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163 lines
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<document id="B41371B782D81AD3F91B045C7AD836C0" ID-DOI="10.11646/zootaxa.5541.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240970" ID-ZooBank="BD9AB41C-245E-4B76-B477-EB92DE6FA63C" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1732834480828" checkinUser="plazi" docAuthor="Zonstein, Sergei & Marusik, Yuri M." docDate="2024" docId="03E987B6FFCAFF8F2BF5FB57FA3EB43C" docLanguage="en" docName="zootaxa.5541.2.6.pdf" docOrigin="Zootaxa 5541 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5541.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brignoliolus caudatus, comb. nov." docType="treatment" docVersion="5" lastPageNumber="217" masterDocId="FFD0FFCEFFC8FF8D2B62FFC8FFB7B224" masterDocTitle="A survey of Brignoliolus Ovchinnikov, 1999 stat. rev. (Araneae, Agelenidae) with the description of a new species from Israel" masterLastPageNumber="226" masterPageNumber="215" pageNumber="217" updateTime="1733421597060" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A7F35AD7F42B72925324702E6308B748">A survey of Brignoliolus Ovchinnikov, 1999 stat. rev. (Araneae, Agelenidae) with the description of a new species from Israel</mods:title>
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<mods:namePart id="77CEAF4085BE267C66277FA66F740B1E">Zonstein, Sergei</mods:namePart>
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<mods:nameIdentifier id="75D3F6A606AAB46A92CA7D704824AAD3" type="ORCID">0000-0002-4515-0630</mods:nameIdentifier>
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<mods:affiliation id="B15B9985E83F739CA94EB00B1A1352EC">The Steinhardt Museum of Natural History, Klausner 12, 69978 Tel-Aviv, Israel.</mods:affiliation>
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<mods:nameIdentifier id="F8C1E3A9D920383FD2F49B6257FE815A" type="email">znn@tauex.tau.ac.il</mods:nameIdentifier>
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<mods:namePart id="1C8CB7C5CA4E055CBD8B46A6EC4BC891">Marusik, Yuri M.</mods:namePart>
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<mods:nameIdentifier id="CC7EFD98A504AB451EC1AE296392D22E" type="ORCID">0000-0002-4499-5148</mods:nameIdentifier>
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<mods:affiliation id="46BCF226E5FDB87D70B268347F9FDE47">Institute for Biological Problems of the North, Portovaya Str. 18, Magadan 685000, Russia. & Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia.</mods:affiliation>
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<mods:nameIdentifier id="CD5DAF897F64D244646AAD9A33D0AACE" type="email">yurmar@mail.ru</mods:nameIdentifier>
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<mods:title id="83958D5514BC38C37AB69B7CA9E1D4B2">Zootaxa</mods:title>
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<mods:date id="3939FFF8DEFAB69EB0C4756655EF6209">2024</mods:date>
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<mods:number id="3A89E186978DAA9718D36C2CDE1D7BD6">2024-11-26</mods:number>
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<mods:number id="1340DA79AF2FF502A74B800454D5E592">5541</mods:number>
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<treatment id="03E987B6FFCAFF8F2BF5FB57FA3EB43C" ID-DOI="http://doi.org/10.5281/zenodo.14248365" ID-Zenodo-Dep="14248365" LSID="urn:lsid:plazi:treatment:03E987B6FFCAFF8F2BF5FB57FA3EB43C" httpUri="http://treatment.plazi.org/id/03E987B6FFCAFF8F2BF5FB57FA3EB43C" lastPageNumber="217" pageId="2" pageNumber="217">
