treatments-xml/data/03/B1/87/03B187C9FFAEFFF8FF0B88E15BDB70BD.xml

<document id="A3B0CFDCA1FA5CC5DF2601F6453CF54B" ID-CLB-Dataset="24172" ID-DOI="10.11646/zootaxa.4731.2.6" ID-GBIF-Dataset="bd7d6423-9948-45b3-8b1f-1c157c6f42ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="3637887" ID-ZooBank="A1005C73-89E1-42AB-9CFE-BA5B6F85D980" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580886629854" checkinUser="plazi" docAuthor="Sohail, Kamran, Huang, Weijian &amp; Zhang, Yalin" docDate="2020" docId="03B187C9FFAEFFF8FF0B88E15BDB70BD" docLanguage="en" docName="zootaxa.4731.2.6.pdf" docOrigin="Zootaxa 4731 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kodaianellissus gibbusis Kamran, Huang et Zhang 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="273" masterDocId="FF88FFB1FFAFFFFBFF9C8C1F5968731A" masterDocTitle="A new species of the genus Kodaianellissus Wang, Bourgoin &amp; Zhang (Hemiptera: Issidae) with morphological and molecular data" masterLastPageNumber="274" masterPageNumber="270" pageNumber="271" updateTime="1698780494147" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AFC1382F5D73899B894FA606CAD0280F">A new species of the genus Kodaianellissus Wang, Bourgoin &amp; Zhang (Hemiptera: Issidae) with morphological and molecular data</mods:title>
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<mods:namePart id="91D527020A007E45521A338775DD9951">Sohail, Kamran</mods:namePart>
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<mods:namePart id="9E7DEA81681FC4980A122ABA24D060DC">Huang, Weijian</mods:namePart>
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<treatment id="03B187C9FFAEFFF8FF0B88E15BDB70BD" ID-DOI="http://doi.org/10.5281/zenodo.3648595" ID-GBIF-Taxon="161944246" ID-Zenodo-Dep="3648595" LSID="urn:lsid:plazi:treatment:03B187C9FFAEFFF8FF0B88E15BDB70BD" httpUri="http://treatment.plazi.org/id/03B187C9FFAEFFF8FF0B88E15BDB70BD" lastPageId="3" lastPageNumber="273" pageId="1" pageNumber="271">
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<paragraph id="8BA736DFFFAEFFFAFF0B88E15A117603" blockId="1.[151,889,1278,1305]" box="[151,889,1278,1305]" pageId="1" pageNumber="271">
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<taxonomicName id="4C184D5CFFAEFFFAFF0B88E15A667603" ID-CoL="3RBD2" authority="Kamran, Huang et Zhang" authorityName="Kamran, Huang et Zhang" authorityYear="2020" box="[151,782,1278,1305]" class="Insecta" family="Issidae" genus="Kodaianellissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="271" phylum="Arthropoda" rank="species" species="gibbusis" status="sp. nov.">
<emphasis id="B96CEACDFFAEFFFAFF0B88E158A87603" bold="true" box="[151,448,1278,1305]" italics="true" pageId="1" pageNumber="271">Kodaianellissus gibbusis</emphasis>
Kamran, Huang et Zhang
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, 
<taxonomicNameLabel id="A25F57B6FFAEFFFAFC8788E05A117603" box="[795,889,1279,1305]" pageId="1" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3026554FFAEFFF8FF0B895A5C1572ED" lastPageId="3" lastPageNumber="273" pageId="1" pageNumber="271" type="description">
<paragraph id="8BA736DFFFAEFFFAFF0B895A5D0C767B" blockId="1.[151,1437,1349,2024]" box="[151,1124,1349,1377]" pageId="1" pageNumber="271">
<emphasis id="B96CEACDFFAEFFFAFF0B895A5843767A" bold="true" box="[151,299,1349,1376]" pageId="1" pageNumber="271">Description:</emphasis>
Length of male (from apex of head to tip of forewing, N=3) 5.5–6.0 mm.
