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<document id="B4D767FAC58F6983B295A2B6F461E8DD" ID="10.11646/zootaxa.4232.2.2SLASH10184" ID-CLB-Dataset="36533" ID-DOI="10.11646/zootaxa.4232.2.2" ID-GBIF-Dataset="45f9becd-c9ea-4670-aca5-270b4c562996" ID-ISSN="1175-5326" ID-Zenodo-Dep="292888" ID-ZooBank="3108C187-8419-46C5-8E69-B731614B9D79" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487232990106" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid &amp; Rafi, Muhammad Ather" docDate="2017" docId="0385B42F9B75FFD5FF2BF9F091BFFD53" docLanguage="en" docName="zootaxa.4232.2.2SLASH10184.pdf" docOrigin="Zootaxa 4232 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Brachinini" docType="key" docVersion="7" lastPageNumber="175" masterDocId="FFBCCC579B74FFD7FFBCFFE09424FFF1" masterDocTitle="Faunistic studies of the tribe Brachinini (Carabidae: Coleoptera) from northern Pakistan" masterLastPageNumber="184" masterPageNumber="173" pageNumber="174" updateTime="1698436961878" updateUser="ExternalLinkService">
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<mods:title id="C91B580BD8A7A0890D92856A81228781">Faunistic studies of the tribe Brachinini (Carabidae: Coleoptera) from northern Pakistan</mods:title>
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<mods:namePart id="2FA723012C99BC6EADE67A4A85E5DC0F">Ullah, Mishkat</mods:namePart>
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<mods:namePart id="24CFAC6018517FB87F727BA40526E69C">Naeem, Muhammad</mods:namePart>
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<mods:namePart id="D5942BE52F6FA4CF70BF3C8DEC0349CD">Mahmood, Khalid</mods:namePart>
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<mods:namePart id="6DC4583816D6A2059743FE3CF094F3FB">Rafi, Muhammad Ather</mods:namePart>
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<mods:title id="906D1B765F6E1A297CF5C33D16D2BF6B">Zootaxa</mods:title>
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<mods:date id="EF8685B56EAB50741C8CE7C5E260E62C">2017</mods:date>
<mods:detail id="5E2E6980987A176045161CC636D7847D" type="volume">
<mods:number id="F15254BAE9E4A2279EDD5F934E71B78B">4232</mods:number>
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<mods:identifier id="43B2BFF6A32AD64924FB3E969F9921A1" type="CLB-Dataset">36533</mods:identifier>
<mods:identifier id="0FF852F9EB87E476993FF1ACADC373B0" type="DOI">10.11646/zootaxa.4232.2.2</mods:identifier>
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<treatment id="0385B42F9B75FFD5FF2BF9F091BFFD53" ID-DOI="http://doi.org/10.5281/zenodo.5999116" ID-GBIF-Taxon="127723673" ID-Zenodo-Dep="5999116" LSID="urn:lsid:plazi:treatment:0385B42F9B75FFD5FF2BF9F091BFFD53" httpUri="http://treatment.plazi.org/id/0385B42F9B75FFD5FF2BF9F091BFFD53" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174">
<subSubSection id="C33656B29B75FFD6FF2BF9F0900CF9DB" box="[151,1064,1552,1578]" pageId="1" pageNumber="174" type="nomenclature">
<paragraph id="8B9305399B75FFD6FF2BF9F0900CF9DB" blockId="1.[151,1064,1552,1578]" box="[151,1064,1552,1578]" pageId="1" pageNumber="174">
<heading id="D0DBB2559B75FFD6FF2BF9F0900CF9DB" bold="true" box="[151,1064,1552,1578]" fontSize="11" level="1" pageId="1" pageNumber="174" reason="1">
<emphasis id="B958D92B9B75FFD6FF2BF9F0900CF9DB" bold="true" box="[151,1064,1552,1578]" pageId="1" pageNumber="174">
Key to the genera and species of tribe
<taxonomicName id="4C2C7EBA9B75FFD6FDCEF9F096D3F9DB" ID-CoL="8KTSZ" box="[626,759,1552,1578]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="tribe" tribe="Brachinini">Brachinini</taxonomicName>
from northern Pakistan
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33656B29B75FFD5FF2BF9B591BFFD53" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174" type="key">
<keyStep id="30D81D9C9B75FFD6FF2BF9B591B8F932" pageId="1" pageNumber="174">
<paragraph id="8B9305399B75FFD6FF2BF9B591BFF978" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF9B591BFF978" pageId="1" pageNumber="174">1 Mandible with scrobe; pronotum without longitudinal ridges on disc; protibia upper spur present; mesocoxae not widely separated; metacoxae conjunct-separate...................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B9305399B75FFD6FF2BF97191B8F932" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF97191B8F932" pageId="1" pageNumber="174">
- Mandible without scrobe; pronotum with two longitudinal ridges on disc; protibia upper spur absent; mesocoxae conjunt-separate; metacoxae widely separated..............................................