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<subSubSection id="C35A652BFFCAFF8F2BF5FB57FCA0B69E" box="[151,791,1183,1210]" pageId="2" pageNumber="217" type="nomenclature">
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<paragraph id="8BFF36A0FFCAFF8F2BF5FB57FCA0B69E" blockId="2.[151,791,1183,1246]" box="[151,791,1183,1210]" pageId="2" pageNumber="217">
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<heading id="D0B781CCFFCAFF8F2BF5FB57FCA0B69E" bold="true" box="[151,791,1183,1210]" fontSize="11" level="1" pageId="2" pageNumber="217" reason="1">
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<emphasis id="B934EAB2FFCAFF8F2BF5FB57FCA0B69E" bold="true" box="[151,791,1183,1210]" pageId="2" pageNumber="217">
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<taxonomicName id="4C404D23FFCAFF8F2BF5FB57FD3EB69E" authority="(de Blauwe, 1973)" baseAuthorityName="de Blauwe" baseAuthorityYear="1973" box="[151,649,1183,1210]" class="Arachnida" family="Agelenidae" genus="Brignoliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="caudatus" status="comb. nov.">
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<emphasis id="B934EAB2FFCAFF8F2BF5FB57FE14B69E" bold="true" box="[151,419,1183,1210]" italics="true" pageId="2" pageNumber="217">Brignoliolus caudatus</emphasis>
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(de
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<bibRefCitation id="EFD14B51FFCAFF8F2ABBFB68FD36B69E" author="Blauwe, R. de" box="[473,641,1184,1210]" pageId="2" pageNumber="217" pagination="1 - 111" refId="ref6569" refString="Blauwe, R. de (1973) Revision de la famille des Agelenidae (Araneae) de la region mediterraneenne. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 49 (2), 1 - 111." type="journal article" year="1973">Blauwe, 1973</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A20757C9FFCAFF8F29F2FB68FCA0B69E" box="[656,791,1184,1210]" pageId="2" pageNumber="217" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A652BFFCAFF8F2BF5FB0CFF50B6FA" box="[151,231,1220,1246]" pageId="2" pageNumber="217" type="description">
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<paragraph id="8BFF36A0FFCAFF8F2BF5FB0CFF50B6FA" blockId="2.[151,791,1183,1246]" box="[151,231,1220,1246]" pageId="2" pageNumber="217">
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<figureCitation id="137B2A25FFCAFF8F2BF5FB0CFF50B6FA" box="[151,231,1220,1246]" captionStart="FIGURE 31" captionStartId="9.[151,253,1344,1368]" captionTargetBox="[151,1435,702,1320]" captionTargetId="figure-69@9.[151,1435,702,1320]" captionTargetPageId="9" captionText="FIGURE 31. Distribution records of Brignoliolus spp.: ✖B. arganoi; ■B. caudatus; B. charitonovi; ✦B. coenobita; B. juglandicola; ¢B. nenilini; B. ovchinnikovi sp. nov.; B. turkestanicus; B. vignai." figureDoi="http://doi.org/10.5281/zenodo.14284953" httpUri="https://zenodo.org/record/14284953/files/figure.png" pageId="2" pageNumber="217">Fig. 31</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A652BFFCAFF8F2BF5FAC0FCB1B71B" pageId="2" pageNumber="217" type="reference_group">
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<paragraph id="8BFF36A0FFCAFF8F2BF5FAC0FCB1B71B" blockId="2.[151,1385,1288,1343]" pageId="2" pageNumber="217">
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<treatmentCitationGroup id="AB50118EFFCAFF8F2BF5FAC0FADEB705" box="[151,1385,1288,1313]" pageId="2" pageNumber="217">
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<treatmentCitation id="0AE110B1FFCAFF8F2BF5FAC0FD9CB705" author="Blauwe, R. de" box="[151,555,1288,1313]" page="31" pageId="2" pageNumber="217" year="1973">
