365428" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E2F0B609FD365428" box="[390,678,583,609]" country="South Africa" county="Saldanha Bay" latitude="-33.08347" longLatPrecision="13" longitude="17.955278" municipality="Jutten Island" name="South African Expedition" pageId="7" pageNumber="550" stateProvince="Western Cape">South African Expedition</location>
, 1950–1951, 
<collectorName id="26B25356FF9CB504E04DB609FC035428" box="[827,915,583,609]" pageId="7" pageNumber="550">P Brink</collectorName>
, 
<collectorName id="26B25356FF9CB504E0EBB609FBB85428" box="[925,1064,583,609]" pageId="7" pageNumber="550">G Rudebeck</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E740B609FB1B5428" box="[1078,1163,583,609]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" lsid="urn:lsid:biocol.org:col:34994" name="Lund University" pageId="7" pageNumber="550">MZLU</collectionCode>
: MZLU-COL00007061, through photographs)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E2EAB622FD5B54E0" ID-GBIF-Occurrence="3062300341" collectingDate="1983-08-30" collectingDateMax="1983-11-01" collectingDateMin="1983-08-30" collectionCode="TMSA" collectorName="S Endrody-Younga &amp; Penrith" country="South Africa" latitude="-33.35" location="Town" longLatPrecision="1205" longitude="18.25" pageId="7" pageNumber="550" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape">
1♁, 
<specimenCount id="9D41FD09FF9CB504E2B9B625FE6254CC" box="[463,498,619,645]" pageId="7" pageNumber="550" type="female">4♀</specimenCount>
<collectingRegion id="4983F862FF9CB504E28AB625FDA654CC" box="[508,566,619,645]" country="South Africa" name="Western Cape" pageId="7" pageNumber="550">Cape</collectingRegion>
<location id="8E98605BFF9CB504E148B625FD1054CC" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E148B625FD1054CC" box="[574,640,619,645]" country="South Africa" latitude="-33.35" longLatPrecision="1205" longitude="18.25" name="Town" pageId="7" pageNumber="550" stateProvince="Western Cape">Town</location>
, 
<quantity id="4CBF9B65FF9CB504E1F8B624FD4554CC" box="[654,725,618,645]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="550" unit="km" value="65.0">65 km</quantity>
N, 
<geoCoordinate id="EE735047FF9CB504E075B622FCCC54CE" box="[771,860,620,647]" degrees="33" direction="south" minutes="21" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.35">33°21′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E013B622FC5254CE" box="[869,962,620,647]" degrees="18" direction="east" minutes="15" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="18.25">18°15′E</geoCoordinate>
, 
<collectingMethod id="52064E97FF9CB504E0A6B625FBC854CC" box="[976,1112,619,645]" pageId="7" pageNumber="550">pitfall traps</collectingMethod>
: 
<date id="FFF91040FF9CB504E712B625FB7E54CC" box="[1124,1262,619,645]" pageId="7" pageNumber="550" value="1983-08-30">
<collectingDate id="EFBDE9A8FF9CB504E712B625FB7E54CC" box="[1124,1262,619,645]" pageId="7" pageNumber="550" value="1983-08-30">30.viii.1983</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E788B625FAE454CC" box="[1278,1396,619,645]" pageId="7" pageNumber="550" value="1983-11-01">
<collectingDate id="EFBDE9A8FF9CB504E788B625FAE454CC" box="[1278,1396,619,645]" pageId="7" pageNumber="550" value="1983-11-01">01.xi.1983</collectingDate>
</date>
, E- Y:1999, 
<collectorName id="26B25356FF9CB504E380B6C1FE5E54E0" box="[246,462,655,681]" pageId="7" pageNumber="550">S Endrödy-Younga</collectorName>
, M-L 
<collectorName id="26B25356FF9CB504E166B6C1FDF154E0" box="[528,609,655,681]" pageId="7" pageNumber="550">Penrith</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E119B6C1FD5354E0" box="[623,707,655,681]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E1A2B6C1FCDF5487" ID-GBIF-Occurrence="3062300322" collectingDate="2009-09-18" collectionCode="UPSA" collectorName="WP Strumpher &amp; CH Scholtz" country="South Africa" latitude="-33.423336" location="Dassen Island" longLatPrecision="20" longitude="18.087223" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
20 unsexed: 
<location id="8E98605BFF9CB504E016B6C1FC6D54E0" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E016B6C1FC6D54E0" box="[864,1021,655,681]" country="South Africa" latitude="-33.423336" longLatPrecision="20" longitude="18.087223" name="Dassen Island" pageId="7" pageNumber="550" stateProvince="Western Cape">Dassen Island</location>
, 
<geoCoordinate id="EE735047FF9CB504E77EB6DEFB1854E2" box="[1032,1160,656,683]" degrees="33" direction="south" minutes="25" orientation="latitude" pageId="7" pageNumber="550" precision="15" seconds="24" value="-33.423336">33°25′24″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E7F8B6DEFA8354E2" box="[1166,1299,656,683]" degrees="18" direction="east" minutes="05" orientation="longitude" pageId="7" pageNumber="550" precision="15" seconds="14" value="18.087223">18°05′14″E</geoCoordinate>
, 
<date id="FFF91040FF9CB504E66BB6C1FA0754E0" box="[1309,1431,655,681]" pageId="7" pageNumber="550" value="2009-09-18">
<collectingDate id="EFBDE9A8FF9CB504E66BB6C1FA0754E0" box="[1309,1431,655,681]" pageId="7" pageNumber="550" value="2009-09-18">18.ix.2009</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E3E1B6FDFEAD5484" box="[151,317,691,717]" pageId="7" pageNumber="550">WP Strümpher</collectorName>
, 
<collectorName id="26B25356FF9CB504E231B6FDFE575484" box="[327,455,691,717]" pageId="7" pageNumber="550">CH Scholtz</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E2A2B6FDFDB05484" box="[468,544,691,717]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="7" pageNumber="550">UPSA</collectionCode>
, preserved in 99% ethanol)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E02EB6FAFA9B5484" ID-GBIF-Occurrence="3062300324" box="[856,1291,691,719]" collectingDate="2009-09-18" collectionCode="SANC" collectorName="WP Strumpher &amp; CH Scholtz" country="South Africa" latitude="-33.423336" location="Dassen Island" longLatPrecision="20" longitude="18.087223" pageId="7" pageNumber="550" specimenCount="3" specimenCount-female="3" stateProvince="Western Cape">
6♁, 
<specimenCount id="9D41FD09FF9CB504E0F1B6FDFC3A5484" box="[903,938,691,717]" pageId="7" pageNumber="550" type="female">3♀</specimenCount>
<emphasis id="B933EA92FF9CB504E0D9B6FDFC755484" box="[943,997,691,717]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but dry-mounted (
<collectionCode id="ED56AE45FF9CB504E7C4B6FDFA935484" box="[1202,1283,691,717]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="7" pageNumber="550" type="Museum">SANC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E662B6FAFD0054B8" ID-GBIF-Occurrence="3062300347" collectingDate="2009-09-18" collectionCode="SAMC" collectorName="WP Strumpher &amp; CH Scholtz" country="South Africa" latitude="-33.423336" location="Dassen Island" longLatPrecision="20" longitude="18.087223" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E64BB6FDFAE35484" box="[1341,1395,691,717]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but dry-mounted (
<collectionCode id="ED56AE45FF9CB504E243B699FE1A54B8" box="[309,394,727,753]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="550" type="Museum">SAMC</collectionCode>
: SAM-COL-A082509)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E1EFB696FAA154B8" ID-GBIF-Occurrence="3062300346" box="[665,1329,727,755]" collectingDate="1974-05" collectionCode="TMSA" collectorName="Honiball" country="South Africa" latitude="-32.583332" location="Citrusdal" longLatPrecision="1209" longitude="19.016666" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
2♁ 
<location id="8E98605BFF9CB504E1B4B699FCBC54B8" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E1B4B699FCBC54B8" box="[706,812,727,753]" country="South Africa" latitude="-32.583332" longLatPrecision="1209" longitude="19.016666" name="Citrusdal" pageId="7" pageNumber="550" stateProvince="Western Cape">Citrusdal</location>
, [
<geoCoordinate id="EE735047FF9CB504E04AB696FC0854BA" box="[828,920,728,755]" degrees="32" direction="south" minutes="35" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-32.583332">32°35′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E0EBB696FC6C54BA" box="[925,1020,728,755]" degrees="19" direction="east" minutes="01" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="19.016666">19°01′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E77BB699FBCC54B8" box="[1037,1116,727,753]" pageId="7" pageNumber="550" value="1974-05">