</paragraph>
<paragraph id="8BA736DFFFAEFFFAFF5B8974588C76EA" blockId="1.[151,1437,1349,2024]" pageId="1" pageNumber="271">Vertex and frons yellowish brown, anterior margin of frons black with small yellow spots, lateral margins yellowish with brown markings. Clypeus yellowish. Eyes greyish with tanned bands. Gena and antennae brown. Pronotum yellowish, mesonotum light brown. Abdomen dark brown dorsally. Tegmina, legs and longitudinal veins brown, cross veins yellowish.</paragraph>
<paragraph id="8BA736DFFFAEFFFAFF5B89E55B4D74F2" blockId="1.[151,1437,1349,2024]" pageId="1" pageNumber="271">
Disc of vertex rectangular, 1.9 times wider than long, centrally and laterally carinate, two rounded markings near lateral margins. Posterior margins concave, anterior margins slightly curved in mid line at apex, lateral margins slightly transverse (
<figureCitation id="13232A5AFFAEFFFAFEE58A5D58BD7546" box="[377,469,1602,1628]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1D</figureCitation>
). Frons slightly globose, broader behind the eyes (
<figureCitation id="13232A5AFFAEFFFAFBBC8A5D5D117546" box="[1056,1145,1602,1628]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1C</figureCitation>
), median carina obvious, lateral margins slightly straight, concavely truncated behind eyes, obliquely narrow near clypeus (
<figureCitation id="13232A5AFFAEFFFAFB4A8A795C44759A" box="[1238,1324,1638,1664]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1C</figureCitation>
). Clypeus with distinct brown transverse strips (
<figureCitation id="13232A5AFFAEFFFAFDA58A955BE775BE" box="[569,655,1674,1700]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1C</figureCitation>
). Large compound eyes (
<figureCitation id="13232A5AFFAEFFFAFC348A955D4475BF" box="[936,1068,1674,1701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Figs. 1C, D</figureCitation>
). Gena smooth, oblique in lateral view (
<figureCitation id="13232A5AFFAEFFFAFF418AB1585C75D2" box="[221,308,1710,1736]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1B</figureCitation>
). Pronotum longer than vertex centrally, anterior margin convex, strongly carinate; posterior margin truncate, central carina yellowish, two black incisions near median carina on the disk (
<figureCitation id="13232A5AFFAEFFFAFBC88ACD5DC575F6" box="[1108,1197,1746,1772]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1D</figureCitation>
). Mesonotum as long as pronotum, two brown spots near lateral margins and near base on each side of the disc, posterior one obviously smaller than apical one. Tegmina, distinctly longer than wide, ScP+R vein forked near base, ScP+RA vein long reaching ⅓ costal margin, RP vein very long, exceeding 1/3 of costa distad at outer margin of tegmina. MP vein bifurcate near basal ¼ long surpassing, MP 
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and MP 
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sinuate at apical margin. Cu forked at apical ⅓, Pcu and A 
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combine at basal half of clavus (
<figureCitation id="13232A5AFFAEFFFAFDAD8B995BE074BA" box="[561,648,1926,1952]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1E</figureCitation>
). Hind wing fully developed tri-lobed. Lobe Pcu-A 
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distinctly longer than wide; A 
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lobe narrower distad at ⅓ of Pcu-A 
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, A 
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lobe simple, longer not forked (
<figureCitation id="13232A5AFFAEFFFAFBD38BB55DCB74DE" box="[1103,1187,1962,1988]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="1" pageNumber="271">Fig. 1F</figureCitation>
). Hind tibiae with two lateral spines, spinal formula 8-7-2.
