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<document id="DA724257DE570ED7C80144188B466E68" ID-CLB-Dataset="26002" ID-DOI="10.11646/zootaxa.4657.1.2" ID-GBIF-Dataset="a7c3cd9d-d4be-4686-a198-3255c948beb8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3369801" ID-ZooBank="F27CB967-C001-436A-9F25-184D5484363C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565968506341" checkinUser="plazi" docAuthor="Perkins, Philip D. &amp; Bergsten, Johannes" docDate="2019" docId="AD076747FFD5FFC4FF6DFBC0B7E697DB" docLanguage="en" docName="zootaxa.4657.1.2.pdf" docOrigin="Zootaxa 4657 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Incoltorrida Perkins &amp; Bergsten 2019" docType="key" docVersion="11" lastPageNumber="64" masterDocId="513E1F3FFFD2FFC3FFFAFFB9B27A9138" masterDocTitle="New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)" masterLastPageNumber="96" masterPageNumber="57" pageNumber="64" updateTime="1698751487288" updateUser="ExternalLinkService">
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<mods:title id="6AAA0972443F8176092B1F2F24678D6C">New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae)</mods:title>
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<mods:namePart id="9AEE596C3A698592E09D86AAE54278B7">Perkins, Philip D.</mods:namePart>
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<mods:namePart id="2C74C1CA02D3F5746B1237D23B530262">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="40D28897F437929303A29F8C3C6B81CB">text</mods:typeOfResource>
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<mods:title id="E42C3F48C7158ED00B1751F8ACF0AEE9">Zootaxa</mods:title>
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<mods:part id="02E90BCFF6492DDEB16C287C179272D0">
<mods:date id="4A7763AB2C5B08A5ED74AD9AC0473BA8">2019</mods:date>
<mods:detail id="95761EFF5F8587E9CC6B2861429D98F1" type="pubDate">
<mods:number id="C9B4A26A80975692167ADBDB7EB1C2D9">2019-08-16</mods:number>
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<mods:number id="86F743F4EB6EE18FB828E8EEA3299A46">4657</mods:number>
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<mods:identifier id="25A15F8C2817248F1FDE2ED2CCD4C581" type="DOI">10.11646/zootaxa.4657.1.2</mods:identifier>
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<treatment id="AD076747FFD5FFC4FF6DFBC0B7E697DB" ID-GBIF-Taxon="159131431" LSID="urn:lsid:plazi:treatment:AD076747FFD5FFC4FF6DFBC0B7E697DB" httpUri="http://treatment.plazi.org/id/AD076747FFD5FFC4FF6DFBC0B7E697DB" lastPageNumber="64" pageId="7" pageNumber="64">
<subSubSection id="6DB485DAFFD5FFC4FF6DFBC0B0D695AC" box="[151,684,1145,1172]" pageId="7" pageNumber="64" type="nomenclature">
<paragraph id="2511D651FFD5FFC4FF6DFBC0B0D695AC" blockId="7.[151,684,1145,1172]" box="[151,684,1145,1172]" pageId="7" pageNumber="64">
<heading id="7E59613DFFD5FFC4FF6DFBC0B0D695AC" bold="true" box="[151,684,1145,1172]" fontSize="11" level="1" pageId="7" pageNumber="64" reason="1">
<emphasis id="17DA0A43FFD5FFC4FF6DFBC0B0D695AC" bold="true" box="[151,684,1145,1172]" pageId="7" pageNumber="64">
Identification key to species of
<taxonomicName id="E2AEADD2FFD5FFC4FDE3FBC0B0D695AC" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[537,684,1145,1172]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="17DA0A43FFD5FFC4FDE3FBC0B0D695AC" bold="true" box="[537,684,1145,1172]" italics="true" pageId="7" pageNumber="64">Incoltorrida</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6DB485DAFFD5FFC4FF6DFB07B7E697DB" pageId="7" pageNumber="64" type="key">
<key id="513F8160FFD5FFC4FF6DFB07B7E697DB" pageId="7" pageNumber="64">
<keyStep id="9E5ACEF4FFD5FFC4FF6DFB07B7E695C8" pageId="7" pageNumber="64">
<paragraph id="2511D651FFD5FFC4FF6DFB07B7E695EB" blockId="7.[151,1437,1213,1764]" box="[151,1436,1213,1235]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFB07B7E695EB" box="[151,1436,1213,1235]" pageId="7" pageNumber="64">1a. Each elytron with five costae............................................................................ 2</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DFB62B7E695C8" blockId="7.[151,1437,1213,1764]" box="[151,1436,1243,1265]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFB62B7E695C8" box="[151,1436,1243,1265]" pageId="7" pageNumber="64">1b. Each elytron with eight costae........................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="9E5ACEF4FFD5FFC4FF6DFB41B7E6945D" pageId="7" pageNumber="64">
<paragraph id="2511D651FFD5FFC4FF6DFB41B7E79413" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFB41B7E79413" pageId="7" pageNumber="64">
2a. Larger species (&gt;
<quantity id="E2567BB4FFD5FFC4FE7DFB41B3C09436" box="[391,442,1272,1294]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="64" unit="mm" value="2.0">2mm</quantity>
) and more elongate body shape. Pronotum lacking distinct midlongitudinal carina anteriorly................................................................................................
