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<mods:title id="B16AAFE7A4CCF7D078C59D51A650B367">Qorimayus, a new genus of relictual, high-altitude harvestmen from western Argentina (Arachnida, Opiliones, Gonyleptidae) reveals trans-Andean phylogenetic links</mods:title>
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<taxonomicName id="4C1A4D30FFDAE012FCDE0D0EFDBAFBF4" ID-CoL="9X5MT" authority="(Hara, 2016)" authorityName="Acosta" authorityYear="2020" baseAuthorityName="Hara" baseAuthorityYear="2016" box="[151,585,1125,1152]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastata" status="comb. nov.">
<emphasis id="B96EEAA1FFDAE012FCDE0D0EFE57FB0B" bold="true" box="[151,420,1125,1152]" italics="true" pageId="23" pageNumber="152">Metabalta polyhastata</emphasis>
(
<bibRefCitation id="EF8B4B42FFDAE012FDFA0D0EFDB3FBF4" author="Hara, M. R." box="[435,576,1125,1151]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara, 2016</bibRefCitation>
)
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<taxonomicNameLabel id="A25D57DAFFDAE012FE190D0EFD25FBF4" box="[592,726,1125,1151]" pageId="23" pageNumber="152" rank="species">comb. nov.</taxonomicNameLabel>
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<emphasis id="B96EEAA1FFDAE012FCDE0DC2FE5AFB4A" box="[151,425,1193,1217]" italics="true" pageId="23" pageNumber="152">Nanophareus polyhastatus</emphasis>
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; 
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Pérez-Schultheiss 
<emphasis id="B96EEAA1FFDAE012FF640DC1FC9AFB4A" box="[813,873,1193,1217]" italics="true" pageId="23" pageNumber="152">et al.,</emphasis>
2019: 10
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. 
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BA536B3FFDAE012FCDE0C65FD70F952" blockId="23.[151,1437,1294,2041]" pageId="23" pageNumber="152">
The results of the cladistic analysis (
<figureCitation id="13212A36FFDAE012FE790C65FD83FAA3" box="[560,624,1294,1320]" captionStart="FIGURE 1" captionStartId="9.[151,250,1687,1712]" captionTargetBox="[155,1430,325,1646]" captionTargetId="figure@9.[151,1436,306,1660]" captionTargetPageId="9" captionText="FIGURE 1. Cladistic relationships of Qorimayus gen. nov.: single most parsimonious tree obtained with implied weights (IW), k=6 (Ci: 0.37, Ri: 0.60, tree length: 376 steps). Character optimization: solid circles depict non-homoplasious states, open circles are homoplasious states. Small numbers on a circle indicate the character number (above the circle) and its state (below); larger numbers under relevant branches display values of bootstrap, jackknife and symmetric resample (B-J-SR), high support values in bold. Abbreviations for major clades (underlined): G: Gonyleptidae; PsG: Pachyloides-group; PG: Pachylus-group; PG+Eu: Pachylus-group plus Eusarcus spp.; MG: Metabalta-group; M+Na(+): Metabalta+Na(+); M: genus Metabalta; Na(+): Nanophareus, plus N+T; Na: genus Nanophareus; e-s: “eyes-separate” clade." figureDoi="http://doi.org/10.5281/zenodo.3605829" httpUri="https://zenodo.org/record/3605829/files/figure.png" pageId="23" pageNumber="152">Fig. 1</figureCitation>
) demonstrated that 
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<emphasis id="B96EEAA1FFDAE012FF1B0C65FB8EFAA3" box="[850,1149,1294,1320]" italics="true" pageId="23" pageNumber="152">Nanophareus polyhastatus</emphasis>
<bibRefCitation id="EF8B4B42FFDAE012F8CA0C64FAF3FAA3" author="Hara, M. R." box="[1155,1280,1294,1321]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara, 2016</bibRefCitation>
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forms a wellsupported clade with species of 
<emphasis id="B96EEAA1FFDAE012FDB30C59FD86FAC7" box="[506,629,1330,1356]" italics="true" pageId="23" pageNumber="152">
<taxonomicName id="4C1A4D30FFDAE012FDB30C59FD82FAC7" ID-CoL="7PNW2" authorityName="Roewer" authorityYear="1913" box="[506,625,1330,1356]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">Metabalta</taxonomicName>
.