</paragraph>
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<paragraph id="8BA736DFFFADFFF9FF0B8B8A5D9A74EC" blockId="2.[151,1436,1941,2039]" pageId="2" pageNumber="272">
<emphasis id="B96CEACDFFADFFF9FF0B8B8A583B74B4" bold="true" box="[151,339,1941,1966]" pageId="2" pageNumber="272">FIGURE 1. A–K.</emphasis>
<taxonomicName id="4C184D5CFFADFFF9FEC68B895B3F74B4" authority="Sohail &amp; Huang &amp; Zhang, 2020" authorityName="Sohail &amp; Huang &amp; Zhang" authorityYear="2020" box="[346,599,1942,1966]" class="Insecta" family="Issidae" genus="Kodaianellissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="gibbusis" status="sp. nov.">
<emphasis id="B96CEACDFFADFFF9FEC68B895B3F74B4" box="[346,599,1942,1966]" italics="true" pageId="2" pageNumber="272">Kodaianellissus gibbusis</emphasis>
</taxonomicName>
<emphasis id="B96CEACDFFADFFF9FDC38B895BD874B4" bold="true" box="[607,688,1942,1966]" pageId="2" pageNumber="272">
<taxonomicNameLabel id="A25F57B6FFADFFF9FDC38B895BD874B4" box="[607,688,1942,1966]" pageId="2" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view.
</paragraph>
</caption>
<paragraph id="8BA736DFFFACFFF8FF5B8C865C1572ED" blockId="3.[151,1437,152,935]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8C86581073A9" bold="true" box="[199,376,152,179]" pageId="3" pageNumber="273">Male genitalia.</emphasis>
Anal segment in dorsal view (
<figureCitation id="13232A5AFFACFFF8FD508C865A4A73A9" box="[716,802,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1H</figureCitation>
) rectangular, longer than wide, laterally straight, slightly convex at basal half, anal foramen short, located in basal half (
<figureCitation id="13232A5AFFACFFF8FCC38CA25AD173CD" box="[863,953,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1H</figureCitation>
). Anal tube in lateral view long, convex at posterior ⅓, ventral margins slightly prominent (
<figureCitation id="13232A5AFFACFFF8FD5F8CFE5A7673E1" box="[707,798,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1G</figureCitation>
). Pygofer in lateral view, long and slender with parallel anterior and posterior margins (
<figureCitation id="13232A5AFFACFFF8FE618D1A5B3F7205" box="[509,599,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1G</figureCitation>
). Genital styles rectangular, widest at basal ⅓, caudoventrally rectangular slightly curved near base of capitilum at posterior margin, dorsolaterally provided with hump-like processes near base of capitulum (
<figureCitation id="13232A5AFFACFFF8FEF68D5258D0727D" box="[362,440,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1J</figureCitation>
). Capitilum very long, tubular, apically sclerotized slightly bent directed anteriorly, posterior margin with lobe-like process (
<figureCitation id="13232A5AFFACFFF8FDBE8D6E5B187291" box="[546,624,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1J</figureCitation>
). Aedeagus in lateral view elevated at both sides, divided into two lobes, dorsolateral lobe slightly membranous in apical portion, centrally convex. Sharp sclerotized spines directed dorsad on each side of ventral apical part (
<figureCitation id="13232A5AFFACFFF8FD878DA65B1B72C9" box="[539,627,441,467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1K</figureCitation>
). In ventral view, periandrium is long U-shaped, somewhat symmetrical, dorsolateral margins twisted in distal half, pair of long hooks arising in distal half directed downwards (