<taxonomicName id="4C2C7EBA9B75FFD6FBEFF94E91A7F932" ID-CoL="8KVTZ" authority="Fischer von Waldheim ." authorityName="Fischer von Waldheim ." box="[1107,1411,1709,1731]" class="Insecta" family="Carabidae" genus="Mastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B958D92B9B75FFD6FBEFF94E90BDF932" box="[1107,1177,1710,1731]" italics="true" pageId="1" pageNumber="174">Mastax</emphasis>
Fischer von Waldheim.
</taxonomicName>
..3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B75FFD6FF2BF92B91B8F8EA" pageId="1" pageNumber="174">
<paragraph id="8B9305399B75FFD6FF2BF92B91BFF910" blockId="1.[151,1436,1621,2025]" box="[151,1435,1739,1761]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF92B91BFF910" box="[151,1435,1739,1761]" pageId="1" pageNumber="174">
2 Mandibular scrobe multisetose; elytral striae flat not well developed; apical elytral membrane present.
<taxonomicName id="4C2C7EBA9B75FFD6FB6DF92B91A6F910" ID-CoL="8KVCP" authority="Weber ." authorityName="Weber ." box="[1233,1410,1739,1761]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="genus">
<emphasis id="B958D92B9B75FFD6FB6DF92B9117F910" box="[1233,1331,1739,1761]" italics="true" pageId="1" pageNumber="174">Brachinus</emphasis>
Weber.
</taxonomicName>
..4
</keyLead>
</paragraph>
<paragraph id="8B9305399B75FFD6FF2BF90991B8F8EA" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF90991B8F8EA" pageId="1" pageNumber="174">
- Mandibular scrobe unisetose; elytral striae carinate; carinae rounded; well defined strongly contrasting with depressions.......................................................................
<taxonomicName id="4C2C7EBA9B75FFD6FBBAF8E591A6F8EA" ID-CoL="8LTW4" authority="Maindron ." authorityName="Maindron ." box="[1030,1410,1797,1819]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="subGenus" subGenus="Stenaptinus">
<emphasis id="B958D92B9B75FFD6FBBAF8E59137F8EA" box="[1030,1299,1797,1819]" italics="true" pageId="1" pageNumber="174">Pheropsophus (Stenaptinus)</emphasis>
Maindron.
</taxonomicName>
..6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B75FFD6FF2BF8C391BFF85F" pageId="1" pageNumber="174">
<paragraph id="8B9305399B75FFD6FF2BF8C391BFF8A7" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF8C391BFF8A7" pageId="1" pageNumber="174">
3 Elytra black with reddish brown dumbbell shaped band along middle suture; two off-white transverse bands which could not touch each other as a result of central dumbbell shape mark.......................
<taxonomicName id="4C2C7EBA9B75FFD6FB91F8A191BFF8A7" ID-CoL="3YCPJ" authority="Fischer von Waldheim" authorityName="Fischer von Waldheim" box="[1069,1435,1856,1878]" class="Insecta" family="Carabidae" genus="Mastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="B958D92B9B75FFD6FB91F8A19098F8A7" box="[1069,1212,1856,1878]" italics="true" pageId="1" pageNumber="174">Mastax histrio</emphasis>
Fischer von Waldheim
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9305399B75FFD6FF2BF8BE91BFF85F" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF8BE91BFF85F" pageId="1" pageNumber="174">
- Elytra matt black; humeral band, light brown quarter a circle shaped end as with square shape mark near epiplura; one white mark near side edge of elytra and one other light brown mark near suture before elytral apical band.....................................................................................................