<collectingDate id="EFBDE9A8FF9CB504E77BB699FBCC54B8" box="[1037,1116,727,753]" pageId="7" pageNumber="550" value="1974-05">v.1974</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E713B699FB5754B8" box="[1125,1223,727,753]" pageId="7" pageNumber="550">Honiball</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E7A3B699FAB954B8" box="[1237,1321,727,753]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E64CB699FDCD555C" ID-GBIF-Occurrence="3062300302" collectionCode="TMSA" country="South Africa" latitude="-33.916668" location="Town" longLatPrecision="1202" longitude="18.416666" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="9D41FD09FF9CB504E64CB699FACD54B8" box="[1338,1373,727,753]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<collectingRegion id="4983F862FF9CB504E615B699FA0D54B8" box="[1379,1437,727,753]" country="South Africa" name="Western Cape" pageId="7" pageNumber="550">Cape</collectingRegion>
<location id="8E98605BFF9CB504E3E1B6B5FF49555C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E3E1B6B5FF49555C" box="[151,217,763,789]" country="South Africa" latitude="-33.916668" longLatPrecision="1202" longitude="18.416666" name="Town" pageId="7" pageNumber="550" stateProvince="Western Cape">Town</location>
, [
<geoCoordinate id="EE735047FF9CB504E39CB6B2FED5555E" box="[234,325,764,791]" degrees="33" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.916668">33°55′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E23DB6B2FE3A555E" box="[331,426,764,791]" degrees="18" direction="east" minutes="25" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="18.416666">18°25′E</geoCoordinate>
], 1913 (
<collectionCode id="ED56AE45FF9CB504E177B6B5FDC5555C" box="[513,597,763,789]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E110B6B2FEFF5570" ID-GBIF-Occurrence="3062300312" collectingDate="1958-06-16" collectionCode="TMSA" collectorName="JA Nagle" country="South Africa" latitude="-33.906944" location="Green Point" longLatPrecision="19" longitude="18.40639" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ CPT [
<collectingRegion id="4983F862FF9CB504E1BBB6B5FC99555C" box="[717,777,763,789]" country="South Africa" name="Western Cape" pageId="7" pageNumber="550">Cape</collectingRegion>
Town], 
<location id="8E98605BFF9CB504E012B6B5FC7C555C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E012B6B5FC7C555C" box="[868,1004,763,789]" country="South Africa" latitude="-33.906944" longLatPrecision="19" longitude="18.40639" name="Green Point" pageId="7" pageNumber="550" stateProvince="Western Cape">Green Point</location>
, [
<geoCoordinate id="EE735047FF9CB504E08BB6B2FB10555E" box="[1021,1152,764,791]" degrees="33" direction="south" minutes="54" orientation="latitude" pageId="7" pageNumber="550" precision="15" seconds="25" value="-33.906944">33°54′25″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E7F3B6B2FA9C555E" box="[1157,1292,764,791]" degrees="18" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="550" precision="15" seconds="23" value="18.40639">18°24′23″E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E66BB6B4FA08555C" box="[1309,1432,762,789]" pageId="7" pageNumber="550" value="1958-06-16">
<collectingDate id="EFBDE9A8FF9CB504E66BB6B4FA08555C" box="[1309,1432,762,789]" pageId="7" pageNumber="550" value="1958-06-16">16.vi.1958</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E3E1B751FE925570" box="[151,258,799,825]" pageId="7" pageNumber="550">JA Nagle</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E265B751FEF75570" box="[275,359,799,825]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E20BB76EFDD95514" ID-GBIF-Occurrence="3062300313" collectingDate="1978-11-08" collectingDateMax="1979-01-17" collectingDateMin="1978-11-08" collectionCode="TMSA" collectorName="S Endrody-Younga" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
2♁ 
<collectingMunicipality id="6B9CACFAFF9CB504E2DFB751FDA55570" box="[425,565,799,825]" pageId="7" pageNumber="550">Cape-Karoo</collectingMunicipality>
, 
<location id="8E98605BFF9CB504E135B751FC815570" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E135B751FC815570" box="[579,785,799,825]" country="South Africa" latitude="-33.166668" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" name="Farm Zwartskraal" pageId="7" pageNumber="550" stateProvince="Western Cape">Farm Zwartskraal</location>
, 
<geoCoordinate id="EE735047FF9CB504E069B76EFCE85572" box="[799,888,800,827]" degrees="33" direction="south" minutes="10" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.166668">33°10′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E0F7B76EFC4E5572" box="[897,990,800,827]" degrees="22" direction="east" minutes="32" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="22.533333">22°32′E</geoCoordinate>
, 
<collectingMethod id="52064E97FF9CB504E09BB751FBE45570" box="[1005,1140,799,825]" pageId="7" pageNumber="550">pitfall traps</collectingMethod>
: 
<date id="FFF91040FF9CB504E7F7B751FB6B5570" box="[1153,1275,799,825]" pageId="7" pageNumber="550" value="1978-11-08">
<collectingDate id="EFBDE9A8FF9CB504E7F7B751FB6B5570" box="[1153,1275,799,825]" pageId="7" pageNumber="550" value="1978-11-08">08.xi.1978</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E67AB751FAE35570" box="[1292,1395,799,825]" pageId="7" pageNumber="550" value="1979-01-17">
<collectingDate id="EFBDE9A8FF9CB504E67AB751FAE35570" box="[1292,1395,799,825]" pageId="7" pageNumber="550" value="1979-01-17">17.i.1979</collectingDate>
</date>
, E- Y:1540, 
<collectorName id="26B25356FF9CB504E38AB70DFE485514" box="[252,472,835,861]" pageId="7" pageNumber="550">S Endrödy-Younga</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E29BB70DFDD15514" box="[493,577,835,861]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E12CB70AFA085514" ID-GBIF-Occurrence="3062300315" box="[602,1432,834,863]" collectingDate="1979-02-01" collectingDateMax="1979-03-02" collectingDateMin="1979-02-01" collectionCode="TMSA" collectorName="R Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E1FFB70DFD2F5514" box="[649,703,835,861]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E18BB70DFCFD5514" box="[765,877,835,861]" pageId="7" pageNumber="550" value="1979-02-01">
<collectingDate id="EFBDE9A8FF9CB504E18BB70DFCFD5514" box="[765,877,835,861]" pageId="7" pageNumber="550" value="1979-02-01">01.ii.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E00BB70DFC685514" box="[893,1016,835,861]" pageId="7" pageNumber="550" value="1979-03-02">
<collectingDate id="EFBDE9A8FF9CB504E00BB70DFC685514" box="[893,1016,835,861]" pageId="7" pageNumber="550" value="1979-03-02">02.iii.1979</collectingDate>
</date>
, E-Y:1545, 
<collectorName id="26B25356FF9CB504E7FFB70DFAB85514" box="[1161,1320,835,861]" pageId="7" pageNumber="550">R Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E64AB70DFA005514" box="[1340,1424,835,861]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E3E1B726FC5655C8" ID-GBIF-Occurrence="3062300327" box="[151,966,870,899]" collectingDate="1979-03-02" collectingDateMax="1979-03-31" collectingDateMin="1979-03-02" collectionCode="TMSA" collectorName="R Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E3B5B729FF6955C8" box="[195,249,871,897]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E246B729FE3C55C8" box="[304,428,871,897]" pageId="7" pageNumber="550" value="1979-03-02">
<collectingDate id="EFBDE9A8FF9CB504E246B729FE3C55C8" box="[304,428,871,897]" pageId="7" pageNumber="550" value="1979-03-02">02.iii.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E2CDB729FDA455C8" box="[443,564,871,897]" pageId="7" pageNumber="550" value="1979-03-31">
<collectingDate id="EFBDE9A8FF9CB504E2CDB729FDA455C8" box="[443,564,871,897]" pageId="7" pageNumber="550" value="1979-03-31">31.iii.1979</collectingDate>
</date>
, E-Y:1552, 
<collectorName id="26B25356FF9CB504E1CBB729FCC955C8" box="[701,857,871,897]" pageId="7" pageNumber="550">R Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E01CB729FC2E55C8" box="[874,958,871,897]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E0A5B726FE6E55EC" ID-GBIF-Occurrence="3062300319" collectingDate="1979-07-23" collectingDateMax="1979-09-05" collectingDateMin="1979-07-23" collectionCode="TMSA" collectorName="R Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E776B729FBA655C8" box="[1024,1078,871,897]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E71AB728FB7C55C8" box="[1132,1260,870,897]" pageId="7" pageNumber="550" value="1979-07-23">
<collectingDate id="EFBDE9A8FF9CB504E71AB728FB7C55C8" box="[1132,1260,870,897]" pageId="7" pageNumber="550" value="1979-07-23">23.vii.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E78AB728FAE455C8" box="[1276,1396,870,897]" pageId="7" pageNumber="550" value="1979-09-05">
<collectingDate id="EFBDE9A8FF9CB504E78AB728FAE455C8" box="[1276,1396,870,897]" pageId="7" pageNumber="550" value="1979-09-05">05.ix.1979</collectingDate>
</date>
, E- Y:1637, 
