treatments-xml/data/C1/7B/87/C17B87FF3C3C1235C0E6FD50FE63F307.xml

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<mods:title id="485E46645CDE322D6EA8D1C221F3E104">Type designations for Neotropical water beetles in the genus Hydrochus Leach 1817 (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="C17B87FF3C3C1235C0E6FD50FE63F307" ID-GBIF-Taxon="160605446" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307" httpUri="http://treatment.plazi.org/id/C17B87FF3C3C1235C0E6FD50FE63F307" lastPageId="3" lastPageNumber="152" pageId="2" pageNumber="151">
<subSubSection id="01C865623C3C1234C0E6FD50FE72F004" box="[151,480,1818,1845]" pageId="2" pageNumber="151" type="nomenclature">
<paragraph id="496D36E93C3C1234C0E6FD50FE72F004" blockId="2.[151,480,1818,1880]" box="[151,480,1818,1845]" pageId="2" pageNumber="151">
<heading id="122581853C3C1234C0E6FD50FE72F004" bold="true" box="[151,480,1818,1845]" fontSize="11" level="1" pageId="2" pageNumber="151" reason="1">
<taxonomicName id="8ED24D6A3C3C1234C0E6FD50FE72F004" ID-CoL="3N65N" authority="Knisch" authorityName="Knisch. Labels" authorityYear="1921" box="[151,480,1818,1845]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3C1234C0E6FD50FE72F004" bold="true" box="[151,480,1818,1845]" pageId="2" pageNumber="151">
<emphasis id="7BA6EAFB3C3C1234C0E6FD50FE13F005" bold="true" box="[151,385,1818,1845]" italics="true" pageId="2" pageNumber="151">Hydrochus argutus</emphasis>
Knisch
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="496D36E93C3C1234C0E6FD74FE8CF068" blockId="2.[151,480,1818,1880]" box="[151,286,1854,1880]" pageId="2" pageNumber="151">
<figureCitation id="D1E92A6C3C3C1234C0E6FD74FF71F068" box="[151,227,1854,1880]" captionStart="FIGURE 1" captionStartId="2.[151,250,1639,1664]" captionTargetBox="[177,1418,249,1576]" captionTargetId="figure@2.[151,1435,234,1611]" captionTargetPageId="2" captionText="FIGURE 1. Hydrochus argutus Knisch, habitus of lectotype, and male genitalia of specimen from Brazil." figureDoi="http://doi.org/10.5281/zenodo.3557929" httpUri="https://zenodo.org/record/3557929/files/figure.png" pageId="2" pageNumber="151">
Figs.
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</figureCitation>
,
<figureCitation id="D1E92A6C3C3C1234C080FD74FE8CF068" box="[241,286,1854,1880]" captionStart="FIGURES 12" captionStartId="15.[151,264,1980,2005]" captionTargetBox="[215,1415,381,1942]" captionTargetId="figure@15.[158,1430,381,1955]" captionTargetPageId="15" captionText="FIGURES 12AE. Labels of lectotypes. (A) Hydrochus ducalis Knisch; (B) Hydrochus drakei Knisch; (C) Hydrochus variabilis Knisch; (D) Hydrochus corruscans Bruch; (E) Hydrochus argutus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557951" httpUri="https://zenodo.org/record/3557951/files/figure.png" pageId="2" pageNumber="151">12E</figureCitation>
</paragraph>
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<paragraph id="496D36E93C3C1234C0E6FDC8FE6DF0AB" blockId="2.[151,764,1922,1977]" box="[151,511,1922,1947]" pageId="2" pageNumber="151">
<treatmentCitationGroup id="69C211C73C3C1234C0E6FDC8FE6DF0AB" box="[151,511,1922,1947]" pageId="2" pageNumber="151">
<taxonomicName id="8ED24D6A3C3C1234C0E6FDC8FE6EF0AB" ID-CoL="3N65N" authority="Knisch 1921: 3" authorityName="Knisch. Labels" authorityPageNumber="3" authorityYear="1921" box="[151,508,1922,1947]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3C1234C0E6FDC8FECAF0AA" box="[151,344,1922,1946]" italics="true" pageId="2" pageNumber="151">Hydrochus argutus</emphasis>