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<taxonomicName id="4C404D23FFCAFF8F2BF5FAC0FD9CB705" ID-CoL="WTMT" authority="de Blauwe, 1973: 31" authorityName="de Blauwe" authorityPageNumber="31" authorityYear="1973" box="[151,555,1288,1313]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="caudatus">
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<emphasis id="B934EAB2FFCAFF8F2BF5FAC0FEE6B704" box="[151,337,1288,1312]" italics="true" pageId="2" pageNumber="217">Coelotes caudatus</emphasis>
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de
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<bibRefCitation id="EFD14B51FFCAFF8F2A14FAC0FD9CB705" author="Blauwe, R. de" box="[374,555,1288,1313]" pageId="2" pageNumber="217" pagination="1 - 111" refId="ref6569" refString="Blauwe, R. de (1973) Revision de la famille des Agelenidae (Araneae) de la region mediterraneenne. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 49 (2), 1 - 111." type="journal article" year="1973">Blauwe, 1973: 31</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 29 (
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<specimenCount id="9D46FD29FFCAFF8F29EBFAC1FD2DB705" box="[649,666,1289,1313]" count="1" pageId="2" pageNumber="217" type="female">♀</specimenCount>
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);
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<treatmentCitation id="0AE110B1FFCAFF8F29D2FAC0FCC4B705" author="Brignoli, P. M." box="[688,883,1288,1313]" page="207" pageId="2" pageNumber="217" year="1978">
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<bibRefCitation id="EFD14B51FFCAFF8F29D2FAC0FCC4B705" author="Brignoli, P. M." box="[688,883,1288,1313]" pageId="2" pageNumber="217" pagination="461 - 541" refId="ref6610" refString="Brignoli, P. M. (1978) Ragni di Turchia V. Specie nuove o interessanti, cavernicole ed epigee, di varie famiglie (Araneae). Revue Suisse de Zoologie, 85 (3), 461 - 541. https: // doi. org / 10.5962 / bhl. part. 82243" type="journal article" year="1978">Brignoli 1978: 207</bibRefCitation>
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</treatmentCitation>
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, fig. 5 (
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<specimenCount id="9D46FD29FFCAFF8F28A6FAC1FC62B705" box="[964,981,1289,1313]" count="1" pageId="2" pageNumber="217" type="female">♀</specimenCount>
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);
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<treatmentCitation id="0AE110B1FFCAFF8F2888FAC1FB6BB705" author="Wang, X. P. & Zhu, M. S." box="[1002,1244,1288,1313]" page="274" pageId="2" pageNumber="217" year="2009">
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<bibRefCitation id="EFD14B51FFCAFF8F2888FAC1FB6BB705" author="Wang, X. P. & Zhu, M. S." box="[1002,1244,1288,1313]" pageId="2" pageNumber="217" pagination="272 - 281" refId="ref7657" refString="Wang, X. P. & Zhu, M. S. (2009) On the charitonovi species group of the spider genus Coelotes (Araneae: Amaurobiidae). Journal of Arachnology, 37, 272 - 281. https: // doi. org / 10.1636 / a 08 - 105.1" type="journal article" year="2009">Wang & Zhu 2009: 274</bibRefCitation>
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</treatmentCitation>
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, figs 4–6 (
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<specimenCount id="9D46FD29FFCAFF8F2E28FAC1FAECB705" box="[1354,1371,1289,1313]" count="1" pageId="2" pageNumber="217" type="female">♀</specimenCount>
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).