<collectorName id="26B25356FF9CB504E38EB7C5FE0355EC" box="[248,403,907,933]" pageId="7" pageNumber="550">R Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E2D4B7C5FE6655EC" box="[418,502,907,933]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E17DB7C2FABE55EC" ID-GBIF-Occurrence="3062300326" box="[523,1326,906,935]" collectingDate="1979-09-05" collectingDateMax="1979-10-25" collectingDateMin="1979-09-05" collectionCode="TMSA" collectorName="R. Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E140B7C5FDFC55EC" box="[566,620,907,933]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E1D7B7C4FC8955EC" box="[673,793,906,933]" pageId="7" pageNumber="550" value="1979-09-05">
<collectingDate id="EFBDE9A8FF9CB504E1D7B7C4FC8955EC" box="[673,793,906,933]" pageId="7" pageNumber="550" value="1979-09-05">05.ix.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E05FB7C4FC0955EC" box="[809,921,906,933]" pageId="7" pageNumber="550" value="1979-10-25">
<collectingDate id="EFBDE9A8FF9CB504E05FB7C4FC0955EC" box="[809,921,906,933]" pageId="7" pageNumber="550" value="1979-10-25">25.x.1979</collectingDate>
</date>
, E-Y:1639, 
<collectorName id="26B25356FF9CB504E757B7C5FB5255EC" box="[1057,1218,907,933]" pageId="7" pageNumber="550">R. Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E7A4B7C5FAB655EC" box="[1234,1318,907,933]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E64DB7C2FCF55580" ID-GBIF-Occurrence="3062300303" collectingDate="1979-10-25" collectingDateMax="1979-12-09" collectingDateMin="1979-10-25" collectionCode="TMSA" collectorName="R. Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E610B7C5FA0C55EC" box="[1382,1436,907,933]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E3B0B7E0FEA55580" box="[198,309,942,969]" pageId="7" pageNumber="550" value="1979-10-25">
<collectingDate id="EFBDE9A8FF9CB504E3B0B7E0FEA55580" box="[198,309,942,969]" pageId="7" pageNumber="550" value="1979-10-25">25.x.1979</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E233B7E0FE565580" box="[325,454,942,969]" pageId="7" pageNumber="550" value="1979-12-09">
<collectingDate id="EFBDE9A8FF9CB504E233B7E0FE565580" box="[325,454,942,969]" pageId="7" pageNumber="550" value="1979-12-09">09.xii.1979</collectingDate>
</date>
, E-Y:1672, 
<collectorName id="26B25356FF9CB504E124B7E1FD675580" box="[594,759,943,969]" pageId="7" pageNumber="550">R. Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E07FB7E1FCCD5580" box="[777,861,943,969]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E002B7FEFE1655A4" ID-GBIF-Occurrence="3062300337" collectingDate="1980-01-01" collectingDateMax="1980-02-18" collectingDateMin="1980-01-01" collectionCode="TMSA" collectorName="R Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E0D7B7E1FC475580" box="[929,983,943,969]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E767B7E1FBEB5580" box="[1041,1147,943,969]" pageId="7" pageNumber="550" value="1980-01-01">
<collectingDate id="EFBDE9A8FF9CB504E767B7E1FBEB5580" box="[1041,1147,943,969]" pageId="7" pageNumber="550" value="1980-01-01">01.i.1980</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E7FDB7E1FB6D5580" box="[1163,1277,943,969]" pageId="7" pageNumber="550" value="1980-02-18">
<collectingDate id="EFBDE9A8FF9CB504E7FDB7E1FB6D5580" box="[1163,1277,943,969]" pageId="7" pageNumber="550" value="1980-02-18">18.ii.1980</collectingDate>
</date>
, E-Y:1699, 
<collectorName id="26B25356FF9CB504E6FCB7E1FE8755A4" pageId="7" pageNumber="550">R Oosthuizen</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E25CB79DFEEE55A4" box="[298,382,979,1005]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E2E3B79AFB8E55A4" ID-GBIF-Occurrence="3062300331" box="[405,1054,979,1007]" collectingDate="1980-09-22" collectingDateMax="1980-11-08" collectingDateMin="1980-09-22" collectionCode="TMSA" collectorName="R Oosthuizen" country="South Africa" latitude="-33.166668" location="Farm Zwartskraal" longLatPrecision="1206" longitude="22.533333" municipality="Cape-Karoo" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<emphasis id="B933EA92FF9CB504E2B5B79DFE6955A4" box="[451,505,979,1005]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
but 
<date id="FFF91040FF9CB504E145B79DFD3C55A4" box="[563,684,979,1005]" pageId="7" pageNumber="550" value="1980-09-22">
<collectingDate id="EFBDE9A8FF9CB504E145B79DFD3C55A4" box="[563,684,979,1005]" pageId="7" pageNumber="550" value="1980-09-22">22.ix.1980</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E1CAB79DFCA455A4" box="[700,820,979,1005]" pageId="7" pageNumber="550" value="1980-11-08">
<collectingDate id="EFBDE9A8FF9CB504E1CAB79DFCA455A4" box="[700,820,979,1005]" pageId="7" pageNumber="550" value="1980-11-08">08.xi.1980</collectingDate>
</date>
, E-Y:1717 (
<collectionCode id="ED56AE45FF9CB504E0B4B79DFB8655A4" box="[962,1046,979,1005]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E75BB79DFC9D5258" ID-GBIF-Occurrence="3062300345" collectionCode="SAMC" collectorName="Touws R &amp; Purcell" country="South Africa" latitude="-33.337498" longLatPrecision="773" longitude="20.033333" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="9D41FD09FF9CB504E75BB79DFBDF55A4" box="[1069,1103,979,1005]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<collectorName id="26B25356FF9CB504E72CB79DFB5455A4" box="[1114,1220,978,1005]" pageId="7" pageNumber="550">Touws R</collectorName>
[iver], [
<geoCoordinate id="EE735047FF9CB504E66FB79AFA0C55A6" box="[1305,1436,980,1007]" degrees="33" direction="south" minutes="20" orientation="latitude" pageId="7" pageNumber="550" precision="15" seconds="15" value="-33.337498">33°20′15″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E3E1B7B6FF67525A" box="[151,247,1016,1043]" degrees="20" direction="east" minutes="02" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="20.033333">20°02′E</geoCoordinate>
], 1896, 
<collectorName id="26B25356FF9CB504E227B7B9FE305258" box="[337,416,1015,1041]" pageId="7" pageNumber="550">Purcell</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E2C7B7B9FD965258" box="[433,518,1015,1041]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="550" type="Museum">SAMC</collectionCode>
: SAM-COL-A082507)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E06FB7B6FC69527C" ID-GBIF-Occurrence="3062300321" collectingDate="1968-12" collectionCode="TMSA" collectorName="RH Jones &amp; JH Potgieter" country="South Africa" latitude="-33.35" location="Prince Albert district" longLatPrecision="1205" longitude="22.05" municipality="Zwartberg Pass" pageId="7" pageNumber="550" specimenCount="2" specimenCount-female="2" stateProvince="Western Cape">
1♁, 
<specimenCount id="9D41FD09FF9CB504E03DB7B9FCFE5258" box="[843,878,1015,1041]" pageId="7" pageNumber="550" type="female">2♀</specimenCount>
<collectingMunicipality id="6B9CACFAFF9CB504E000B7B9FBB55258" box="[886,1061,1015,1041]" pageId="7" pageNumber="550">Zwartberg Pass</collectingMunicipality>
[
<emphasis id="B933EA92FF9CB504E740B7B9FBFD5259" box="[1078,1133,1015,1040]" italics="true" pageId="7" pageNumber="550">recte</emphasis>
Swartberg Pass], [
<geoCoordinate id="EE735047FF9CB504E637B7B6FA0C525A" box="[1345,1436,1016,1043]" degrees="33" direction="south" minutes="21" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.35">33°21′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E3E1B052FF66527E" box="[151,246,1052,1079]" degrees="22" direction="east" minutes="03" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="22.05">22°03′E</geoCoordinate>
], 
<location id="8E98605BFF9CB504E27AB055FE6F527C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E27AB055FE6F527C" box="[268,511,1051,1077]" country="South Africa" latitude="-33.35" longLatPrecision="1205" longitude="22.05" municipality="Zwartberg Pass" name="Prince Albert district" pageId="7" pageNumber="550" stateProvince="Western Cape">Prince Albert district</location>
, 
<date id="FFF91040FF9CB504E178B055FDFC527C" box="[526,620,1051,1077]" pageId="7" pageNumber="550" value="1968-12">
<collectingDate id="EFBDE9A8FF9CB504E178B055FDFC527C" box="[526,620,1051,1077]" pageId="7" pageNumber="550" value="1968-12">xii.1968</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E10DB055FD7E527C" box="[635,750,1051,1077]" pageId="7" pageNumber="550">RH Jones</collectorName>
, 
<collectorName id="26B25356FF9CB504E18AB055FC1B527C" box="[764,907,1051,1077]" pageId="7" pageNumber="550">JH Potgieter</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E0EBB055FC61527C" box="[925,1009,1051,1077]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E77EB052FD9B5210" ID-GBIF-Occurrence="3062300343" collectingDate="1968-12" collectionCode="SAMC" collectorName="RH Jones &amp; JH Potgieter" country="South Africa" latitude="-33.35" location="Prince Albert district" longLatPrecision="1205" longitude="22.05" municipality="Zwartberg Pass" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
1♁, 
<specimenCount id="9D41FD09FF9CB504E74AB055FBCF527C" box="[1084,1119,1051,1077]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<emphasis id="B933EA92FF9CB504E71FB055FB0F527C" box="[1129,1183,1051,1077]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