<figureCitation id="13232A5AFFACFFF8FABC8DC25C0472ED" box="[1312,1388,477,503]" captionStart="FIGURE 1" captionStartId="2.[151,250,1941,1966]" captionTargetBox="[202,1403,174,1909]" captionTargetId="figure@2.[175,1412,166,1922]" captionTargetPageId="2" captionText="FIGURE 1. A–K. Kodaianellissus gibbusis sp. nov. A. adult, dorsal view; B. adult, right lateral view; C. frons, ventral view; D. vertex, pronotum &amp; mesonotum dorsal view; E, F. fore and hind wings left lateral view; G. pygofer, lateral view; H. anal segment, dorsal view; I. aedeagus, ventral view; J. genital style, lateral view; K. phallic complex, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3637889" httpUri="https://zenodo.org/record/3637889/files/figure.png" pageId="3" pageNumber="273">Fig. 1I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3026554FFACFFF8FF5B8E1E58CD7179" pageId="3" pageNumber="273" type="materials_examined">
<paragraph id="8BA736DFFFACFFF8FF5B8E1E5D5B7125" blockId="3.[151,1437,152,935]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8E1E58EB7101" bold="true" box="[199,387,512,539]" pageId="3" pageNumber="273">Type materials:</emphasis>
<materialsCitation id="3B703C82FFACFFF8FE168E1E5B1D7125" ID-GBIF-Occurrence="2563048133" collectingDate="2018-07-14" collectorName="Azad Jammu &amp; Kashmir &amp; Kamran Sohail." country="Pakistan" elevation="1628" latitude="33.858936" location="Rawlakot" longLatPrecision="1" longitude="73.759705" pageId="3" pageNumber="273" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A3887DFFACFFF8FE168E1E589F7101" box="[394,503,513,539]" pageId="3" pageNumber="273" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D1EFD56FFACFFF8FD9E8E1E5B7F7101" box="[514,535,513,539]" pageId="3" pageNumber="273" type="male">♂</specimenCount>
<location id="8EC76004FFACFFF8FD828E1E5BE47101" LSID="urn:lsid:plazi:treatment:03B187C9FFAEFFF8FF0B88E15BDB70BD:8EC76004FFACFFF8FD828E1E5BE47101" box="[542,652,513,539]" country="Pakistan" latitude="33.858936" longLatPrecision="1" longitude="73.759705" name="Rawlakot" pageId="3" pageNumber="273">Rawlakot</location>
, 
<collectorName id="26ED5309FFACFFF8FD0A8E1E5A427101" box="[662,810,513,539]" pageId="3" pageNumber="273">Azad Jammu</collectorName>
&amp; 
<collectorName id="26ED5309FFACFFF8FCD38E1E5AD97101" box="[847,945,513,539]" pageId="3" pageNumber="273">Kashmir</collectorName>
, 
<collectingCountry id="F30F764FFFACFFF8FC218E1E5D777101" box="[957,1055,513,539]" name="Pakistan" pageId="3" pageNumber="273">Pakistan</collectingCountry>
, 
<geoCoordinate id="EE2C5018FFACFFF8FBB68E1F5DB17101" box="[1066,1241,512,539]" degrees="33" direction="north" minutes="51" orientation="latitude" pageId="3" pageNumber="273" precision="1" seconds="32.18" value="33.858936">33°51’32.18”N</geoCoordinate>
, 
<geoCoordinate id="EE2C5018FFACFFF8FB798E1E5CF07101" box="[1253,1432,512,539]" degrees="73" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="273" precision="1" seconds="34.93" value="73.759705">73° 45’34.93”E</geoCoordinate>
, 
<quantity id="4CE09B3AFFACFFF8FF0B8E3A59867125" box="[151,238,549,575]" metricMagnitude="3" metricUnit="m" metricValue="1.628" pageId="3" pageNumber="273" unit="m" value="1628.0">
<elevation id="0035D1ECFFACFFF8FF0B8E3A59867125" box="[151,238,549,575]" metricMagnitude="3" metricUnit="m" metricValue="1.628" pageId="3" pageNumber="273" unit="m" value="1628.0">1628 m</elevation>
</quantity>
, 
<date id="FFA6101FFFACFFF8FF668E3A58E97125" box="[250,385,549,575]" pageId="3" pageNumber="273" value="2018-07-14">
<collectingDate id="EFE2E9F7FFACFFF8FF668E3A58E97125" box="[250,385,549,575]" pageId="3" pageNumber="273" value="2018-07-14">14-vii-2018</collectingDate>
</date>
, coll. 
