127 lines
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127 lines
16 KiB
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<document id="28615C400B19CBC6634736FC41A9369F" ID-CLB-Dataset="2406" ID-DOI="10.11646/zootaxa.5093.2.2" ID-GBIF-Dataset="ffbad455-ad57-4fa5-8efa-d18cc2d12e40" ID-ISSN="1175-5326" ID-Zenodo-Dep="5905141" ID-ZooBank="31913C5B-A8D5-4A30-8D9E-4C7C9E953E1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643182084586" checkinUser="plazi" docAuthor="Antić, Dragan & Mauriès, Jean-Paul" docDate="2022" docId="03E387B4FFC3CD0856C40817FAD0FA73" docLanguage="en" docName="zootaxa.5093.2.2.pdf" docOrigin="Zootaxa 5093 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guipuzcosomatidae Antic & Mauries 2022, new family" docType="treatment" docVersion="7" lastPageNumber="146" masterDocId="FFDAFFCCFFC7CD0C56530A51FFA7FFC8" masterDocTitle="Two new species of the previously monospecific genus Guipuzcosoma Vicente & Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)" masterLastPageNumber="168" masterPageNumber="142" pageNumber="146" updateTime="1699304320109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="14440890EE5E9241C1894987D21C9678" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="452DEECE70374163FAD65A34B05BB462">
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<mods:title id="301FFF33856EEE8B3DB6473305AF8FAA">Two new species of the previously monospecific genus Guipuzcosoma Vicente & Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)</mods:title>
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</mods:titleInfo>
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<mods:name id="0ABCA2D5D59FA843A90572D260D9FCFC" type="personal">
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<mods:roleTerm id="1C7E15660CF39D6B10CFC272963E6900">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A72D0E98EBC61996DD8CCBEECBEF6310">Antić, Dragan</mods:namePart>
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<mods:affiliation id="733B72A2A42882818680C94E789D1BE7">University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia</mods:affiliation>
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</mods:name>
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<mods:name id="823A9E5BA71D401AD2D1426DFD53FB17" type="personal">
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<mods:role id="354D0A85039F85F133CBE90675D3AAEF">
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<mods:roleTerm id="ADBDF05EDB1C20AE6AC16E1CF9B119E3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A87F28B7A7D36009DB88D7DB0D48A3E">Mauriès, Jean-Paul</mods:namePart>
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<mods:nameIdentifier id="F9026528E276FE753EE87DCA21478421" type="ORCID">0000-0002-0575-6937</mods:nameIdentifier>
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<mods:affiliation id="4CB99121524443D6CD8088D228863526">Muséum National d’Histoire Naturelle, Dpt. Origines & Evolution, Section Arthropodes, 61 rue Buffon F- 75005 Paris, France mauries. jean-paul @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 0575 - 6937</mods:affiliation>
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<mods:nameIdentifier id="5922F82AF8135A0720528D2D66D2281B" type="email">mauries.jean-paul@wanadoo.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="F5D82FEF42B85B7C9D418F5A7F1846B2">text</mods:typeOfResource>
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<mods:titleInfo id="ECCFF1F5DF0135CCE9A500C68ADCF755">
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<mods:title id="4CABD30550E42EABD44E0B9EBB0AF10F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B1F3CA7DD5144958A9606B4719AFB4D5">
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<mods:date id="0239522C3D4996828DDD19EF1D85BA8D">2022</mods:date>
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<mods:detail id="2C1151DD1860BF422B9775D68FDBC96E" type="pubDate">
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<mods:number id="275D8EF4F5EE3CA13FB8F63BBD9BA249">2022-01-26</mods:number>
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</mods:detail>
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<mods:detail id="D732951250BE9109FE2E1E884B80471E" type="volume">