<taxonomicName id="E2AEADD2FFD5FFC4FAFCFAAFB7E79413" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1286,1437,1302,1323]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="quintacostata">
<emphasis id="17DA0A43FFD5FFC4FAFCFAAFB7E79413" box="[1286,1437,1302,1323]" italics="true" pageId="7" pageNumber="64">I. quintacostata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DFA8AB7E6945D" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFA8AB7E6945D" pageId="7" pageNumber="64">
2b. Smaller species (&lt;
<quantity id="E2567BB4FFD5FFC4FE68FA8AB3BC9471" box="[402,454,1331,1353]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="64" unit="mm" value="2.0">2mm</quantity>
) and shorter broader body shape. Pronotum with distinct midlongitudinal carina anteriorly......................................................................................................
<taxonomicName id="E2AEADD2FFD5FFC4FABDFAE8B7E6945D" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1351,1436,1360,1382]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="galoko">
<emphasis id="17DA0A43FFD5FFC4FABDFAE8B7E6945D" box="[1351,1436,1360,1382]" italics="true" pageId="7" pageNumber="64">I. galoko</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9E5ACEF4FFD5FFC4FF6DFAD7B7E69499" pageId="7" pageNumber="64">
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<keyLead id="9E5F7564FFD5FFC4FF6DFAD7B7E694BB" box="[151,1436,1389,1411]" pageId="7" pageNumber="64">
3a. Apical part of male aedeagus without spines. Body length&gt;
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............................................... 4
</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DFA32B7E69499" blockId="7.[151,1437,1213,1764]" box="[151,1436,1419,1441]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFA32B7E69499" box="[151,1436,1419,1441]" pageId="7" pageNumber="64">
3b. Apical part of male aedeagus spinose. Body length above or below
<quantity id="E2567BB4FFD5FFC4FC90FA32B1E19498" box="[874,923,1419,1440]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="64" unit="mm" value="2.0">2mm</quantity>
......................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9E5ACEF4FFD5FFC4FF6DFA11B7E6970B" pageId="7" pageNumber="64">
<paragraph id="2511D651FFD5FFC4FF6DFA11B7E694C0" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DFA11B7E694C0" pageId="7" pageNumber="64">
4a. Basal part of pronotum with two carinae. The two transverse ridges of elytron between costae #3-#5 and between costae #5 and #8 offset from each other. Aedeagus evenly wide along most of its length in ventral view and tips not diverging (
<figureCitation id="BD95CAD4FFD5FFC4FAE0FA7CB71A94E4" box="[1306,1376,1477,1500]" captionStart="FIGURES 19–20" captionStartId="17.[151,264,1761,1786]" captionTargetBox="[235,1353,486,1744]" captionTargetId="figure@17.[235,1353,486,1744]" captionTargetPageId="17" captionText="FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype." httpUri="https://zenodo.org/record/3369829/files/figure.png" pageId="7" pageNumber="64">Fig. 19</figureCitation>
)............................................................................................
<taxonomicName id="E2AEADD2FFD5FFC4FAF1FA5DB7E694C0" authorityName="Steffan" authorityYear="1973" box="[1291,1436,1507,1529]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="madagassica">
<emphasis id="17DA0A43FFD5FFC4FAF1FA5DB7E694C0" box="[1291,1436,1507,1529]" italics="true" pageId="7" pageNumber="64">I. madagassica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DF9B9B7E6970B" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DF9B9B7E6970B" pageId="7" pageNumber="64">
4b. Basal part of pronotum without carinae. The two transverse ridges of elytron between costae #3-#5 and between costae #5 and #8 continuous. Aedeagus widest preapically and tips diverging (
<figureCitation id="BD95CAD4FFD5FFC4FCB2F9A4B1F4970C" box="[840,910,1565,1588]" captionStart="FIGURES 21–22" captionStartId="18.[151,264,1508,1533]" captionTargetBox="[151,1410,214,1442]" captionTargetId="figure@18.[151,1436,204,1459]" captionTargetPageId="18" captionText="FIGURES 21–22. Incoltorrida, male genitalia of holotypes. (21) I. magna; (22) I. marojejy." httpUri="https://zenodo.org/record/3369831/files/figure.png" pageId="7" pageNumber="64">Fig. 21</figureCitation>
)....................................