<collectorName id="26ED5309FFACFFF8FE588E3A5B1D7125" box="[452,629,549,575]" pageId="3" pageNumber="273">Kamran Sohail.</collectorName>
</materialsCitation>
<materialsCitation id="3B703C82FFACFFF8FDE78E3A5D477125" ID-GBIF-Occurrence="2563048134" box="[635,1071,549,575]" collectingDate="2018-07-14" collectorName="Azad Jammu &amp; Kashmir &amp; Kamran Sohail." country="Pakistan" elevation="1628" latitude="33.858936" location="Rawlakot" longLatPrecision="1" longitude="73.759705" pageId="3" pageNumber="273" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54A3887DFFACFFF8FDE78E3A5B807125" box="[635,744,549,575]" pageId="3" pageNumber="273" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D1EFD56FFACFFF8FD738E3A5A427125" box="[751,810,549,575]" pageId="3" pageNumber="273" type="male">2♂♂</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BA736DFFFACFFF8FF5B8E5658CD7179" blockId="3.[151,1437,152,935]" box="[199,421,585,611]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8E5658407179" bold="true" box="[199,296,585,611]" pageId="3" pageNumber="273">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3026554FFACFFF8FF5B8E72583F71B1" pageId="3" pageNumber="273" type="etymology">
<paragraph id="8BA736DFFFACFFF8FF5B8E72583F71B1" blockId="3.[151,1437,152,935]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8E72583A719D" bold="true" box="[199,338,621,647]" pageId="3" pageNumber="273">Etymology:</emphasis>
Latin word “ 
<emphasis id="B96CEACDFFACFFF8FE6D8E725B54719D" box="[497,572,621,647]" italics="true" pageId="3" pageNumber="273">gibbus</emphasis>
” means hump, referring to hump-like processes near base of capitilum on dorsolateral side.
</paragraph>
</subSubSection>
<subSubSection id="C3026554FFACFFF8FF5B8EAA58BE71D5" box="[199,470,693,719]" pageId="3" pageNumber="273" type="biology_ecology">
<paragraph id="8BA736DFFFACFFF8FF5B8EAA58BE71D5" blockId="3.[151,1437,152,935]" box="[199,470,693,719]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8EAA583171D5" bold="true" box="[199,345,693,719]" pageId="3" pageNumber="273">Host plants:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3026554FFACFFF8FF5B8EC75863700D" pageId="3" pageNumber="273" type="diagnosis">
<paragraph id="8BA736DFFFACFFF8FF5B8EC75863700D" blockId="3.[151,1437,152,935]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8EC7582C71E9" bold="true" box="[199,324,728,755]" pageId="3" pageNumber="273">Diagnosis:</emphasis>
<taxonomicName id="4C184D5CFFACFFF8FED68EC65B0871E9" authority="Sohail &amp; Huang &amp; Zhang, 2020" authorityName="Sohail &amp; Huang &amp; Zhang" authorityYear="2020" box="[330,608,729,755]" class="Insecta" family="Issidae" genus="Kodaianellissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="gibbusis" status="sp. nov.">
<emphasis id="B96CEACDFFACFFF8FED68EC65B0871E9" box="[330,608,729,755]" italics="true" pageId="3" pageNumber="273">Kodaianellissus gibbusis</emphasis>
</taxonomicName>
<emphasis id="B96CEACDFFACFFF8FDFB8EC65BA871E9" bold="true" box="[615,704,729,755]" pageId="3" pageNumber="273">
<taxonomicNameLabel id="A25F57B6FFACFFF8FDFB8EC65BA871E9" box="[615,704,729,755]" pageId="3" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from 
<taxonomicName id="4C184D5CFFACFFF8FC728EC55DED71E9" authorityName="Wang, Bourgoin &amp; Zhang" authorityYear="2017" box="[1006,1157,730,755]" class="Insecta" family="Issidae" genus="Kodaianellissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="intorqueus">
<emphasis id="B96CEACDFFACFFF8FC728EC55DED71E9" box="[1006,1157,730,755]" italics="true" pageId="3" pageNumber="273">K. intorqueus</emphasis>
</taxonomicName>
by characters mentioned in the key.