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<mods:number id="0E679198DDE7020E58FBAD5936809AF0">5093</mods:number>
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</mods:detail>
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<mods:detail id="55F23D2D26A7F44B3CF579F6F585AC31" type="issue">
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<mods:number id="FABB77994349D61ADF6B505266033B46">2</mods:number>
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<mods:identifier id="4BA6CFE597DF5AF05A6F0731D2508BA8" type="DOI">10.11646/zootaxa.5093.2.2</mods:identifier>
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<mods:identifier id="157E30105255FF18F689FD2722B2A615" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03E387B4FFC3CD0856C40817FAD0FA73" ID-DOI="http://doi.org/10.5281/zenodo.5906896" ID-GBIF-Taxon="192626921" ID-Zenodo-Dep="5906896" LSID="urn:lsid:plazi:treatment:03E387B4FFC3CD0856C40817FAD0FA73" httpUri="http://treatment.plazi.org/id/03E387B4FFC3CD0856C40817FAD0FA73" lastPageNumber="146" pageId="4" pageNumber="146">
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<subSubSection id="C3506529FFC3CD0856C40817FCC7FDA8" box="[151,864,582,609]" pageId="4" pageNumber="146" type="nomenclature">
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<paragraph id="8BF536A2FFC3CD0856C40817FCC7FDA8" blockId="4.[151,864,582,609]" box="[151,864,582,609]" pageId="4" pageNumber="146">
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<heading id="D0BD81CEFFC3CD0856C40817FCC7FDA8" bold="true" box="[151,864,582,609]" fontSize="11" level="1" pageId="4" pageNumber="146" reason="1">
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<emphasis id="B93EEAB0FFC3CD0856C40817FCC7FDA8" bold="true" box="[151,864,582,609]" pageId="4" pageNumber="146">
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Family
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<taxonomicName id="4C4A4D21FFC3CD0856A40817FD6EFDA9" ID-CoL="b55a17c1-98c6-4541-8339-faca4d732973" authority="Antic & Mauries" authorityName="Antic & Mauries" authorityYear="2022" box="[247,713,582,609]" family="Guipuzcosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" rank="family" status="fam. nov.">Guipuzcosomatidae Antić & Mauriès</taxonomicName>
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,
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<taxonomicNameLabel id="A20D57CBFFC3CD0854850816FCC7FDA8" box="[726,864,582,609]" pageId="4" pageNumber="146" rank="family">new family</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3506529FFC3CD0856C408DAFCF2FD6C" box="[151,853,651,676]" pageId="4" pageNumber="146" type="type_taxon">
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<paragraph id="8BF536A2FFC3CD0856C408DAFCF2FD6C" blockId="4.[151,853,651,676]" box="[151,853,651,676]" pageId="4" pageNumber="146">
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<emphasis id="B93EEAB0FFC3CD0856C408DAFEBEFD6B" bold="true" box="[151,281,651,675]" pageId="4" pageNumber="146">
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<typeStatus id="54F18800FFC3CD0856C408DAFF6BFD6B" box="[151,204,651,675]" pageId="4" pageNumber="146">Type</typeStatus>
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genus:
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</emphasis>
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<taxonomicName id="4C4A4D21FFC3CD08574C08DDFD19FD6B" authority="Vicente & Mauries, 1980" authorityName=", Vicente & Mauries" authorityYear="1980" box="[287,702,651,675]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" phylum="Arthropoda" rank="genus">
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<emphasis id="B93EEAB0FFC3CD08574C08DDFE11FD6B" box="[287,438,652,675]" italics="true" pageId="4" pageNumber="146">Guipuzcosoma</emphasis>
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<bibRefCitation id="EFDB4B53FFC3CD0857EF08DAFD19FD6B" author="Vicente, M. C. & Mauries, J. - P." box="[444,702,651,675]" pageId="4" pageNumber="146" pagination="9 - 13" refId="ref15742" refString="Vicente, M. C. & Mauries, J. - P. (1980) Un genero y una especie nuevos de Diplopodos Cavernicolas de Guipuzcoa, Espana (Myriapoda - Diplopoda). Speleon, Barcelona, 25, 9 - 13." type="journal article" year="1980">Vicente & Mauriès, 1980</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3506529FFC3CD0856C40880FAD0FA73" pageId="4" pageNumber="146" type="diagnosis">