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</treatmentCitationGroup>
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<treatmentCitationGroup id="AB50118EFFCAFF8F2BF5FAEFFCB1B71B" box="[151,774,1319,1343]" pageId="2" pageNumber="217">
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<taxonomicName id="4C404D23FFCAFF8F2BF5FAEFFE50B71B" ID-CoL="WTMT" authorityName="de Blauwe" authorityYear="1973" box="[151,487,1319,1343]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="caudatus" subGenus="Brignoliolus">
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<emphasis id="B934EAB2FFCAFF8F2BF5FAEFFF47B71B" box="[151,240,1319,1343]" italics="true" pageId="2" pageNumber="217">Coelotes</emphasis>
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(
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<emphasis id="B934EAB2FFCAFF8F2B9DFAEFFEC9B71B" box="[255,382,1319,1343]" italics="true" pageId="2" pageNumber="217">Brignoliolus</emphasis>
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)
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<emphasis id="B934EAB2FFCAFF8F2AEEFAEFFE50B71B" box="[396,487,1319,1343]" italics="true" pageId="2" pageNumber="217">caudatus</emphasis>
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</taxonomicName>
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:
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<treatmentCitation id="0AE110B1FFCAFF8F2A97FAEFFD6BB71B" author="Ovchinnikov, S. V." box="[501,732,1319,1343]" page="74" pageId="2" pageNumber="217" year="1999">
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<bibRefCitation id="EFD14B51FFCAFF8F2A97FAEFFD6BB71B" author="Ovchinnikov, S. V." box="[501,732,1319,1343]" pageId="2" pageNumber="217" pagination="63 - 80" refId="ref7423" refString="Ovchinnikov, S. V. (1999) On the supraspecific systematics of the subfamily Coelotinae (Araneae, Amaurobiidae) in the former USSR fauna. Tethys Entomological Research, 1, 63 - 80. [in Russian]" type="journal article" year="1999">Ovchinnikov 1999: 74</bibRefCitation>
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</treatmentCitation>
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, 75.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A652BFFCAFF8F2BF5FAA5FF02B7D7" pageId="2" pageNumber="217" type="discussion">
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<paragraph id="8BFF36A0FFCAFF8F2BF5FAA5FF02B7D7" blockId="2.[151,1436,1389,1560]" pageId="2" pageNumber="217">
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<emphasis id="B934EAB2FFCAFF8F2BF5FAA5FEA8B7A3" bold="true" box="[151,287,1389,1415]" pageId="2" pageNumber="217">Comments.</emphasis>
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Females of this species can be distinguished from those of
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<taxonomicName id="4C404D23FFCAFF8F28A9FAA6FBCDB7A3" authorityName="Zonstein & Marusik" authorityYear="2024" box="[971,1146,1389,1416]" class="Arachnida" family="Agelenidae" genus="Brignoliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="ovchinnikovi" status="sp. nov.">
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<emphasis id="B934EAB2FFCAFF8F28A9FAA6FC6BB7A3" box="[971,988,1390,1415]" italics="true" pageId="2" pageNumber="217">B</emphasis>
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.
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<emphasis id="B934EAB2FFCAFF8F2889FAA5FBCDB7A3" box="[1003,1146,1389,1415]" italics="true" pageId="2" pageNumber="217">ovchinnikovi</emphasis>
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</taxonomicName>
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<emphasis id="B934EAB2FFCAFF8F2FE1FAA5FB69B7A3" bold="true" box="[1155,1246,1389,1415]" pageId="2" pageNumber="217">
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<taxonomicNameLabel id="A20757C9FFCAFF8F2FE1FAA5FB69B7A3" box="[1155,1246,1389,1415]" pageId="2" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName id="4C404D23FFCAFF8F2E75FAA6FAC9B7A3" authorityName="Zonstein & Marusik" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[1303,1406,1390,1416]" class="Arachnida" family="Agelenidae" genus="Brignoliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="vignai">
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<emphasis id="B934EAB2FFCAFF8F2E75FAA6FA9FB7A3" box="[1303,1320,1390,1415]" italics="true" pageId="2" pageNumber="217">B</emphasis>
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.