(
<collectionCode id="ED56AE45FF9CB504E7C4B055FA97527C" box="[1202,1287,1051,1077]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="550" type="Museum">SAMC</collectionCode>
: SAM-COL- A082508, SAM-COL-A082510)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E16EB071FB705234" ID-GBIF-Occurrence="3062300304" collectingDate="1976-12-16" collectingDateMax="1977-03-02" collectingDateMin="1976-12-16" collectionCode="TMSA" collectorName="S Endrody-Younga" country="South Africa" elevation="2000" latitude="-33.416668" location="Blesberg East" longLatPrecision="1205" longitude="22.683332" municipality="Swartberge" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="9D41FD09FF9CB504E16EB071FDAB5210" box="[536,571,1087,1113]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<collectingMunicipality id="6B9CACFAFF9CB504E132B071FD535210" box="[580,707,1087,1113]" pageId="7" pageNumber="550">Swartberge</collectingMunicipality>
[Swartberg Mountains], 
<location id="8E98605BFF9CB504E097B071FB105210" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E097B071FB105210" box="[993,1152,1087,1113]" country="South Africa" latitude="-33.416668" longLatPrecision="1205" longitude="22.683332" municipality="Swartberge" name="Blesberg East" pageId="7" pageNumber="550" stateProvince="Western Cape">Blesberg East</location>
, 
<quantity id="4CBF9B65FF9CB504E7FBB071FB7D5210" box="[1165,1261,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="550" unit="m" value="2000.0">
<elevation id="006AD1B3FF9CB504E7FBB071FB7D5210" box="[1165,1261,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="550" unit="m" value="2000.0">2,000 m</elevation>
</quantity>
, 23°25′S [
<emphasis id="B933EA92FF9CB504E613B071FA0C5211" box="[1381,1436,1087,1112]" italics="true" pageId="7" pageNumber="550">recte</emphasis>
<geoCoordinate id="EE735047FF9CB504E3E1B02AFF625236" box="[151,242,1124,1151]" degrees="33" direction="south" minutes="25" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.416668">33°25′S</geoCoordinate>
] 
<geoCoordinate id="EE735047FF9CB504E275B02AFEF15236" box="[259,353,1124,1151]" degrees="22" direction="east" minutes="41" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="22.683332">22°41′E</geoCoordinate>
, 
<collectingMethod id="52064E97FF9CB504E219B02DFE675234" box="[367,503,1123,1149]" pageId="7" pageNumber="550">pitfall traps</collectingMethod>
: 
<date id="FFF91040FF9CB504E172B02DFD135234" box="[516,643,1123,1149]" pageId="7" pageNumber="550" value="1976-12-16">
<collectingDate id="EFBDE9A8FF9CB504E172B02DFD135234" box="[516,643,1123,1149]" pageId="7" pageNumber="550" value="1976-12-16">16.xii.1976</collectingDate>
</date>
– 
<date id="FFF91040FF9CB504E1E5B02DFC9D5234" box="[659,781,1123,1149]" pageId="7" pageNumber="550" value="1977-03-02">
<collectingDate id="EFBDE9A8FF9CB504E1E5B02DFC9D5234" box="[659,781,1123,1149]" pageId="7" pageNumber="550" value="1977-03-02">02.iii.1977</collectingDate>
</date>
, E-Y:1532, 
<collectorName id="26B25356FF9CB504E0EFB02DFBE25234" box="[921,1138,1123,1149]" pageId="7" pageNumber="550">S Endrödy-Younga</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E7F2B02DFB485234" box="[1156,1240,1123,1149]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E79BB02DFC1952E8" ID-GBIF-Occurrence="3062300317" collectingDate="1994-02" collectingDateMax="1994-03-31" collectingDateMin="1994-02" collectionCode="TMSA" collectorName="T Berry" country="South Africa" latitude="-33.716667" location="Gamkaberg Nature Reserve" longLatPrecision="1203" longitude="21.9" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount id="9D41FD09FF9CB504E79BB02DFA805234" box="[1261,1296,1123,1149]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<location id="8E98605BFF9CB504E66CB02DFEDB52E8" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E66CB02DFEDB52E8" country="South Africa" latitude="-33.716667" longLatPrecision="1203" longitude="21.9" name="Gamkaberg Nature Reserve" pageId="7" pageNumber="550" stateProvince="Western Cape">Gamkaberg Nature Reserve</location>
, [
<geoCoordinate id="EE735047FF9CB504E216B0C6FE2C52EA" box="[352,444,1160,1187]" degrees="33" direction="south" minutes="43" orientation="latitude" pageId="7" pageNumber="550" precision="925" value="-33.716667">33°43′S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E2B0B0C6FDB552EA" box="[454,549,1160,1187]" degrees="21" direction="east" minutes="54" orientation="longitude" pageId="7" pageNumber="550" precision="925" value="21.9">21°54′E</geoCoordinate>
], 
<date id="FFF91040FF9CB504E14DB0C9FD2152E8" box="[571,689,1159,1185]" pageId="7" pageNumber="550" value="1994-02" valueMax="1994-03-31" valueMin="1994-02-00">
<collectingDate id="EFBDE9A8FF9CB504E14DB0C9FD2152E8" box="[571,689,1159,1185]" pageId="7" pageNumber="550" value="1994-02" valueMax="1994-03-31" valueMin="1994-02-00">ii–iii.1994</collectingDate>
</date>
, 
<collectorName id="26B25356FF9CB504E1B6B0C9FC8B52E8" box="[704,795,1159,1185]" pageId="7" pageNumber="550">T Berry</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E05BB0C9FC1152E8" box="[813,897,1159,1185]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2F3CDDFF9CB504E0E1B0C6FE4F528C" ID-GBIF-Occurrence="3062300333" collectionCode="TMSA" collectorName="RJ Power" country="South Africa" latitude="-34.045834" location="Riversdale" longLatPrecision="19" longitude="21.2625" pageId="7" pageNumber="550" specimenCount="1" stateProvince="Western Cape">
1♁ 
<location id="8E98605BFF9CB504E0B2B0C9FBAF52E8" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF9CB504E0B2B0C9FBAF52E8" box="[964,1087,1159,1185]" country="South Africa" latitude="-34.045834" longLatPrecision="19" longitude="21.2625" name="Riversdale" pageId="7" pageNumber="550" stateProvince="Western Cape">Riversdale</location>
, [
<geoCoordinate id="EE735047FF9CB504E722B0C6FB4752EA" box="[1108,1239,1160,1187]" degrees="34" direction="south" minutes="02" orientation="latitude" pageId="7" pageNumber="550" precision="15" seconds="45" value="-34.045834">34°02′45″S</geoCoordinate>
<geoCoordinate id="EE735047FF9CB504E794B0C6FAF852EA" box="[1250,1384,1160,1187]" degrees="21" direction="east" minutes="15" orientation="longitude" pageId="7" pageNumber="550" precision="15" seconds="45" value="21.2625">21°15′45″E</geoCoordinate>
], 
<collectorName id="26B25356FF9CB504E609B0C9FF4F528C" pageId="7" pageNumber="550">RJ Power</collectorName>
(
<collectionCode id="ED56AE45FF9CB504E39BB0E5FEAE528C" box="[237,318,1195,1221]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="550">TMSA</collectionCode>
) (
<figureCitation id="137C2A05FF9CB504E223B0E5FE46528C" box="[341,470,1195,1221]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">Figs 11–14</figureCitation>
)
</materialsCitation>
; 
<materialsCitation id="3B2F3CDDFF9CB504E29DB0E2FBCD528C" ID-GBIF-Occurrence="3062300323" box="[491,1117,1194,1223]" collectionCode="SAMC" collectorName="RJ Power" country="South Africa" latitude="-34.045834" location="Riversdale" longLatPrecision="19" longitude="21.2625" pageId="7" pageNumber="550" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
2♁, 
<specimenCount id="9D41FD09FF9CB504E16AB0E5FDAF528C" box="[540,575,1195,1221]" pageId="7" pageNumber="550" type="female">1♀</specimenCount>
<emphasis id="B933EA92FF9CB504E130B0E5FDEC528C" box="[582,636,1195,1221]" italics="true" pageId="7" pageNumber="550">idem</emphasis>
(
<collectionCode id="ED56AE45FF9CB504E1FDB0E5FD70528C" box="[651,736,1195,1221]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="7" pageNumber="550" type="Museum">SAMC</collectionCode>
: SAM-COL-A082503–A082505)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF9CB509E3B1B080FB045026" lastPageId="10" lastPageNumber="553" pageId="7" pageNumber="550" type="description">
<paragraph id="8BF83680FF9CB504E3B1B080FC6752A0" blockId="7.[151,1437,150,1977]" box="[199,1015,1230,1257]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B080FEE552A1" bold="true" box="[199,373,1230,1256]" pageId="7" pageNumber="550">Redescription.</emphasis>
<emphasis id="B933EA92FF9CB504E20AB081FE3952A0" box="[380,425,1231,1257]" italics="true" pageId="7" pageNumber="550">Size</emphasis>
: Length: 
<quantity id="4CBF9B65FF9CB504E160B081FD0252A0" box="[534,658,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="7" pageNumber="550" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
. Width across humeri: 
<quantity id="4CBF9B65FF9CB504E0E5B081FC6352A0" box="[915,1011,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="7" pageNumber="550" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B0BDFB2E5344" blockId="7.[151,1437,150,1977]" box="[199,1214,1267,1294]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B0BDFE865344" box="[199,278,1267,1293]" italics="true" pageId="7" pageNumber="550">Colour</emphasis>