</emphasis>
This relationship is underpinned α-taxonomically too, as evidenced by a close examination of the original description and illustrations of 
<taxonomicName id="4C1A4D30FFDAE012FF200C3CFBE4FAFB" ID-CoL="7XDTR" authorityName="Hara, 2016 Hara" authorityYear="2016" box="[873,1047,1366,1392]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastatus">
<emphasis id="B96EEAA1FFDAE012FF200C3CFBE4FAFB" box="[873,1047,1366,1392]" italics="true" pageId="23" pageNumber="152">N. polyhastatus</emphasis>
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: the exomorphological and genital features of this species (
<bibRefCitation id="EF8B4B42FFDAE012FDEA0C11FDD0FA1E" author="Hara, M. R." box="[419,547,1402,1429]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara, 2016</bibRefCitation>
) best match the character states present in 
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<emphasis id="B96EEAA1FFDAE012FFB60C11FB81FA1F" box="[1023,1138,1402,1428]" italics="true" pageId="23" pageNumber="152">Metabalta</emphasis>
</taxonomicName>
, as defined in 
<tableCitation id="C6980308FFDAE012F95C0C11FA99FA1F" box="[1301,1386,1402,1428]" captionStart="TABLE 8" captionStartId="22.[151,239,874,898]" captionTargetBox="[166,1425,947,2011]" captionTargetPageId="22" captionText="TABLE 8. Comparative overview of diagnostic features of genera Qorimayus gen. nov., Metabalta Roewer and Nano- phareus Roewer. Shared character states are indicated as (*)" httpUri="http://table.plazi.org/id/DF65663BFFDBE013FCDE0A01FCE8FC2C" pageId="23" pageNumber="152" tableUuid="DF65663BFFDBE013FCDE0A01FCE8FC2C">Table 8</tableCitation>
. For example, fig. 6A by 
<bibRefCitation id="EF8B4B42FFDAE012FD320CF4FDF0FA32" author="Hara, M. R." box="[379,515,1438,1465]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara (2016)</bibRefCitation>
clearly displays pedipalp coxae with ‘normal’ appearance, 
<emphasis id="B96EEAA1FFDAE012F8D30CF4FB30FA33" box="[1178,1219,1439,1464]" italics="true" pageId="23" pageNumber="152">i.e.,</emphasis>
not elongated as in 
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<emphasis id="B96EEAA1FFDAE012FCDE0CA9FEDEFA57" box="[151,301,1474,1500]" italics="true" pageId="23" pageNumber="152">Nanophareus</emphasis>
</taxonomicName>
. In addition, 
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<emphasis id="B96EEAA1FFDAE012FD810CA8FD8BFA57" box="[456,632,1474,1500]" italics="true" pageId="23" pageNumber="152">N. polyhastatus</emphasis>
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has a ‘normal’ development of basichelicerite and pedipap femur, and the retrolateral apical-subapical spines on tibia are placed in a bifid basal socket, not as large and furcate-tipped as in 
<taxonomicName id="4C1A4D30FFDAE012FCFD0F61FEB9F9AF" ID-CoL="7PPQZ" box="[180,330,1546,1572]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA1FFDAE012FCFD0F61FEB9F9AF" box="[180,330,1546,1572]" italics="true" pageId="23" pageNumber="152">Nanophareus</emphasis>
</taxonomicName>
(
<tableCitation id="C6980308FFDAE012FD110F61FE5DF9AF" box="[344,430,1546,1572]" captionStart="TABLE 8" captionStartId="22.[151,239,874,898]" captionTargetBox="[166,1425,947,2011]" captionTargetPageId="22" captionText="TABLE 8. Comparative overview of diagnostic features of genera Qorimayus gen. nov., Metabalta Roewer and Nano- phareus Roewer. Shared character states are indicated as (*)" httpUri="http://table.plazi.org/id/DF65663BFFDBE013FCDE0A01FCE8FC2C" pageId="23" pageNumber="152" tableUuid="DF65663BFFDBE013FCDE0A01FCE8FC2C">Table 8</tableCitation>