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<emphasis id="B934EAB2FFCAFF8F2E5AFAA6FAC9B7A3" box="[1336,1406,1390,1415]" italics="true" pageId="2" pageNumber="217">vignai</emphasis>
|
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</taxonomicName>
|
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|
|
in having longer and slightly diverging epigynal teeth and a considerably shorter septum (
|
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|
|
<figureCitation id="137B2A25FFCAFF8F2F06FA59FB54B788" box="[1124,1251,1425,1452]" captionStart="FIGURES 24–27" captionStartId="8.[151,264,1167,1191]" captionTargetBox="[151,1435,181,1143]" captionTargetId="figure-28@8.[151,1435,181,1143]" captionTargetPageId="8" captionText="FIGURES 24–27. Brignoliolus ovchinnikovi sp. nov., paratype female. 24 habitus, lateral; 25 epigyne, ventral; 26 cleaned epigyne, ventral; 27 vulva, dorsal. Scale bars: 24 = 2 mm, 25–27 = 0.5 mm. Abbreviations: Am anterior margin of fovea; Et tooth of epigyne; Fo fovea; Re receptacle; Rh head of receptacle." figureDoi="http://doi.org/10.5281/zenodo.14240980" httpUri="https://zenodo.org/record/14240980/files/figure.png" pageId="2" pageNumber="217">Figs 25–27</figureCitation>
|
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|
|
<emphasis id="B934EAB2FFCAFF8F2F88FA59FB49B78F" box="[1258,1278,1425,1451]" italics="true" pageId="2" pageNumber="217">cf</emphasis>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation id="EFD14B51FFCAFF8F2E69FA5AFF65B7EB" author="Wang, X. P. & Zhu, M. S." pageId="2" pageNumber="217" pagination="272 - 281" refId="ref7657" refString="Wang, X. P. & Zhu, M. S. (2009) On the charitonovi species group of the spider genus Coelotes (Araneae: Amaurobiidae). Journal of Arachnology, 37, 272 - 281. https: // doi. org / 10.1636 / a 08 - 105.1" type="journal article" year="2009">Wang & Zhu 2009</bibRefCitation>
|
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|
|
, figs 4, 34), and from that of
|
|||
|
|
<taxonomicName id="4C404D23FFCAFF8F2944FA7EFD28B7EB" authorityName="Brignoli" authorityYear="1978" box="[550,671,1462,1488]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="arganoi">
|
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|
|
<emphasis id="B934EAB2FFCAFF8F2944FA7EFD80B7EB" box="[550,567,1462,1487]" italics="true" pageId="2" pageNumber="217">B</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B934EAB2FFCAFF8F2925FA7EFD28B7EB" box="[583,671,1462,1487]" italics="true" pageId="2" pageNumber="217">arganoi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by distinctly shorter epigynal teeth (see
|
|||
|
|
<bibRefCitation id="EFD14B51FFCAFF8F2F12FA7EFAFFB7EB" author="Wang, X. P. & Zhu, M. S." box="[1136,1352,1461,1488]" pageId="2" pageNumber="217" pagination="272 - 281" refId="ref7657" refString="Wang, X. P. & Zhu, M. S. (2009) On the charitonovi species group of the spider genus Coelotes (Araneae: Amaurobiidae). Journal of Arachnology, 37, 272 - 281. https: // doi. org / 10.1636 / a 08 - 105.1" type="journal article" year="2009">Wang & Zhu 2009</bibRefCitation>
|
|||
|
|
, figs 1, 4).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C35A652BFFCAFF8F2BA5FA35FA3EB43C" box="[199,1417,1533,1560]" pageId="2" pageNumber="217" type="distribution">
|
|||
|
|
<paragraph id="8BFF36A0FFCAFF8F2BA5FA35FA3EB43C" blockId="2.[151,1436,1389,1560]" box="[199,1417,1533,1560]" pageId="2" pageNumber="217">
|
|||
|
|
<emphasis id="B934EAB2FFCAFF8F2BA5FA35FED6B433" bold="true" box="[199,353,1533,1559]" pageId="2" pageNumber="217">Distribution.</emphasis>
|
|||
|
|
The species is known from several localities in
|
|||
|
|
<collectingCountry id="F3577630FFCAFF8F2819FA35FC57B433" box="[891,992,1533,1559]" name="Lebanon" pageId="2" pageNumber="217">Lebanon</collectingCountry>
|
|||
|
|
, at an altitude between
|
|||
|
|
<quantity id="4CB89B45FFCAFF8F2F85FA35FA32B43C" box="[1255,1413,1533,1560]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.6" metricValueMin="1.4" pageId="2" pageNumber="217" unit="m" value="2000.0" valueMax="2600.0" valueMin="1400.0">1400–2600 m</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|