(
<figureCitation id="137C2A05FF9CB504E253B0BDFEFD5344" box="[293,365,1267,1293]" captionStart="FIGURES 8–10" captionStartId="8.[151,264,1932,1956]" captionTargetBox="[182,1402,193,1906]" captionTargetId="figure-31@8.[167,1419,181,1908]" captionTargetPageId="8" captionText="FIGURES 8–10. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." figureDoi="http://doi.org/10.5281/zenodo.4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="7" pageNumber="550">Figs 8</figureCitation>
, 
<figureCitation id="137C2A05FF9CB504E20CB0BDFE545344" box="[378,452,1267,1293]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">11–12</figureCitation>
): Matt black. Body setae yellow-brown (amber) to light rust-brown.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B159FA92531C" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B159FE945378" box="[199,260,1303,1329]" italics="true" pageId="7" pageNumber="550">Head</emphasis>
(
<figureCitation id="137C2A05FF9CB504E260B159FE0B5378" box="[278,411,1303,1329]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">Figs 11–12</figureCitation>
): As in 
<taxonomicName id="4C474D03FF9CB504E28EB159FD225379" box="[504,690,1303,1329]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E28EB159FD225379" box="[504,690,1303,1329]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
, except surface coarsely sculpted and densely punctate, surfaces between punctures smooth, dull matt; antennal scape with long yellowish-brown to rust-brown setae.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B111FD7F53D4" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B111FEA65331" box="[199,310,1375,1400]" italics="true" pageId="7" pageNumber="550">Pronotum</emphasis>
(
<figureCitation id="137C2A05FF9CB504E23EB111FE5C5330" box="[328,460,1375,1401]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">Figs 11–12</figureCitation>
, 
<figureCitation id="137C2A05FF9CB504E2ACB111FE6A5333" box="[474,506,1375,1402]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1442,1466]" captionTargetBox="[171,1416,181,1419]" captionTargetId="figure-31@10.[170,1416,181,1419]" captionTargetPageId="10" captionText="FIGURES 15–16. Pronotal disc, dorsal view. 15, Phoberus cyrtus (Haaf, 1953); 16, Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612465" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="7" pageNumber="550">16</figureCitation>
): As in 
<taxonomicName id="4C474D03FF9CB504E12FB111FC835331" box="[601,787,1375,1401]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E12FB111FC835331" box="[601,787,1375,1401]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
, except surface coarsely punctate; punctures prominent, round; surfaces between punctures smooth, dull matt.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B1E9FEA353AC" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<taxonomicName id="4C474D03FF9CB504E3B1B1E9FEA55388" box="[199,309,1447,1473]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="CoL" kingdom="Fungi" order="Corynexochida" pageId="7" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="B933EA92FF9CB504E3B1B1E9FEA55388" box="[199,309,1447,1473]" italics="true" pageId="7" pageNumber="550">Scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="137C2A05FF9CB504E230B1E9FE595388" box="[326,457,1447,1473]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">Figs 11–12</figureCitation>
): Oval, rounded; surface smooth; lateral margins flat; mediobasally depressed, depression punctate.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B1A1FE015138" blockId="7.[151,1437,150,1977]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B1A1FE9D5040" box="[199,269,1519,1545]" italics="true" pageId="7" pageNumber="550">Elytra</emphasis>
(
<figureCitation id="137C2A05FF9CB504E268B1A1FEF75040" box="[286,359,1519,1545]" captionStart="FIGURES 8–10" captionStartId="8.[151,264,1932,1956]" captionTargetBox="[182,1402,193,1906]" captionTargetId="figure-31@8.[167,1419,181,1908]" captionTargetPageId="8" captionText="FIGURES 8–10. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." figureDoi="http://doi.org/10.5281/zenodo.4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="7" pageNumber="550">Figs 8</figureCitation>
, 
<figureCitation id="137C2A05FF9CB504E200B1A1FE505040" box="[374,448,1519,1545]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">11–12</figureCitation>
): Relative dimensions, margins and profiles as in 
<taxonomicName id="4C474D03FF9CB504E08DB1A1FB235041" box="[1019,1203,1519,1545]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E08DB1A1FB235041" box="[1019,1203,1519,1545]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
. Humeral calli as in 
<taxonomicName id="4C474D03FF9CB504E3E1B25DFEDE5065" box="[151,334,1555,1581]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E3E1B25DFEDE5065" box="[151,334,1555,1581]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
. Sutural margin raised, surface smooth, with distinct round to oval tubercles regularly distributed along margin, roughly of similar size for 0.75 elytral length, thereafter decreasing in size until barely discernible; these tubercles as in 
<taxonomicName id="4C474D03FF9CB504E2F2B215FDAC503D" box="[388,572,1627,1653]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E2F2B215FDAC503D" box="[388,572,1627,1653]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
. Elytral costae distinct; even-numbered costae with prominently raised oval to elongate tubercles, roughly of equal size and height, evenly spaced along elytral length; these tubercles smooth, dull, each with a posterior setal tuft. Apical callosity as in 
<taxonomicName id="4C474D03FF9CB504E040B2EDFC7F50F5" box="[822,1007,1699,1725]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E040B2EDFC7F50F5" box="[822,1007,1699,1725]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
. Elytral costae 2 and 4 with 8–12 tubercles for 0.75 elytral length (up to apical callosity); odd-numbered costae marked by small round to oval, dull shiny tubercles like in 
<taxonomicName id="4C474D03FF9CB504E2E7B2A5FDD6514D" box="[401,582,1771,1797]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E2E7B2A5FDD6514D" box="[401,582,1771,1797]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
, but these tubercles closely spaced. Costae 1–8 separated from intercostae as well-defined elevated bands. Intercostae with undulating depressions separated by transverse ridges, the margins of intercostae 1–7 marked by distinct, narrow longitudinal ridges, thereafter intercostae with carinae vestigial, barely discernible or lacking.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B335FD3051DC" blockId="7.[151,1437,150,1977]" box="[199,672,1915,1941]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B335FF6C51DD" box="[199,252,1915,1940]" italics="true" pageId="7" pageNumber="550">Legs</emphasis>
(
<figureCitation id="137C2A05FF9CB504E27DB335FE1E51DC" box="[267,398,1915,1941]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="7" pageNumber="550">Figs 11–12</figureCitation>
): As in 
<taxonomicName id="4C474D03FF9CB504E294B335FD0851DD" box="[482,664,1915,1941]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF9CB504E294B335FD0851DD" box="[482,664,1915,1941]" italics="true" pageId="7" pageNumber="550">Phoberus cyrtus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF83680FF9CB504E3B1B3D1FDBA51F0" blockId="7.[151,1437,150,1977]" box="[199,554,1951,1977]" pageId="7" pageNumber="550">
<emphasis id="B933EA92FF9CB504E3B1B3D1FE3651F1" box="[199,422,1951,1977]" italics="true" pageId="7" pageNumber="550">Metathoracic wings</emphasis>
: Complete.
</paragraph>
<caption id="DF386608FF93B50BE3E1B3C2FCD65181" ID-DOI="http://doi.org/10.5281/zenodo.4612459" ID-Zenodo-Dep="4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="8" pageNumber="551" startId="8.[151,264,1932,1956]" targetBox="[182,1402,193,1906]" targetPageId="8">
<paragraph id="8BF83680FF93B50BE3E1B3C2FCD65181" blockId="8.[151,1437,1932,1993]" pageId="8" pageNumber="551">
<emphasis id="B933EA92FF93B50BE3E1B3C2FED851ED" bold="true" box="[151,328,1932,1956]" pageId="8" pageNumber="551">FIGURES 8–10.</emphasis>
Holotype of 
<taxonomicName id="4C474D03FF93B50BE2B9B3C3FD4551ED" authority="Haaf, 1953" authorityName="Haaf" authorityYear="1953" box="[463,725,1932,1956]" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="551" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF93B50BE2B9B3C3FDCD51ED" box="[463,605,1932,1956]" italics="true" pageId="8" pageNumber="551">Trox fumarius</emphasis>
<bibRefCitation id="EFD64B71FF93B50BE112B3C2FD4551ED" author="Haaf, E." box="[612,725,1932,1956]" pageId="8" pageNumber="551" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
</taxonomicName>
(NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB.