, chars. b, d, h). Several genital characters support the transfer of 
<taxonomicName id="4C1A4D30FFDAE012F8CE0F60FAC5F9AF" ID-CoL="7XDTR" authorityName="Hara, 2016 Hara" authorityYear="2016" box="[1159,1334,1546,1572]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastatus">
<emphasis id="B96EEAA1FFDAE012F8CE0F60FAC5F9AF" box="[1159,1334,1546,1572]" italics="true" pageId="23" pageNumber="152">N. polyhastatus</emphasis>
</taxonomicName>
into 
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<emphasis id="B96EEAA1FFDAE012F9260F60FF15F9C3" italics="true" pageId="23" pageNumber="152">Metabalta</emphasis>
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, like the sigmoid curvature of stylus, its orientation and apical narrowing, and the apical end of VPS (
<tableCitation id="C6980308FFDAE012F9080F45FA65F9C3" box="[1345,1430,1582,1608]" captionStart="TABLE 8" captionStartId="22.[151,239,874,898]" captionTargetBox="[166,1425,947,2011]" captionTargetPageId="22" captionText="TABLE 8. Comparative overview of diagnostic features of genera Qorimayus gen. nov., Metabalta Roewer and Nano- phareus Roewer. Shared character states are indicated as (*)" httpUri="http://table.plazi.org/id/DF65663BFFDBE013FCDE0A01FCE8FC2C" pageId="23" pageNumber="152" tableUuid="DF65663BFFDBE013FCDE0A01FCE8FC2C">Table 8</tableCitation>
, chars. o, p, r, s); drawings of 
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are not detailed enough to see if also char. t (subapical spines on VPS shaft) applies. Based on this evidence, I hereby formally propose the new combination 
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<emphasis id="B96EEAA1FFDAE012F8020F1DFAB8F91B" box="[1099,1355,1654,1680]" italics="true" pageId="23" pageNumber="152">Metabalta polyhastata</emphasis>
(
<bibRefCitation id="EF8B4B42FFDAE012F9140F1DFF22F93E" author="Hara, M. R." pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara, 2016</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25D57DAFFDAE012FCA90FF1FE91F93E" box="[224,354,1690,1717]" pageId="23" pageNumber="152" rank="species">
<emphasis id="B96EEAA1FFDAE012FCA90FF1FEA8F93F" bold="true" box="[224,347,1690,1716]" pageId="23" pageNumber="152">comb. nov</emphasis>
.
</taxonomicNameLabel>
(the original spelling 
<taxonomicName id="4C1A4D30FFDAE012FE100FF1FD14F93F" ID-CoL="7XDTR" authorityName="Hara, 2016 Hara" authorityYear="2016" box="[601,743,1690,1716]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastatus">
<emphasis id="B96EEAA1FFDAE012FE100FF1FD14F93F" box="[601,743,1690,1716]" italics="true" pageId="23" pageNumber="152">polyhastatus</emphasis>
</taxonomicName>
changed to 
<taxonomicName id="4C1A4D30FFDAE012FF260FF1FC01F93F" ID-CoL="9X5MT" authorityName="Acosta" authorityYear="2020" baseAuthorityName="Hara" baseAuthorityYear="2016" box="[879,1010,1690,1716]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastata">
<emphasis id="B96EEAA1FFDAE012FF260FF1FC01F93F" box="[879,1010,1690,1716]" italics="true" pageId="23" pageNumber="152">polyhastata</emphasis>
</taxonomicName>
to ensure the agreement in grammatical gender with the genus name 
<taxonomicName id="4C1A4D30FFDAE012FE490FD5FD80F953" ID-CoL="7PNW2" authorityName="Roewer" authorityYear="1913" box="[512,627,1726,1752]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA1FFDAE012FE490FD5FD80F953" box="[512,627,1726,1752]" italics="true" pageId="23" pageNumber="152">Metabalta</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3006538FFDAE012FC8E0F89FB8AF85E" pageId="23" pageNumber="152" type="description">