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<paragraph id="8BF536A2FFC3CD0856C40880FDC3FC2F" blockId="4.[151,1437,721,1468]" pageId="4" pageNumber="146">
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<emphasis id="B93EEAB0FFC3CD0856C40880FEB5FD23" bold="true" box="[151,274,721,747]" pageId="4" pageNumber="146">Diagnosis.</emphasis>
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The newly established family undoubtedly belongs to the suborder
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<taxonomicName id="4C4A4D21FFC3CD0852120880FA87FD23" authorityName="Cook" authorityYear="1895" box="[1089,1312,721,747]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="146" phylum="Arthropoda" rank="subOrder" subOrder="Craspedosomatidea">Craspedosomatidea</taxonomicName>
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. However, considering that the higher classification of the entire order is obviously very chaotic, it is difficult to assign it to a suprafamiliar place at the moment. In any case, it seems to us that
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<taxonomicName id="4C4A4D21FFC3CD0855D10948FBC7FCFB" authority="Antić & Mauriès, 2022" authorityName="Antić & Mauriès" authorityYear="2022" box="[898,1120,793,819]" family="Guipuzcosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" rank="family" status="fam. nov.">Guipuzcosomatidae</taxonomicName>
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<emphasis id="B93EEAB0FFC3CD08523B0948FB71FCFB" bold="true" box="[1128,1238,793,819]" pageId="4" pageNumber="146">
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<taxonomicNameLabel id="A20D57CBFFC3CD08523B0948FB71FCFB" box="[1128,1238,793,819]" pageId="4" pageNumber="146" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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can find its place within any of the two superfamilies
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<taxonomicName id="4C4A4D21FFC3CD08546A096CFCE2FC9F" authorityName="Verhoeff" authorityYear="1899" box="[569,837,829,855]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="superFamily" superFamily="Anthroleucosomatoidea">Anthroleucosomatoidea</taxonomicName>
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(primarily
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<taxonomicName id="4C4A4D21FFC3CD085594096CFBC1FC9F" box="[967,1126,829,855]" class="Diplopoda" family="Anthogonidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" phylum="Arthropoda" rank="family">Anthogonidae</taxonomicName>
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and
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<taxonomicName id="4C4A4D21FFC3CD0852F3096CFAC4FC90" authorityName="Mauries" authorityYear="1970" box="[1184,1379,829,856]" class="Diplopoda" family="Vandeleumatidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" phylum="Arthropoda" rank="family">Vandeleumatidae</taxonomicName>
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) and
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<taxonomicName id="4C4A4D21FFC3CD0856C40930FEEFFCB3" authorityName="Cook" authorityYear="1896" box="[151,328,865,891]" pageId="4" pageNumber="164" rank="superFamily" superFamily="Cleidogonoidea">Cleidogonoidea</taxonomicName>
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since it can be compared to some of their families. We believe that it is superfluous to compare this family with the representatives of other superfamilies of the suborder
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<taxonomicName id="4C4A4D21FFC3CD0855ED09D4FB3EFC57" authorityName="Cook" authorityYear="1895" box="[958,1177,901,927]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="146" phylum="Arthropoda" rank="subOrder" subOrder="Craspedosomatidea">Craspedosomatidea</taxonomicName>
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or with the remaining three suborders, viz., Chordeumatidea, Heterochordeumatidea and Striariidea, since it seems they have nothing in common to the newly established family.