</paragraph>
</caption>
<caption id="DF386608FF92B50AE3E1B3D9FA06519A" ID-DOI="http://doi.org/10.5281/zenodo.4612463" ID-Zenodo-Dep="4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="9" pageNumber="552" startId="9.[151,264,1943,1967]" targetBox="[203,1379,197,1904]" targetPageId="9">
<paragraph id="8BF83680FF92B50AE3E1B3D9FA06519A" blockId="9.[151,1436,1943,2003]" pageId="9" pageNumber="552">
<emphasis id="B933EA92FF92B50AE3E1B3D9FEC351E6" bold="true" box="[151,339,1943,1967]" pageId="9" pageNumber="552">FIGURES 11–14.</emphasis>
<taxonomicName id="4C474D03FF92B50AE22EB3D9FD3451E6" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[344,676,1943,1967]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="552" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF92B50AE22EB3D9FD8A51E6" box="[344,538,1943,1967]" italics="true" pageId="9" pageNumber="552">Phoberus fumarius</emphasis>
(
<bibRefCitation id="EFD64B71FF92B50AE151B3D9FD0C51E6" author="Haaf, E." box="[551,668,1943,1967]" pageId="9" pageNumber="552" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
, 
<emphasis id="B933EA92FF92B50AE1D8B3D9FB8551E6" bold="true" box="[686,1045,1943,1967]" pageId="9" pageNumber="552">revalidated and new combination</emphasis>
, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label.
</paragraph>
</caption>
<caption id="DF386608FF91B509E3E1B1ECFAD353F3" ID-DOI="http://doi.org/10.5281/zenodo.4612465" ID-Zenodo-Dep="4612465" box="[151,1347,1442,1467]" httpUri="https://zenodo.org/record/4612465/files/figure.png" pageId="10" pageNumber="553" startId="10.[151,264,1442,1466]" targetBox="[171,1416,181,1419]" targetPageId="10">
<paragraph id="8BF83680FF91B509E3E1B1ECFAD353F3" blockId="10.[151,1347,1442,1467]" box="[151,1347,1442,1467]" pageId="10" pageNumber="553">
<emphasis id="B933EA92FF91B509E3E1B1ECFEC453F3" bold="true" box="[151,340,1442,1466]" pageId="10" pageNumber="553">FIGURES 15–16.</emphasis>
Pronotal disc, dorsal view. 15, 
<taxonomicName id="4C474D03FF91B509E1E4B1ECFC5353F3" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[658,963,1442,1466]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF91B509E1E4B1ECFCA853F3" box="[658,824,1442,1466]" italics="true" pageId="10" pageNumber="553">Phoberus cyrtus</emphasis>
(
<bibRefCitation id="EFD64B71FF91B509E033B1ECFC2A53F3" author="Haaf, E." box="[837,954,1442,1466]" pageId="10" pageNumber="553" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
; 16, 
<taxonomicName id="4C474D03FF91B509E085B1ECFAAF53F3" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1011,1343,1442,1467]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF91B509E085B1ECFB2453F2" box="[1011,1204,1442,1467]" italics="true" pageId="10" pageNumber="553">Phoberus fumarius</emphasis>
(
<bibRefCitation id="EFD64B71FF91B509E7B4B1ECFAA753F3" author="Haaf, E." box="[1218,1335,1442,1466]" pageId="10" pageNumber="553" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF83680FF91B509E3B1B1A7FB045026" blockId="10.[151,1437,1513,2044]" pageId="10" pageNumber="553">
<emphasis id="B933EA92FF91B509E3B1B1A7FEFE504A" box="[199,366,1513,1539]" italics="true" pageId="10" pageNumber="553">Male genitalia</emphasis>
(
<figureCitation id="137C2A05FF91B509E208B1A7FE59504A" box="[382,457,1513,1539]" captionStart="FIGURES 8–10" captionStartId="8.[151,264,1932,1956]" captionTargetBox="[182,1402,193,1906]" captionTargetId="figure-31@8.[167,1419,181,1908]" captionTargetPageId="8" captionText="FIGURES 8–10. Holotype of Trox fumarius Haaf, 1953 (NHMB): 8, habitus, dorsal view; 9, aedeagus, dorsal view; 10, holotype labels. Photographs by Christoph Germann, copyright NHMB." figureDoi="http://doi.org/10.5281/zenodo.4612459" httpUri="https://zenodo.org/record/4612459/files/figure.png" pageId="10" pageNumber="553">Figs 9</figureCitation>
, 
<figureCitation id="137C2A05FF91B509E2A1B1A7FE66504A" box="[471,502,1513,1539]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[203,1379,197,1904]" captionTargetId="figure-17@9.[188,1399,181,1919]" captionTargetPageId="9" captionText="FIGURES 11–14. Phoberus fumarius (Haaf, 1953), revalidated and new combination, specimen with obscuring layer of dirt removed (TMSA). 11, habitus, dorsal view; 12, habitus, lateral view; 13, aedeagus, dorsal and lateral views; 14, locality label." figureDoi="http://doi.org/10.5281/zenodo.4612463" httpUri="https://zenodo.org/record/4612463/files/figure.png" pageId="10" pageNumber="553">13</figureCitation>
): Symmetrical, slender, phallobase arched, as long as parameres. Parameres long, extending beyond median lobe, apices pointed and curved. Median lobe divided into two symmetrical parts, these broad, not parallel, with distinct basolateral swelling, in dorsal view concave upward; apices angularly rounded and incurved; two bladelike projections and hooks visible between parameres and median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C35D650BFF91B50FE3B1B236FE2D51C8" lastPageId="12" lastPageNumber="555" pageId="10" pageNumber="553" type="diagnosis">
<paragraph id="8BF83680FF91B509E3B1B236FD0250B6" blockId="10.[151,1437,1513,2044]" pageId="10" pageNumber="553">
<emphasis id="B933EA92FF91B509E3B1B236FED250DB" bold="true" box="[199,322,1656,1682]" pageId="10" pageNumber="553">Diagnosis.</emphasis>
<taxonomicName id="4C474D03FF91B509E23EB237FD8F50DA" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[328,543,1657,1683]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF91B509E23EB237FD8F50DA" box="[328,543,1657,1683]" italics="true" pageId="10" pageNumber="553">Phoberus fumarius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C474D03FF91B509E123B237FC9B50DA" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[597,779,1657,1683]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF91B509E123B237FC9B50DA" box="[597,779,1657,1683]" italics="true" pageId="10" pageNumber="553">Phoberus cyrtus</emphasis>
</taxonomicName>
are very similar to each other. The two species can be told apart as elaborated under the diagnosis of 
<taxonomicName id="4C474D03FF91B509E104B2D3FCB950FE" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[626,809,1693,1719]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF91B509E104B2D3FCB950FE" box="[626,809,1693,1719]" italics="true" pageId="10" pageNumber="553">Phoberus cyrtus</emphasis>
</taxonomicName>
, above. Whereas both species can be confused with the common and widespread 
<taxonomicName id="4C474D03FF91B509E2C1B28FFD165092" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[439,646,1729,1755]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF91B509E2C1B28FFD165092" box="[439,646,1729,1755]" italics="true" pageId="10" pageNumber="553">Phoberus sulcatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C474D03FF91B509E1E2B28FFCFB5092" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[660,875,1729,1755]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF91B509E1E2B28FFCFB5092" box="[660,875,1729,1755]" italics="true" pageId="10" pageNumber="553">Phoberus fumarius</emphasis>
</taxonomicName>
is more likely to cause such confusion. Again see the discussion under 
<taxonomicName id="4C474D03FF91B509E2F5B2ABFDA950B6" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[387,569,1765,1791]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF91B509E2F5B2ABFDA950B6" box="[387,569,1765,1791]" italics="true" pageId="10" pageNumber="553">Phoberus cyrtus</emphasis>
</taxonomicName>
, above.