<paragraph id="8BA536B3FFDAE012FC8E0F89FB8AF85E" blockId="23.[151,1437,1294,2041]" pageId="23" pageNumber="152">
Aside, 
<bibRefCitation id="EF8B4B42FFDAE012FD5C0F89FD99F977" author="Perez-Schultheiss, J. &amp; Urra, F. &amp; Otarola, A." box="[277,618,1762,1788]" pageId="23" pageNumber="152" pagination="1 - 24" refId="ref18157" refString="Perez-Schultheiss, J., Urra, F. &amp; Otarola, A. (2019) Opiliones Laniatores (Arachnida) de la Cordillera de Nahuelbuta: un desconocido hotspot de diversidad. Boletin Nahuelbuta Natural, 4, 1 - 24." type="journal article" year="2019">
Pérez-Schultheiss 
<emphasis id="B96EEAA1FFDAE012FDAB0F88FDEAF977" box="[482,537,1762,1788]" italics="true" pageId="23" pageNumber="152">et al.</emphasis>
(2019)
</bibRefCitation>
suggested that 
<taxonomicName id="4C1A4D30FFDAE012FF5A0F88FC32F977" authorityName="Hara, 2016 Hara" authorityYear="2016" box="[787,961,1762,1788]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastatus">
<emphasis id="B96EEAA1FFDAE012FF5A0F88FC32F977" box="[787,961,1762,1788]" italics="true" pageId="23" pageNumber="152">N. polyhastatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1A4D30FFDAE012FFBD0F89FA6FF977" authority="Mello-Leitao, 1931" authorityName="Mello-Leitao" authorityYear="1931" box="[1012,1436,1762,1788]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="B96EEAA1FFDAE012FFBD0F89FB4DF977" box="[1012,1214,1762,1788]" italics="true" pageId="23" pageNumber="152">Metabalta albipes</emphasis>
<bibRefCitation id="EF8B4B42FFDAE012F88D0F89FA6FF977" author="Mello-Leitao, C. de" box="[1220,1436,1762,1788]" pageId="23" pageNumber="152" pagination="83 - 97" refId="ref17992" refString="Mello-Leitao, C. de (1931) Notas sobre arachnideos argentinos. Annaes da Academia Brasileira de Ciencias, 3 (2), 83 - 97." type="journal article" year="1931">Mello-Leitão, 1931</bibRefCitation>
</taxonomicName>
might prove to be the same. Although the latter species has no original figure available for a comparison, that of 
<taxonomicName id="4C1A4D30FFDAE012FCDE0E41FDCDF8CF" authority="Mello-Leitao, 1936" authorityName="Mello-Leitao" authorityYear="1936" box="[151,574,1834,1860]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="porteri">
<emphasis id="B96EEAA1FFDAE012FCDE0E41FEACF8CF" box="[151,351,1834,1860]" italics="true" pageId="23" pageNumber="152">Metabalta porteri</emphasis>
<bibRefCitation id="EF8B4B42FFDAE012FD2C0E41FDCDF8CF" author="Mello-Leitao, C. de" box="[357,574,1834,1860]" pageId="23" pageNumber="152" pagination="112 - 129" refId="ref18021" refString="Mello-Leitao, C. de (1936) Etude sur les arachnides de Papudo et Constitucion (Chili), recueillis par le Prof. Dr. Carlos E. Porter. Revista Chilena de Historia Natural, 40, 112 - 129." type="journal article" year="1936">Mello-Leitão, 1936</bibRefCitation>
</taxonomicName>
(currently under synonymy of 
<taxonomicName id="4C1A4D30FFDAE012FFD10E40FBFDF8CF" authorityName="Mello-Leitao" authorityYear="1931" box="[920,1038,1834,1860]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="B96EEAA1FFDAE012FFD10E40FBFDF8CF" box="[920,1038,1834,1860]" italics="true" pageId="23" pageNumber="152">M. albipes</emphasis>
</taxonomicName>
, after 
<bibRefCitation id="EF8B4B42FFDAE012F81C0E41FAF2F8CF" author="Ringuelet, R. A." box="[1109,1281,1833,1860]" pageId="23" pageNumber="152" pagination="127 - 439" refId="ref18325" refString="Ringuelet, R. A. (1959) Los aracnidos argentinos del Orden Opiliones. Revista del Museo Argentino de Ciencias Naturales, Ciencias Zoologicas, 5 (2), 127 - 439, pls. I-XX." type="journal article" year="1959">Ringuelet 1959</bibRefCitation>
) looks identical to 