<treatmentCitation id="C87310F83C3C1234C12EFDC8FE6EF0AB" author="Knisch, A." box="[351,508,1922,1947]" page="3" pageId="2" pageNumber="151" year="1921">
<bibRefCitation id="2D434B183C3C1234C12EFDC8FE6EF0AB" author="Knisch, A." box="[351,508,1922,1947]" pageId="2" pageNumber="151" pagination="1 - 24" refId="ref5602" refString="Knisch, A. (1921) Hydrophiliden aus Matto Grosso (Erster Teil). Archiv fur Naturegeschichte, 87 A, 1 - 24." type="journal article" year="1921">Knisch 1921: 3</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="496D36E93C3C1235C0E6FDEBFCA1F553" blockId="2.[151,764,1922,1977]" lastBlockId="3.[151,1437,153,1079]" lastPageId="3" lastPageNumber="152" pageId="2" pageNumber="151">
<taxonomicName id="8ED24D6A3C3C1234C0E6FDEBFDFBF089" ID-CoL="3N68X" authority="Makhan 1995" authorityName="Makhan" authorityYear="1995" box="[151,617,1952,1977]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="johannapietersenae">
<emphasis id="7BA6EAFB3C3C1234C0E6FDEBFE46F089" box="[151,468,1953,1977]" italics="true" pageId="2" pageNumber="151">Hydrochus johannapietersenae</emphasis>
Makhan 1995
</taxonomicName>
;
<bibRefCitation id="2D434B183C3C1234C202FDEAFD6BF089" author="Oliva, A." box="[627,761,1952,1977]" pageId="2" pageNumber="151" pagination="301 - 341" refId="ref6127" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
. The RBINS kindly provided eight
<typeStatus id="9669884B3C3D1235C268FAD3FDE9F783" box="[537,635,153,179]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
<materialsCitation id="F9BA3CB43C3D1235C2F3FAD3FDA7F7E7" ID-GBIF-Occurrence="2460219433" county="Five" location="Orchymont" municipality="Knisch" pageId="3" pageNumber="152" specimenCount="1" typeStatus="syntype">
of
<taxonomicName id="8ED24D6A3C3D1235C2D1FAD3FC5CF783" ID-CoL="3N65N" authority="Knisch" authorityName="Knisch. Labels" authorityYear="1921" box="[672,974,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3D1235C2D1FAD3FCE4F783" box="[672,886,153,179]" italics="true" pageId="3" pageNumber="152">Hydrochus argutus</emphasis>
Knisch
</taxonomicName>
, which are part of the
<location id="4C0D60323C3D1235C4B7FAD3FAD4F783" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C4B7FAD3FAD4F783" box="[1222,1350,153,179]" county="Five" municipality="Knisch" name="Orchymont" pageId="3" pageNumber="152">Orchymont</location>
/
<collectingMunicipality id="A909AC933C3D1235C53FFAD3FA0FF783" box="[1358,1437,153,179]" pageId="3" pageNumber="152">Knisch</collectingMunicipality>
types collection.
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of the
<typeStatus id="9669884B3C3D1235C1A2FAF7FDA7F7E7" box="[467,565,189,215]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
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<materialsCitation id="F9BA3CB43C3D1235C24AFAF7FBF3F7CB" ID-GBIF-Occurrence="2460219444" collectorName="This" location="One" pageId="3" pageNumber="152" specimenCount="1" typeStatus="syntype">
are females, one of which is in three pieces.
<collectorName id="E427533F3C3D1235C457FAF7FBCAF7E7" box="[1062,1112,189,215]" pageId="3" pageNumber="152">This</collectorName>
specimen has been dissected by a prior worker, but the genital segments are not with the specimen.