</paragraph>
<paragraph id="8BF83680FF91B509E3B1B346FB9551B5" blockId="10.[151,1437,1513,2044]" pageId="10" pageNumber="553">
<emphasis id="B933EA92FF91B509E3B1B346FE61516B" bold="true" box="[199,497,1800,1826]" pageId="10" pageNumber="553">Geographic distribution.</emphasis>
<taxonomicName id="4C474D03FF91B509E28CB347FD42516A" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[506,722,1801,1827]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF91B509E28CB347FD42516A" box="[506,722,1801,1827]" italics="true" pageId="10" pageNumber="553">Phoberus fumarius</emphasis>
</taxonomicName>
is presently known only from the 
<collectingRegion id="4983F862FF91B509E716B347FA98516A" box="[1120,1288,1801,1827]" country="South Africa" name="Northern Cape" pageId="10" pageNumber="553">Northern Cape</collectingRegion>
and 
<collectingRegion id="4983F862FF91B509E637B347FF41510E" country="South Africa" name="Western Cape" pageId="10" pageNumber="553">Western Cape</collectingRegion>
Provinces of 
<collectingCountry id="F3507610FF91B509E218B363FE6F510E" box="[366,511,1837,1863]" name="South Africa" pageId="10" pageNumber="553">South Africa</collectingCountry>
, apart from its 
<typeStatus id="54FC8822FF91B509E1DAB363FD4C510E" box="[684,732,1837,1863]" pageId="10" pageNumber="553">type</typeStatus>
locality that marginally falls in the westernmost 
<collectingRegion id="4983F862FF91B509E671B363FF6E5122" country="South Africa" name="Eastern Cape" pageId="10" pageNumber="553">Eastern Cape Province</collectingRegion>
. The known distribution of the species follows the 
<collectingRegion id="4983F862FF91B509E022B31FFC1E5122" box="[852,910,1873,1899]" country="South Africa" name="Western Cape" pageId="10" pageNumber="553">Cape</collectingRegion>
Fold Mountains, along the western mountain chain from the Richtersveld southwards through Namaqualand to the West Coast Peninsula and the 
<collectingRegion id="4983F862FF91B509E79EB33BFAB251C6" box="[1256,1314,1909,1935]" country="South Africa" name="Western Cape" pageId="10" pageNumber="553">Cape</collectingRegion>
Peninsula; and from there eastwards along the southern mountain chains as far east as the town of Willowmore (
<figureCitation id="137C2A05FF91B509E798B3D7FAD051FA" box="[1262,1344,1945,1971]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="10" pageNumber="553">Fig. 21</figureCitation>
). This is a distribution pattern that broadly repeats among many beetle species and higher categories. Another example from the 
<taxonomicName id="4C474D03FF91B509E3B6B3AFFEB451B2" baseAuthorityName="Strumpher &amp; Scholtz" baseAuthorityYear="2011" box="[192,292,2017,2043]" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
is the 
<taxonomicName id="4C474D03FF91B509E218B3AFFDD051B2" baseAuthorityName="Scholtz" baseAuthorityYear="1979" box="[366,576,2017,2043]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="553" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="B933EA92FF91B509E218B3AFFDD051B2" box="[366,576,2017,2043]" italics="true" pageId="10" pageNumber="553">Phoberus capensis</emphasis>
</taxonomicName>
species-group (
<bibRefCitation id="EFD64B71FF91B509E187B3AFFC6951B5" author="Strumpher, W. P. &amp; Sole, C. L. &amp; Villet, M. H. &amp; Scholtz, C. H." box="[753,1017,2017,2044]" pageId="10" pageNumber="553" pagination="149 - 179" refId="ref8465" refString="Strumpher, W. P., Sole, C. L., Villet, M. H. &amp; Scholtz, C. H. (2016 a) Allopatric speciation in the flightless Phoberus capensis (Coleoptera: Trogidae) group, with description of two new species. Insect Systematics &amp; Evolution, 47 (2), 149 - 179. https: // doi. org / 10.1163 / 1876312 X- 47022138" type="journal article" year="2016">
Strümpher 
<emphasis id="B933EA92FF91B509E007B3ACFC3951B2" box="[881,937,2017,2043]" italics="true" pageId="10" pageNumber="553">et al.</emphasis>
2016a
</bibRefCitation>
).
</paragraph>
<caption id="DF386608FF90B508E3E1B3D9FA8C519A" ID-DOI="http://doi.org/10.5281/zenodo.4612471" ID-Zenodo-Dep="4612471" httpUri="https://zenodo.org/record/4612471/files/figure.png" pageId="11" pageNumber="554" startId="11.[151,264,1943,1967]" targetBox="[185,1404,195,1919]" targetPageId="11">
<paragraph id="8BF83680FF90B508E3E1B3D9FA8C519A" blockId="11.[151,1436,1943,2003]" pageId="11" pageNumber="554">
<emphasis id="B933EA92FF90B508E3E1B3D9FEC651E6" bold="true" box="[151,342,1943,1967]" pageId="11" pageNumber="554">FIGURES 17–20.</emphasis>
<taxonomicName id="4C474D03FF90B508E228B3D9FD4A51E6" authority="(Thunberg, 1787)" baseAuthorityName="Thunberg" baseAuthorityYear="1787" box="[350,730,1943,1967]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="554" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B933EA92FF90B508E228B3D9FD8A51E6" box="[350,538,1943,1967]" italics="true" pageId="11" pageNumber="554">Phoberus sulcatus</emphasis>
(Thunberg, 1787)
</taxonomicName>
, cleaned specimen (BMSA). 17, habitus, dorsal view; 18, aedeagus, dorsal view; 19, pronotal disc, with red arrow indicating distinct median pronotal depression; 20, specimen labels.
</paragraph>
</caption>
<caption id="DF386608FF97B50FE3E1B03AFB2C52C4" ID-DOI="http://doi.org/10.5281/zenodo.4612475" ID-Zenodo-Dep="4612475" box="[151,1212,1140,1165]" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="12" pageNumber="555" startId="12.[151,250,1140,1164]" targetBox="[151,1434,181,1117]" targetPageId="12">
<paragraph id="8BF83680FF97B50FE3E1B03AFB2C52C4" blockId="12.[151,1212,1140,1165]" box="[151,1212,1140,1165]" pageId="12" pageNumber="555">
<emphasis id="B933EA92FF97B50FE3E1B03AFEB052C4" bold="true" box="[151,288,1140,1165]" pageId="12" pageNumber="555">FIGURE 21.</emphasis>
Known distribution of 
<taxonomicName id="4C474D03FF97B50FE178B03BFCAA52C4" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[526,826,1141,1165]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF97B50FE178B03BFD2352C5" box="[526,691,1141,1165]" italics="true" pageId="12" pageNumber="555">Phoberus cyrtus</emphasis>
(
<bibRefCitation id="EFD64B71FF97B50FE1B6B03BFCA452C4" author="Haaf, E." box="[704,820,1141,1165]" pageId="12" pageNumber="555" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C474D03FF97B50FE01AB03BFB2952C4" authority="(Haaf, 1953)" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[876,1209,1141,1165]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF97B50FE01AB03BFBBD52C4" box="[876,1069,1141,1165]" italics="true" pageId="12" pageNumber="555">Phoberus fumarius</emphasis>
(
<bibRefCitation id="EFD64B71FF97B50FE74DB03BFB2052C4" author="Haaf, E." box="[1083,1200,1141,1165]" pageId="12" pageNumber="555" pagination="309 - 346" refId="ref7655" refString="Haaf, E. (1953) Die afrikanischen Arten der Gattung Trox F. (Col. Scarab.). 1. Beitrag zur Kenntnis der Subfam. Troginae. Entomologische Arbeiten aus dem Museum G. Frey, 4, 309 - 346." type="journal article" year="1953">Haaf, 1953</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF83680FF97B50FE3B1B0F5FA8C532C" blockId="12.[151,1437,1211,1921]" pageId="12" pageNumber="555">
<materialsCitation id="3B2F3CDDFF97B50FE3B1B0F5FA85532C" ID-GBIF-Occurrence="3062300349" country="Namibia" county="Saldanha Bay" latitude="-33.052776" location="The" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" pageId="12" pageNumber="555" specimenCount="1">
<location id="8E98605BFF97B50FE3B1B0F5FF63529C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE3B1B0F5FF63529C" box="[199,243,1211,1237]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="The" pageId="12" pageNumber="555">The</location>
distribution of this species may well reach further east into the 
<location id="8E98605BFF97B50FE0B0B0F5FA88529C" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE0B0B0F5FA88529C" box="[966,1304,1211,1237]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="eastern Cape Floristic Region" pageId="12" pageNumber="555">eastern Cape Floristic Region</location>
; and it may well extend into southern 
<collectingCountry id="F3507610FF97B50FE2C0B091FD8952B0" box="[438,537,1247,1273]" name="Namibia" pageId="12" pageNumber="555">Namibia</collectingCountry>
, since it is known from the 
<location id="8E98605BFF97B50FE03FB091FC4752B0" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE03FB091FC4752B0" box="[841,983,1247,1273]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="Richtersveld" pageId="12" pageNumber="555">Richtersveld</location>
on the southern side of the 
<location id="8E98605BFF97B50FE670B091FA0852B0" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE670B091FA0852B0" box="[1286,1432,1247,1273]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="Gariep River" pageId="12" pageNumber="555">Gariep River</location>
, while identical habitat is present on the northern side of the river in 
<collectingCountry id="F3507610FF97B50FE0F5B14DFC735354" box="[899,995,1283,1309]" name="Namibia" pageId="12" pageNumber="555">Namibia</collectingCountry>
(
<figureCitation id="137C2A05FF97B50FE084B14DFBD45354" box="[1010,1092,1283,1309]" captionStart="FIGURE 21" captionStartId="12.[151,250,1140,1164]" captionTargetBox="[151,1434,181,1117]" captionTargetId="figure-31@12.[151,1435,181,1117]" captionTargetPageId="12" captionText="FIGURE 21. Known distribution of Phoberus cyrtus (Haaf, 1953) and Phoberus fumarius (Haaf, 1953)." figureDoi="http://doi.org/10.5281/zenodo.4612475" httpUri="https://zenodo.org/record/4612475/files/figure.png" pageId="12" pageNumber="555">Fig. 21</figureCitation>
). 