<taxonomicName id="4C1A4D30FFDAE012FC930E24FE72F8E3" authorityName="Acosta" authorityYear="2020" baseAuthorityName="Hara" baseAuthorityYear="2016" box="[218,385,1870,1896]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastata">
<emphasis id="B96EEAA1FFDAE012FC930E24FE72F8E3" box="[218,385,1870,1896]" italics="true" pageId="23" pageNumber="152">M. polyhastata</emphasis>
</taxonomicName>
, as described and drawn by 
<bibRefCitation id="EF8B4B42FFDAE012FEF30E24FCB7F8E2" author="Hara, M. R." box="[698,836,1870,1897]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara (2016)</bibRefCitation>
; key similarities refer to the general apophysis pattern on femur IV, and the shape (and thickness) of apophysis of coxa IV. Until the relevant 
<typeStatus id="54A18811FFDAE012F8150E18FB64F806" box="[1116,1175,1907,1933]" pageId="23" pageNumber="152">types</typeStatus>
are studied I prefer not to formalize this presumed specific synonymy. It should be noted that 
<taxonomicName id="4C1A4D30FFDAE012FFE00EFCFBA2F83B" authorityName="Acosta" authorityYear="2020" baseAuthorityName="Hara" baseAuthorityYear="2016" box="[937,1105,1942,1968]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="polyhastata">
<emphasis id="B96EEAA1FFDAE012FFE00EFCFBA2F83B" box="[937,1105,1942,1968]" italics="true" pageId="23" pageNumber="152">M. polyhastata</emphasis>
</taxonomicName>
was included by 
<bibRefCitation id="EF8B4B42FFDAE012F95D0EFCFA6FF83A" author="Hara, M. R." box="[1300,1436,1942,1969]" pageId="23" pageNumber="152" pagination="101 - 123" refId="ref17163" refString="Hara, M. R. (2016) Cladistic analysis and description of three new species of the Chilean genus Nanophareus (Opiliones: Gonyleptidae: Pachylinae). Zootaxa, 4105 (2), 101 - 123. https: // doi. org / 10.11646 / zootaxa. 4105.2.1" type="journal article" year="2016">Hara (2016)</bibRefCitation>
in his cladistic analysis of 
<taxonomicName id="4C1A4D30FFDAE012FDF60ED1FDA6F85F" box="[447,597,1978,2004]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA1FFDAE012FDF60ED1FDA6F85F" box="[447,597,1978,2004]" italics="true" pageId="23" pageNumber="152">Nanophareus</emphasis>
</taxonomicName>
, but resulted in an internal position in the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3006538FFDAE01DFC8E0EB6FEF4FE94" lastPageId="24" lastPageNumber="153" pageId="23" pageNumber="152" type="materials_examined">
<paragraph id="8BA536B3FFDAE01DFC8E0EB6FEF4FE94" blockId="23.[151,1437,1294,2041]" lastBlockId="24.[151,1436,152,287]" lastPageId="24" lastPageNumber="153" pageId="23" pageNumber="152">
<materialsCitation id="3B723CEEFFDAE012FC8E0EB6FA6BF873" ID-GBIF-Occurrence="2549107390" box="[199,1432,2013,2041]" collectionCode="SMF" collectorName="de Chillan" country="Chile" location="Sierra de Chillan" pageId="23" pageNumber="152" specimenCode="RII 952" specimenCount="3" specimenCount-male="2" typeStatus="syntype">
<emphasis id="B96EEAA1FFDAE012FC8E0EB6FDC5F873" bold="true" box="[199,566,2013,2040]" pageId="23" pageNumber="152">Additional material examined.</emphasis>
<taxonomicName id="4C1A4D30FFDAE012FE740EB5FCD6F873" authorityName="Roewer" authorityYear="1929" box="[573,805,2014,2040]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="efformata">
<emphasis id="B96EEAA1FFDAE012FE740EB5FCD6F873" box="[573,805,2014,2040]" italics="true" pageId="23" pageNumber="152">Metabalta efformata</emphasis>
</taxonomicName>
: 
<collectingCountry id="F30D7623FFDAE012FF7D0EB5FC86F873" box="[820,885,2014,2040]" name="Chile" pageId="23" pageNumber="152">Chile</collectingCountry>
: 