<location id="4C0D60323C3D1235C3BFFAABFC6FF7CB" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C3BFFAABFC6FF7CB" box="[974,1021,225,251]" name="One" pageId="3" pageNumber="152">One</location>
<typeStatus id="9669884B3C3D1235C476FAABFBF3F7CB" box="[1031,1121,225,251]" pageId="3" pageNumber="152" type="syntype">syntype</typeStatus>
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,
<materialsCitation id="F9BA3CB43C3D1235C41FFAABFE6FF673" ID-GBIF-Occurrence="2460219434" location="One" pageId="3" pageNumber="152" specimenCount="1" typeStatus="syntype">
labeled as male by a prior worker, is missing the head and prothorax, has been dissected, but the genital segments and male genitalia are not with the specimen.
<location id="4C0D60323C3D1235C11FFB63FE0FF673" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C11FFB63FE0FF673" box="[366,413,297,323]" name="One" pageId="3" pageNumber="152">One</location>
<typeStatus id="9669884B3C3D1235C1D2FB63FE6FF673" box="[419,509,297,323]" pageId="3" pageNumber="152" type="syntype">syntype</typeStatus>
</materialsCitation>
,
<materialsCitation id="F9BA3CB43C3D1235C276FB63FD2AF657" ID-GBIF-Occurrence="2460219438" collectorName="The" location="One" pageId="3" pageNumber="152" specimenCount="1" typeStatus="syntype">
gender unknown, is in two pieces.
<collectorName id="E427533F3C3D1235C3F6FB63FC21F673" box="[903,947,297,323]" pageId="3" pageNumber="152">The</collectorName>
genital segments are not with the specimen, hence the gender is unknown.
<location id="4C0D60323C3D1235C19AFB07FD88F657" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C19AFB07FD88F657" box="[491,538,333,359]" name="One" pageId="3" pageNumber="152">One</location>
male
<typeStatus id="9669884B3C3D1235C22FFB07FD2AF657" box="[606,696,333,359]" pageId="3" pageNumber="152" type="syntype">syntype</typeStatus>
</materialsCitation>
, dissected by a prior worker, is in six pieces. The male genitalia is partially crushed, and approximately the distal ½ of both parameres are missing, making the aedeagus appear to be much longer than the parameres. A complete male specimen, with complete male genitalia, is not available to serve as
<typeStatus id="9669884B3C3D1235C0C6FBF3FEB3F6E3" box="[183,289,441,467]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
. Therefore, I have chosen a relatively good
<typeStatus id="9669884B3C3D1235C369FBF3FCFDF6E3" box="[792,879,441,467]" pageId="3" pageNumber="152" type="syntype">syntype</typeStatus>
female to serve as
<typeStatus id="9669884B3C3D1235C43EFBF3FB2BF6E3" box="[1103,1209,441,467]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
,
<materialsCitation id="F9BA3CB43C3D1235C4B4FBF3FD6AF6C7" ID-GBIF-Occurrence="2460219437" collectorName="This" location="The" pageId="3" pageNumber="152" specimenCount="1" typeStatus="lectotype">
and have labeled it such.
<collectorName id="E427533F3C3D1235C0A8FB97FE99F6C7" box="[217,267,477,503]" pageId="3" pageNumber="152">This</collectorName>
specimen is imaged (
<figureCitation id="D1E92A6C3C3D1235C273FB97FDD6F6C7" box="[514,580,477,503]" captionStart="FIGURE 1" captionStartId="2.[151,250,1639,1664]" captionTargetBox="[177,1418,249,1576]" captionTargetId="figure@2.[151,1435,234,1611]" captionTargetPageId="2" captionText="FIGURE 1. Hydrochus argutus Knisch, habitus of lectotype, and male genitalia of specimen from Brazil." figureDoi="http://doi.org/10.5281/zenodo.3557929" httpUri="https://zenodo.org/record/3557929/files/figure.png" pageId="3" pageNumber="152">Fig. 1</figureCitation>
).