<taxonomicName id="4C2C7EBA9B75FFD6FB25F87991BFF85F" authority="Jedlicka" authorityName="Jedlicka" box="[1177,1435,1944,1966]" class="Insecta" family="Carabidae" genus="Mastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="174" phylum="Arthropoda" rank="species" species="pakistana">
<emphasis id="B958D92B9B75FFD6FB25F8799163F85F" box="[1177,1351,1944,1966]" italics="true" pageId="1" pageNumber="174">Mastax pakistana</emphasis>
Jedlička
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B75FFD5FF2BF85591BFFF3B" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174">
<paragraph id="8B9305399B75FFD6FF2BF85591B3F818" blockId="1.[151,1436,1621,2025]" pageId="1" pageNumber="174">
<keyLead id="30DDA60C9B75FFD6FF2BF85591B3F818" pageId="1" pageNumber="174">4 Elytra spotted or variegated color pattern (yellow and black); two subhumeral round marks narrowly extend to the shoulders and a subapical transverse band with three posterior spots merged to form a continuous sinuated band...................</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF5FFF7891BFFF3B" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
..................................................................................
<taxonomicName id="4C2C7EBA9B76FFD5FB02FF7891B8FF5D" ID-CoL="MSWH" authority="Hope" authorityName="Hope" box="[1214,1436,151,173]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="pictus" subGenus="Aploa">
<emphasis id="B958D92B9B76FFD5FB02FF789145FF5C" box="[1214,1377,151,173]" italics="true" pageId="2" pageNumber="175">B. (Aploa) pictus</emphasis>
Hope
</taxonomicName>
<keyLead id="30DDA60C9B76FFD5FF2BFF5591BFFF3B" box="[151,1435,180,202]" pageId="2" pageNumber="175">- Elytra uniformly colored with metallic tinge; elytral striae contrasting in color with disc.............................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B76FFD5FF2BFF3191B9FECE" pageId="2" pageNumber="175">
<paragraph id="8B9305399B76FFD5FF2BFF3191BFFEF4" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFF3191BFFEF4" pageId="2" pageNumber="175">
5 Antennal segment III subequal to segment I and II combined; elytral apical membrane at apex with a row of long setae (longer than the width of apical membrane)..................................
<taxonomicName id="4C2C7EBA9B76FFD5FC72FF1091BFFEF4" ID-CoL="MSXF" authority="Fischer von Waldheim" authorityName="Fischer von Waldheim" box="[974,1435,239,261]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="ejaculans" subGenus="Brachinus">
<emphasis id="B958D92B9B76FFD5FC72FF109099FEF4" box="[974,1213,239,261]" italics="true" pageId="2" pageNumber="175">B. (Brachinus) ejaculans</emphasis>
Fischer von Waldheim
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF2BFEED91B9FECE" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFEED91B9FECE" pageId="2" pageNumber="175">
- Antennal segment III longer than segment I and II combined; elytral apical membrane apex without row of long setae....................................................................
<taxonomicName id="4C2C7EBA9B76FFD5FC75FECA910DFECE" ID-CoL="MSYT" authority="Reitter" authorityName="Reitter" box="[969,1321,297,319]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="pallidipes" status="comb. nov." subGenus="Neobrachinus">
<emphasis id="B958D92B9B76FFD5FC75FECA90FBFECE" box="[969,1247,297,319]" italics="true" pageId="2" pageNumber="175">B. (Neobrachinus) pallidipes</emphasis>
Reitter
</taxonomicName>
,
<taxonomicNameLabel id="A26B64509B76FFD5FA8EFEC991B9FECE" box="[1330,1437,297,319]" pageId="2" pageNumber="175">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B76FFD5FF2BFEA791BFFE8B" pageId="2" pageNumber="175">
<paragraph id="8B9305399B76FFD5FF2BFEA791B8FEAD" blockId="2.[151,1437,151,674]" box="[151,1436,327,349]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFEA791B8FEAD" box="[151,1436,327,349]" pageId="2" pageNumber="175">6 Elytra with humeral marks; elytral epipleura without fine tubercles..............................................7</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF2BFE8591BFFE8B" blockId="2.[151,1437,151,674]" box="[151,1435,356,378]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFE8591BFFE8B" box="[151,1435,356,378]" pageId="2" pageNumber="175">
- Elytral without yellow humeral spot and epipleura with fine tubercles............