<location id="8EC56068FFDAE012FFC80EB5FBB1F873" LSID="urn:lsid:plazi:treatment:03B387A5FFDAE01DFCDE0D0EFEF4FE94:8EC56068FFDAE012FFC80EB5FBB1F873" box="[897,1090,2014,2040]" country="Chile" name="Sierra de Chillan" pageId="23" pageNumber="152">
Sierra 
<collectorName id="26EF5365FFDAE012FF820EB5FBB1F873" box="[971,1090,2014,2040]" pageId="23" pageNumber="152">de Chillán</collectorName>
</location>
, 
<specimenCount id="9D1CFD3AFFDAE012F8060EB5FB8AF872" box="[1103,1145,2014,2041]" pageId="23" pageNumber="152" type="male">2 ♂</specimenCount>
<typeStatus id="54A18811FFDAE012F8C80EB4FB10F872" box="[1153,1251,2015,2041]" pageId="23" pageNumber="152" type="syntype">syntypes</typeStatus>
(
<collectionCode id="ED0BAE76FFDAE012F8BA0EB5FADEF873" box="[1267,1325,2014,2040]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="152">SMF</collectionCode>
<specimenCode id="DBBC9EC8FFDAE012F97D0EB4FA7CF873" box="[1332,1423,2013,2041]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="152">RII 952</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B723CEEFFD5E01DFCDE09F2FC0AFF38" ID-GBIF-Occurrence="2549107387" box="[151,1017,152,179]" collectionCode="SMF" country="Chile" location="Valparaiso" pageId="24" pageNumber="153" specimenCode="RII 951" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Valparaiso" typeStatus="syntype">
<taxonomicName id="4C1A4D30FFD5E01DFCDE09F2FE7AFF38" authorityName="Roewer" authorityYear="1929" box="[151,393,153,179]" class="Arachnida" family="Gonyleptidae" genus="Metabalta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="geniculata">
<emphasis id="B96EEAA1FFD5E01DFCDE09F2FE7AFF38" box="[151,393,153,179]" italics="true" pageId="24" pageNumber="153">Metabalta geniculata</emphasis>
</taxonomicName>
: 
<collectingCountry id="F30D7623FFD5E01DFDD309F2FE28FF38" box="[410,475,153,179]" name="Chile" pageId="24" pageNumber="153">Chile</collectingCountry>
: 
<collectingRegion id="49DEF851FFD5E01DFDAE09F2FD93FF38" box="[487,608,153,179]" country="Chile" name="Valparaiso" pageId="24" pageNumber="153">Valparaíso</collectingRegion>
, 
<specimenCount id="9D1CFD3AFFD5E01DFE2709F2FD6FFF38" box="[622,668,153,179]" pageId="24" pageNumber="153" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D1CFD3AFFD5E01DFEE309F2FD25FF38" box="[682,726,153,179]" pageId="24" pageNumber="153" type="female">1 ♀</specimenCount>
<typeStatus id="54A18811FFD5E01DFE9609F2FCB2FF38" box="[735,833,153,179]" pageId="24" pageNumber="153" type="syntype">syntypes</typeStatus>
(
<collectionCode id="ED0BAE76FFD5E01DFF1809F2FC78FF38" box="[849,907,153,179]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="153">SMF</collectionCode>
<specimenCode id="DBBC9EC8FFD5E01DFFDD09F2FC02FF38" box="[916,1009,152,179]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="153">RII 951</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B723CEEFFD5E01DF84E09F2FEF0FE94" ID-GBIF-Occurrence="2549107386" collectingDate="1985-01-09" collectionCode="AMNH" collectorName="Los Molles &amp; N. I. Platnick &amp; O. F. Francke" country="Chile" elevation="2" location="Chile'" pageId="24" pageNumber="153" specimenCode="SMF 986" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="lectotype">
<taxonomicName id="4C1A4D30FFD5E01DF84E09F2FAF3FF38" authorityName="Roewer" authorityYear="1929" box="[1031,1280,153,179]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="B96EEAA1FFD5E01DF84E09F2FAF3FF38" box="[1031,1280,153,179]" italics="true" pageId="24" pageNumber="153">Nanophareus palpalis</emphasis>
</taxonomicName>
: ‘ 