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</paragraph>
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<paragraph id="8BF536A2FFC3CD08569409A0FEF5FA5F" blockId="4.[151,1437,721,1468]" pageId="4" pageNumber="146">
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<taxonomicName id="4C4A4D21FFC3CD08569409A0FE02FBC3" authority="Antić & Mauriès, 2022" authorityName="Antić & Mauriès" authorityYear="2022" box="[199,421,1009,1035]" family="Guipuzcosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" rank="family" status="fam. nov.">Guipuzcosomatidae</taxonomicName>
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<emphasis id="B93EEAB0FFC3CD0857E009A0FD80FBC3" bold="true" box="[435,551,1009,1035]" pageId="4" pageNumber="146">
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<taxonomicNameLabel id="A20D57CBFFC3CD0857E009A0FD80FBC3" box="[435,551,1009,1035]" pageId="4" pageNumber="146" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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differs from the member of the superfamilies
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<taxonomicName id="4C4A4D21FFC3CD08520809A0FAC0FBC3" authorityName="Verhoeff" authorityYear="1899" box="[1115,1383,1009,1035]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="superFamily" superFamily="Anthroleucosomatoidea">Anthroleucosomatoidea</taxonomicName>
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and
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<taxonomicName id="4C4A4D21FFC3CD0856C40E44FEEFFBE7" authorityName="Cook" authorityYear="1896" box="[151,328,1045,1071]" pageId="4" pageNumber="164" rank="superFamily" superFamily="Cleidogonoidea">Cleidogonoidea</taxonomicName>
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by the following combination of characters: male pregonopodal legs 3–7 not incrassate and without tarsal papillae; postgonopodal legs 10 and 11 with coxal sacks, additionally leg 10 with short trochanteral and leg 11 with short coxal process, no other peculiarities; gonopodal sternum of both anterior and posterior gonopods poorly developed and poorly chitinized, mainly membranous; anterior gonopods with striking second pair of angiocoxites starting laterally from coxites, with two branches one with a few very long, flagelliform processes; colpocoxites of the anterior gonopods strongly developed, divided, strongly chitinized, only with rounded membranous remnants; posterior gonopods modified, not leg-like, without articulation and with well-developed, partially chitinized colpocoxites. Fortunately, in many taxa from the superfamilies
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<taxonomicName id="4C4A4D21FFC3CD08550B0F40FBC3FAE3" authorityName="Verhoeff" authorityYear="1899" box="[856,1124,1297,1323]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="superFamily" superFamily="Anthroleucosomatoidea">Anthroleucosomatoidea</taxonomicName>
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and
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<taxonomicName id="4C4A4D21FFC3CD0852C90F40FAECFAE3" authorityName="Cook" authorityYear="1896" box="[1178,1355,1297,1323]" pageId="4" pageNumber="164" rank="superFamily" superFamily="Cleidogonoidea">Cleidogonoidea</taxonomicName>
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vulvae are known, but in none of them they are medially fused, as it is in
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<taxonomicName id="4C4A4D21FFC3CD0855F10F64FB27FA87" authority="Antić & Mauriès, 2022" authorityName="Antić & Mauriès" authorityYear="2022" box="[930,1152,1333,1359]" family="Guipuzcosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" rank="family" status="fam. nov.">Guipuzcosomatidae</taxonomicName>
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<emphasis id="B93EEAB0FFC3CD0852D90F64FB5DFA87" bold="true" box="[1162,1274,1333,1359]" pageId="4" pageNumber="146">
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<taxonomicNameLabel id="A20D57CBFFC3CD0852D90F64FB5DFA87" box="[1162,1274,1333,1359]" pageId="4" pageNumber="146" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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where vulvae are characterized with a striking, chitinized and massive medial bursal structure posteriorly, as one of the possible autapomorphies.
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</paragraph>
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<paragraph id="8BF536A2FFC3CD0856940FF3FAD0FA73" blockId="4.[151,1437,721,1468]" box="[199,1399,1441,1468]" pageId="4" pageNumber="146">
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For more detailed comparison of the new family with some members of
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<taxonomicName id="4C4A4D21FFC3CD0855A00FF0FB07FA73" authorityName="Pocock" authorityYear="1894" box="[1011,1184,1441,1467]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="146" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
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see the discussion.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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