<taxonomicName id="E2AEADD2FFD5FFC4FABDF9A7B7E6970B" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1351,1436,1566,1587]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="17DA0A43FFD5FFC4FABDF9A7B7E6970B" box="[1351,1436,1566,1587]" italics="true" pageId="7" pageNumber="64">I. magna</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9E5ACEF4FFD5FFC4FF6DF982B7E69756" pageId="7" pageNumber="64">
<paragraph id="2511D651FFD5FFC4FF6DF982B7E79768" blockId="7.[151,1437,1213,1764]" box="[151,1437,1595,1617]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DF982B7E79768" box="[151,1437,1595,1617]" pageId="7" pageNumber="64">
5a. Small species (&lt;
<quantity id="E2567BB4FFD5FFC4FE7AF982B3CC9769" box="[384,438,1595,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="64" unit="mm" value="2.0">2mm</quantity>
). Basal part of pronotum with both a midlongitudinal carina and two lateral carinae.....
<taxonomicName id="E2AEADD2FFD5FFC4FAECF985B7E79768" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1302,1437,1595,1617]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="benesculpta">
<emphasis id="17DA0A43FFD5FFC4FAECF985B7E79768" box="[1302,1437,1595,1617]" italics="true" pageId="7" pageNumber="64">I. benesculpta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DF9E1B7E69756" blockId="7.[151,1437,1213,1764]" box="[151,1436,1624,1646]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DF9E1B7E69756" box="[151,1436,1624,1646]" pageId="7" pageNumber="64">
5b. Medium-sized species (approx.
<quantity id="E2567BB4FFD5FFC4FDEEF9E1B0339756" box="[532,585,1624,1646]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="64" unit="mm" value="2.0">2mm</quantity>
). Basal part of pronotum without carinae..................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9E5ACEF4FFD5FFC4FF6DF9CCB7E697DB" pageId="7" pageNumber="64">
<paragraph id="2511D651FFD5FFC4FF6DF9CCB7E69791" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DF9CCB7E69791" pageId="7" pageNumber="64">
6a. Elytral costae strongly developed and anterior oblique carinae of pronotum distinct. Aedeagus in lateral view with apex distinctly narrowed (
<figureCitation id="BD95CAD4FFD5FFC4FE77F92AB3A99791" box="[397,467,1683,1705]" captionStart="FIGURES 21–22" captionStartId="18.[151,264,1508,1533]" captionTargetBox="[151,1410,214,1442]" captionTargetId="figure@18.[151,1436,204,1459]" captionTargetPageId="18" captionText="FIGURES 21–22. Incoltorrida, male genitalia of holotypes. (21) I. magna; (22) I. marojejy." httpUri="https://zenodo.org/record/3369831/files/figure.png" pageId="7" pageNumber="64">Fig. 22</figureCitation>
).......................................................................
<taxonomicName id="E2AEADD2FFD5FFC4FACEF92DB7E69791" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1332,1436,1684,1705]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="marojejy">
<emphasis id="17DA0A43FFD5FFC4FACEF92DB7E69791" box="[1332,1436,1684,1705]" italics="true" pageId="7" pageNumber="64">I. marojejy</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2511D651FFD5FFC4FF6DF909B7E697DB" blockId="7.[151,1437,1213,1764]" pageId="7" pageNumber="64">
<keyLead id="9E5F7564FFD5FFC4FF6DF909B7E697DB" pageId="7" pageNumber="64">
6b. Elytral costa less strong and anterior oblique carinae of pronotum not very distinct. Aedeagus in lateral view with apex expanded (
<figureCitation id="BD95CAD4FFD5FFC4FECEF974B30097DC" box="[308,378,1741,1764]" captionStart="FIGURES 11–12" captionStartId="14.[151,264,2000,2025]" captionTargetId="figure@14.[267,1320,166,1980]" captionTargetPageId="14" captionText="FIGURES 11–12. Incoltorrida zahamena, holotype, habitus and aedeagus." httpUri="https://zenodo.org/record/3387376/files/figure.png" pageId="7" pageNumber="64">Fig. 12</figureCitation>
).............................................................................
<taxonomicName id="E2AEADD2FFD5FFC4FADDF977B7E697DB" authorityName="Perkins &amp; Bergsten" authorityYear="2019" box="[1319,1436,1742,1763]" class="Insecta" family="Torridincolidae" genus="Incoltorrida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="zahamena">
<emphasis id="17DA0A43FFD5FFC4FADDF977B7E697DB" box="[1319,1436,1742,1763]" italics="true" pageId="7" pageNumber="64">I. zahamena</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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