<taxonomicName id="4C2C7EBA9B76FFD5FBB5FE8591BFFE8B" ID-CoL="8MHWJ" authority="Dejean" authorityName="Dejean" box="[1033,1435,356,378]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="subSpecies" species="hilaris" subGenus="Stenaptinus" subSpecies="sobrinus">
<emphasis id="B958D92B9B76FFD5FBB5FE859166FE8B" box="[1033,1346,356,378]" italics="true" pageId="2" pageNumber="175">P. (Stenaptinus) hilaris sobrinus</emphasis>
(Dejean)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B76FFD5FF2BFE6291BFFE44" pageId="2" pageNumber="175">
<paragraph id="8B9305399B76FFD5FF2BFE6291BFFE66" blockId="2.[151,1437,151,674]" box="[151,1435,385,407]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFE6291BFFE66" box="[151,1435,385,407]" pageId="2" pageNumber="175">7 Frons with black mark................................................................................. 8</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF2BFE7F91BFFE44" blockId="2.[151,1437,151,674]" box="[151,1435,415,437]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFE7F91BFFE44" box="[151,1435,415,437]" pageId="2" pageNumber="175">- Frons without black mark.............................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B76FFD5FF2BFE5C91B8FDDB" pageId="2" pageNumber="175">
<paragraph id="8B9305399B76FFD5FF2BFE5C91BFFE1E" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFE5C91BFFE1E" pageId="2" pageNumber="175">
8 Head yellowish; frons with black inverted “Y” shaped mark; pronotum black with two longitudinal yellow stripes.........................................................................
<taxonomicName id="4C2C7EBA9B76FFD5FC54FE3A91BFFE1E" ID-CoL="4G395" authority="Schmidt-Goebel" authorityName="Schmidt-Goebel" box="[1000,1435,473,495]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="consularis" subGenus="Stenaptinus">
<emphasis id="B958D92B9B76FFD5FC54FE3A90CCFE1E" box="[1000,1256,473,495]" italics="true" pageId="2" pageNumber="175">P. (Stenaptinus) consularis</emphasis>
(Schmidt-Goebel)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF2BFE1791B8FDDB" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFE1791B8FDDB" pageId="2" pageNumber="175">
- Head reddish brown; frons with dark brown longitudinal stripe; pronotum lateral bead black...................................................................................................
<taxonomicName id="4C2C7EBA9B76FFD5FBF9FDF591B8FDDB" ID-CoL="778J7" authority="Chaudoir" authorityName="Chaudoir" box="[1093,1436,532,554]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="lineifrons" subGenus="Stenaptinus">
<emphasis id="B958D92B9B76FFD5FBF9FDF59119FDDB" box="[1093,1341,532,554]" italics="true" pageId="2" pageNumber="175">P. (Stenaptinus) lineifrons</emphasis>
Chaudoir
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D81D9C9B76FFD5FF2BFDD491BFFD53" pageId="2" pageNumber="175">
<paragraph id="8B9305399B76FFD5FF2BFDD491B8FD96" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFDD491B8FD96" pageId="2" pageNumber="175">
9 Elytral humeral mark longitudinal; median transverse band strongly serrated at edges and ends at 8th stria; elytral epipleuron black...................................................................
<taxonomicName id="4C2C7EBA9B76FFD5FB85FDB291B8FD96" ID-CoL="6V8J8" authority="Chaudoir" authorityName="Chaudoir" box="[1081,1436,593,615]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="lissoderus" subGenus="Stenaptinus">
<emphasis id="B958D92B9B76FFD5FB85FDB29118FD96" box="[1081,1340,593,615]" italics="true" pageId="2" pageNumber="175">P. (Stenaptinus) lissoderus</emphasis>
Chaudoir
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9305399B76FFD5FF2BFD8F91BFFD53" blockId="2.[151,1437,151,674]" pageId="2" pageNumber="175">
<keyLead id="30DDA60C9B76FFD5FF2BFD8F91BFFD53" pageId="2" pageNumber="175">
- Elytral humeral mark round; median transverse band round near elytral suture and narrow towards epipleuron where it ends; epipleuron black except where median transverse band touches.......................
<taxonomicName id="4C2C7EBA9B76FFD5FBE7FD6D91BFFD53" ID-CoL="4G38V" authority="Dejean" authorityName="Dejean" box="[1115,1435,652,674]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="175" phylum="Arthropoda" rank="species" species="catoirei" subGenus="Stenaptinus">
<emphasis id="B958D92B9B76FFD5FBE7FD6D9166FD53" box="[1115,1346,652,674]" italics="true" pageId="2" pageNumber="175">P. (Stenaptinus) catoirei</emphasis>
(Dejean)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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