</paragraph>
</subSubSection>
<subSubSection id="C3026554FFACFFF8FF5B8F3E5BDB70BD" pageId="3" pageNumber="273" type="discussion">
<paragraph id="8BA736DFFFACFFF8FF5B8F3E5BDB70BD" blockId="3.[151,1437,152,935]" pageId="3" pageNumber="273">
<emphasis id="B96CEACDFFACFFF8FF5B8F3E58557021" bold="true" box="[199,317,801,827]" pageId="3" pageNumber="273">Remarks:</emphasis>
Phylogenetic analyses of 5 species of 
<taxonomicName id="4C184D5CFFACFFF8FD6D8F3E5ADA7021" authorityName="Melichar" authorityYear="1906" box="[753,946,801,827]" class="Insecta" family="Issidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="subFamily" subFamily="Hemisphaeriinae">Hemisphaeriinae</taxonomicName>
, including two out-groups (
<taxonomicName id="4C184D5CFFACFFF8FB778F3E59837045" authorityName="Yang" authorityYear="1994" class="Insecta" family="Issidae" genus="Gelastyrella" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="litaoensis">
<emphasis id="B96CEACDFFACFFF8FB778F3E59837045" italics="true" pageId="3" pageNumber="273">Gelastyrella litaoensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C184D5CFFACFFF8FEB88F5A58907045" authorityName="Gnezdilov &amp; Bourgoin" authorityYear="2014" box="[292,504,837,863]" class="Insecta" family="Issidae" genus="Darwallia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="barbata">
<emphasis id="B96CEACDFFACFFF8FEB88F5A58907045" box="[292,504,837,863]" italics="true" pageId="3" pageNumber="273">Darwallia barbata</emphasis>
</taxonomicName>
), based on ML and BI analyses of nucleotide sequence data of two genes, yielded almost same topologies with strong support (
<figureCitation id="13232A5AFFACFFF8FD168F765BA77099" box="[650,719,873,899]" captionStart="FIGURE 2" captionStartId="3.[151,250,1570,1595]" captionTargetBox="[154,1427,955,1548]" captionTargetId="figure@3.[151,1435,949,1550]" captionTargetPageId="3" captionText="FIGURE 2. Phylogenetic tree based on COI and cytb sequences with Gelastyrella litaoensis (Hemisphaeriinae, Parahiraciini) and Darwallia barbata (Hemisphaeriinae, Sarimini) as outgroup to test the position of Kodaianellissus gibbusis sp. nov. in the classification with the genus Kodaianella. At each node, values indicate Bayesian PP and ML support: Bayesian posterior probability (PP) / ultrafast bootstrap (UFB) values." figureDoi="http://doi.org/10.5281/zenodo.3637891" httpUri="https://zenodo.org/record/3637891/files/figure.png" pageId="3" pageNumber="273">Fig. 2</figureCitation>
). The molecular analyses (
<figureCitation id="13232A5AFFACFFF8FC648F765D527099" box="[1016,1082,873,899]" captionStart="FIGURE 2" captionStartId="3.[151,250,1570,1595]" captionTargetBox="[154,1427,955,1548]" captionTargetId="figure@3.[151,1435,949,1550]" captionTargetPageId="3" captionText="FIGURE 2. Phylogenetic tree based on COI and cytb sequences with Gelastyrella litaoensis (Hemisphaeriinae, Parahiraciini) and Darwallia barbata (Hemisphaeriinae, Sarimini) as outgroup to test the position of Kodaianellissus gibbusis sp. nov. in the classification with the genus Kodaianella. At each node, values indicate Bayesian PP and ML support: Bayesian posterior probability (PP) / ultrafast bootstrap (UFB) values." figureDoi="http://doi.org/10.5281/zenodo.3637891" httpUri="https://zenodo.org/record/3637891/files/figure.png" pageId="3" pageNumber="273">Fig. 2</figureCitation>
) with morphological characters indicates its placement into the 
<taxonomicName id="4C184D5CFFACFFF8FE658F925BC470BD" box="[505,684,909,935]" class="Insecta" family="Issidae" genus="Kodaianellissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEACDFFACFFF8FE658F925BC470BD" box="[505,684,909,935]" italics="true" pageId="3" pageNumber="273">Kodaianellissus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>