<taxonomicName id="4C474D03FF97B50FE72EB14DFABD5354" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1112,1325,1283,1309]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF97B50FE72EB14DFABD5354" box="[1112,1325,1283,1309]" italics="true" pageId="12" pageNumber="555">Phoberus fumarius</emphasis>
</taxonomicName>
is present on off-shore islands on the 
<location id="8E98605BFF97B50FE2B0B169FD515308" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE2B0B169FD515308" box="[454,705,1319,1345]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="Western Cape Atlantic" pageId="12" pageNumber="555">Western Cape Atlantic</location>
coast. 
<location id="8E98605BFF97B50FE079B169FCD05308" LSID="urn:lsid:plazi:treatment:03EE8796FF9DB50FE3E1B174FE2D51C8:8E98605BFF97B50FE079B169FCD05308" box="[783,832,1319,1345]" country="Namibia" county="Saldanha Bay" latitude="-33.052776" longLatPrecision="20" longitude="17.925278" municipality="Malgas Island" name="Two" pageId="12" pageNumber="555">Two</location>
of these ‘guano islands’ are noted above; 
<bibRefCitation id="EFD64B71FF97B50FE662B169FA0C530B" author="Haaf, E." box="[1300,1436,1319,1346]" pageId="12" pageNumber="555" refId="ref7702" refString="Haaf, E. (1959) Coleoptera: Trogidae. In: Hanstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life. Results of the Lund University Expedition in 1950 - 1951. Vol. 5. Almqvist &amp; Wiksell, Uppsala, pp. 473 - 478. [1958]" type="journal volume" year="1959">Haaf (1959)</bibRefCitation>
additionally reported this beetle species from 
<collectingMunicipality id="6B9CACFAFF97B50FE1E1B105FCA4532C" box="[663,820,1355,1381]" pageId="12" pageNumber="555">Malgas Island</collectingMunicipality>
(
<geoCoordinate id="EE735047FF97B50FE032B102FC56532E" box="[836,966,1356,1383]" degrees="33" direction="south" minutes="03" orientation="latitude" pageId="12" pageNumber="555" precision="15" seconds="10" value="-33.052776">33°03′10″S</geoCoordinate>
<geoCoordinate id="EE735047FF97B50FE0BBB102FBC0532E" box="[973,1104,1356,1383]" degrees="17" direction="east" minutes="55" orientation="longitude" pageId="12" pageNumber="555" precision="15" seconds="31" value="17.925278">17°55′31″E</geoCoordinate>
) in 
<collectingCounty id="62994E0CFF97B50FE70AB105FA85532C" box="[1148,1301,1355,1381]" pageId="12" pageNumber="555">Saldanha Bay</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF83680FF97B50FE3B1B121FDDD5028" blockId="12.[151,1437,1211,1921]" pageId="12" pageNumber="555">
<emphasis id="B933EA92FF97B50FE3B1B121FD9F53C0" bold="true" box="[199,527,1391,1417]" pageId="12" pageNumber="555">Broad habitat associations.</emphasis>
The known distribution of 
<taxonomicName id="4C474D03FF97B50FE039B121FBB853C0" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[847,1064,1391,1417]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF97B50FE039B121FBB853C0" box="[847,1064,1391,1417]" italics="true" pageId="12" pageNumber="555">Phoberus fumarius</emphasis>
</taxonomicName>
is confined to the Greater 
<collectingRegion id="4983F862FF97B50FE615B121FA0D53C0" box="[1379,1437,1391,1417]" country="South Africa" name="Western Cape" pageId="12" pageNumber="555">Cape</collectingRegion>
Floristic Region of 
<bibRefCitation id="EFD64B71FF97B50FE202B1DDFDD153E4" author="Born, J. &amp; Linder, H. P. &amp; Desmet, P." box="[372,577,1427,1453]" pageId="12" pageNumber="555" pagination="147 - 162" refId="ref7204" refString="Born, J., Linder, H. P. &amp; Desmet, P. (2007) The Greater Cape Floristic Region. Journal of Biogeography, 34 (1), 147 - 162. https: // doi. org / 10.1111 / j. 1365 - 2699.2006.01595. x" type="journal article" year="2007">
Born 
<emphasis id="B933EA92FF97B50FE2C2B1DAFE7D53E4" box="[436,493,1427,1453]" italics="true" pageId="12" pageNumber="555">et al.</emphasis>
(2007)
</bibRefCitation>
, falling entirely in the winter rainfall region of southern Africa, whereas the range of the putative sister species 
<taxonomicName id="4C474D03FF97B50FE150B1F9FD4D5398" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[550,733,1463,1489]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="cyrtus">
<emphasis id="B933EA92FF97B50FE150B1F9FD4D5398" box="[550,733,1463,1489]" italics="true" pageId="12" pageNumber="555">Phoberus cyrtus</emphasis>
</taxonomicName>
is confined to the summer rainfall region. 
<taxonomicName id="4C474D03FF97B50FE7B3B1F9FA0D5398" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[1221,1437,1463,1489]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF97B50FE7B3B1F9FA0D5398" box="[1221,1437,1463,1489]" italics="true" pageId="12" pageNumber="555">Phoberus fumarius</emphasis>
</taxonomicName>
has been recorded at an altitude of 
<quantity id="4CBF9B65FF97B50FE16FB195FDE553BC" box="[537,629,1499,1525]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="555" unit="m" value="2000.0">2,000 m</quantity>
above sea level in the Swartberg Mountains and close to sea level on the West Coast and the 
<collectingRegion id="4983F862FF97B50FE207B1B1FE3B5050" box="[369,427,1535,1561]" country="South Africa" name="Western Cape" pageId="12" pageNumber="555">Cape</collectingRegion>
Peninsula. No field-collected information about habitat associations exists for this species, and the possible vegetation 
<typeStatus id="54FC8822FF97B50FE2A5B26DFD9E5074" box="[467,526,1571,1597]" pageId="12" pageNumber="555">types</typeStatus>
in which it occurs are geographically complexly arranged, necessitating caution when inferring where the species lives.
</paragraph>
<paragraph id="8BF83680FF97B50FE3B1B225FE2D51C8" blockId="12.[151,1437,1211,1921]" pageId="12" pageNumber="555">
The distribution of 
<taxonomicName id="4C474D03FF97B50FE2E9B225FDE550CC" baseAuthorityName="Haaf" baseAuthorityYear="1953" box="[415,629,1643,1669]" class="Insecta" family="Trogidae" genus="Phoberus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="555" phylum="Arthropoda" rank="species" species="fumarius">
<emphasis id="B933EA92FF97B50FE2E9B225FDE550CC" box="[415,629,1643,1669]" italics="true" pageId="12" pageNumber="555">Phoberus fumarius</emphasis>
</taxonomicName>
is in the northwest centred on the Succulent Karoo Biome (Richtersveld to Namaqualand) and on the Fynbos Biome in the south and towards the east (biomes, bioregions and vegetation units 
<emphasis id="B933EA92FF97B50FE3A0B2FAFE845084" box="[214,276,1716,1741]" italics="true" pageId="12" pageNumber="555">sensu</emphasis>
<bibRefCitation id="EFD64B71FF97B50FE26AB2FDFDC75084" author="Mucina, L. &amp; Rutherford, M. C." box="[284,599,1715,1741]" pageId="12" pageNumber="555" pagination="1 - 807" refId="ref7875" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. South African National Biodiversity Institute, Pretoria, Strelitzia, 19, i - viii + 1 - 807." type="journal article" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
; 
<bibRefCitation id="EFD64B71FF97B50FE115B2FDFCC05084" author="Dayaram, A. &amp; Harris, L. R. &amp; Grobler, B. A. &amp; Van der Merwe, S. &amp; Rebelo, A. G. &amp; Powrie, L. W. &amp; Vlok, J. H. J. &amp; Desmet, P. G. &amp; Qabaqaba, M. &amp; Hlahane, K. M. &amp; Skowno, A. L." box="[611,848,1715,1741]" pageId="12" pageNumber="555" pagination="1 - 11" refId="ref7440" refString="Dayaram, A., Harris, L. R., Grobler, B. A., Van der Merwe, S., Rebelo, A. G., Powrie, L. W., Vlok, J. H. J., Desmet, P. G., Qabaqaba, M., Hlahane, K. M. &amp; Skowno, A. L. (2019) Vegetation map of South Africa, Lesotho and Swaziland 2018: a description of changes since 2006. Bothalia, 49 (1), a 2452, 1 - 11. https: // doi. org / 10.4102 / abc. v 49 i 1.2452" type="journal article" year="2019">
Dayaram 
<emphasis id="B933EA92FF97B50FE1A7B2FAFC9B5084" box="[721,779,1715,1741]" italics="true" pageId="12" pageNumber="555">et al.</emphasis>
2019
</bibRefCitation>
). In the Fynbos Biome the species may likely occur in both Fynbos and Renosterveld vegetation units, but dedicated collecting is required to confirm or refute this. At higher altitudes along the southern mountains the species may again be found in Succulent Karoo habitats, and specifically in the Rainshadow Valley Karoo Bioregion. In the Swartberg and Little Karoo it possibly occurs in intrusions of the Albany Thicket Biome. On the off-shore islands the species occurs in Western Strandveld, a vegetation unit of the Fynbos Biome.
</paragraph>
</subSubSection>
</treatment>
</document>