<location id="8EC56068FFD5E01DF95C09F2FA91FF38" LSID="urn:lsid:plazi:treatment:03B387A5FFDAE01DFCDE0D0EFEF4FE94:8EC56068FFD5E01DF95C09F2FA91FF38" box="[1301,1378,153,179]" country="Chile" name="Chile'" pageId="24" pageNumber="153">Chile’</location>
, 
<specimenCount id="9D1CFD3AFFD5E01DF93909F2FA6FFF38" box="[1392,1436,153,179]" pageId="24" pageNumber="153" type="male">1 ♂</specimenCount>
<typeStatus id="54A18811FFD5E01DFCDE09D6FEF2FF5C" box="[151,257,189,215]" pageId="24" pageNumber="153" type="lectotype">lectotype</typeStatus>
, 
<specimenCount id="9D1CFD3AFFD5E01DFD5909D6FECCFF5C" box="[272,319,189,215]" pageId="24" pageNumber="153" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D1CFD3AFFD5E01DFD0609D6FE8EFF5C" box="[335,381,189,215]" pageId="24" pageNumber="153" type="female">2 ♀</specimenCount>
<typeStatus id="54A18811FFD5E01DFDCE09D6FDDAFF5C" box="[391,553,189,215]" pageId="24" pageNumber="153" type="paralectotype">paralectotypes</typeStatus>
(designated by 
<bibRefCitation id="EF8B4B42FFD5E01DFEAF09D6FC59FF5C" author="Hara, M. R. &amp; Pinto-da-Rocha, R. &amp; Kury, A. B." box="[742,938,189,215]" pageId="24" pageNumber="153" pagination="37 - 66" refId="ref17272" refString="Hara, M. R., Pinto-da-Rocha, R. &amp; Kury, A. B. (2012) Revision of Nanophareus, a mysterious harvestman genus from Chile, with descriptions of three new species (Opiliones: Laniatores: Gonyleptidae). Zootaxa, 3579 (1), 37 - 66. https: // doi. org / 10.11646 / zootaxa. 3579.1.2" type="journal article" year="2012">
Hara 
<emphasis id="B96EEAA1FFD5E01DFF6E09D5FC90FF5C" box="[807,867,189,215]" italics="true" pageId="24" pageNumber="153">et al.</emphasis>
2012
</bibRefCitation>
; 
<specimenCode id="DBBC9EC8FFD5E01DFFF109D6FBD8FF53" box="[952,1067,189,216]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="24" pageNumber="153">SMF 986</specimenCode>
/1); 
<taxonomicName id="4C1A4D30FFD5E01DF81E09D6FA67FF5C" box="[1111,1428,189,215]" class="Arachnida" family="Gonyleptidae" genus="Nanophareus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="bosqenublado">
<emphasis id="B96EEAA1FFD5E01DF81E09D6FA67FF5C" box="[1111,1428,189,215]" italics="true" pageId="24" pageNumber="153">Nanophareus bosqenublado</emphasis>
</taxonomicName>
: 
<collectingCountry id="F30D7623FFD5E01DFCDE098AFF2BFF70" box="[151,216,225,251]" name="Chile" pageId="24" pageNumber="153">Chile</collectingCountry>
: 
<collectorName id="26EF5365FFD5E01DFCAC098AFE9AFF70" box="[229,361,225,251]" pageId="24" pageNumber="153">Los Molles</collectorName>
, elev. 
<quantity id="4CE29B56FFD5E01DFDFB098AFE12FF70" box="[434,481,225,251]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="153" unit="m" value="2.0">
<elevation id="0037D180FFD5E01DFDFB098AFE12FF70" box="[434,481,225,251]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="153" unit="m" value="2.0">2 m</elevation>
</quantity>
, under succulent rock cover along coast, 
<date id="FFA41073FFD5E01DFFF4098AFBB7FF70" box="[957,1092,224,251]" pageId="24" pageNumber="153" value="1985-01-09">
<collectingDate id="EFE0E99BFFD5E01DFFF4098AFBB7FF70" box="[957,1092,224,251]" pageId="24" pageNumber="153" value="1985-01-09">9 Jan. 1985</collectingDate>
</date>
, 
<collectorName id="26EF5365FFD5E01DF818098AFB11FF70" box="[1105,1250,225,251]" pageId="24" pageNumber="153">N.I. Platnick</collectorName>
&amp; 
<collectorName id="26EF5365FFD5E01DF940098AFA6FFF70" box="[1289,1436,225,251]" pageId="24" pageNumber="153">O.F. Francke</collectorName>
(
<collectionCode id="ED0BAE76FFD5E01DFCD6086EFF08FE94" box="[159,251,261,287]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="153">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>