<location id="4C0D60323C3D1235C22CFB97FD1BF6C7" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C22CFB97FD1BF6C7" box="[605,649,477,503]" name="The" pageId="3" pageNumber="152">The</location>
<typeStatus id="9669884B3C3D1235C2E0FB97FD6AF6C7" box="[657,760,477,503]" pageId="3" pageNumber="152" type="lectotype">lectotype</typeStatus>
</materialsCitation>
<materialsCitation id="F9BA3CB43C3D1235C28EFB97FE21F553" ID-GBIF-Occurrence="2460219441" collectorName="Coll. A. Knisch &amp; Ex &amp; P. D. Perkins &amp; P. D. Perkins." determinerName="Kniz" location="Bras. Corumba" municipality="Matto Grosso" pageId="3" pageNumber="152" specimenCount="2" typeStatus="lectotype">
has the following labels (
<figureCitation id="D1E92A6C3C3D1235C46CFB97FB16F6C7" box="[1053,1156,477,503]" captionStart="FIGURES 12" captionStartId="15.[151,264,1980,2005]" captionTargetBox="[215,1415,381,1942]" captionTargetId="figure@15.[158,1430,381,1955]" captionTargetPageId="15" captionText="FIGURES 12AE. Labels of lectotypes. (A) Hydrochus ducalis Knisch; (B) Hydrochus drakei Knisch; (C) Hydrochus variabilis Knisch; (D) Hydrochus corruscans Bruch; (E) Hydrochus argutus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557951" httpUri="https://zenodo.org/record/3557951/files/figure.png" pageId="3" pageNumber="152">Fig. 12E</figureCitation>
): “
<location id="4C0D60323C3D1235C4D2FB97FADAF6C7" LSID="urn:lsid:plazi:treatment:C17B87FF3C3C1235C0E6FD50FE63F307:4C0D60323C3D1235C4D2FB97FADAF6C7" box="[1187,1352,477,503]" municipality="Matto Grosso" name="Bras. Corumba" pageId="3" pageNumber="152">Bras.Corumba</location>
[
<collectingMunicipality id="A909AC933C3D1235C527FB97FF7BF52B" pageId="3" pageNumber="152">Matto Grosso</collectingMunicipality>
] //
<determinerName id="BCB1DA2D3C3D1235C17DF84BFED0F52B" box="[268,322,513,539]" pageId="3" pageNumber="152">Kniž</determinerName>
det. argutus m. //
<collectorName id="E427533F3C3D1235C279F84BFD3CF52B" box="[520,686,513,539]" pageId="3" pageNumber="152">Coll. A.Knisch</collectorName>
COTYPUS //
<collectorName id="E427533F3C3D1235C33EF84BFCFCF52B" box="[847,878,513,539]" pageId="3" pageNumber="152">Ex</collectorName>
coll. A.dOrchymont // HABITUS/GENIT. DIGI- TAL IMAGES captured 2019
<collectorName id="E427533F3C3D1235C19DF86FFDE4F50F" box="[492,630,549,575]" pageId="3" pageNumber="152">P.D. Perkins</collectorName>
// [red]
<typeStatus id="9669884B3C3D1235C2A3F86FFCE0F50F" box="[722,882,549,575]" pageId="3" pageNumber="152" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="8ED24D6A3C3D1235C30BF86FFB34F50F" ID-CoL="3N65N" authority="Knisch" authorityName="Knisch. Labels" authorityYear="1921" box="[890,1190,549,575]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3D1235C30BF86FFBC2F50F" box="[890,1104,549,575]" italics="true" pageId="3" pageNumber="152">Hydrochus argutus</emphasis>
Knisch
</taxonomicName>
desig.
<collectorName id="E427533F3C3D1235C486F86FFA01F50F" box="[1271,1427,549,575]" pageId="3" pageNumber="152">P. D. Perkins.</collectorName>
” The seven other
<typeStatus id="9669884B3C3D1235C120F803FE21F553" box="[337,435,585,611]" pageId="3" pageNumber="152" type="syntype">syntypes</typeStatus>
</materialsCitation>
have been labeled
<typeStatus id="9669884B3C3D1235C2FBF803FCBDF553" box="[650,815,585,611]" pageId="3" pageNumber="152" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="01C865623C3D1235C0B6F827FAE0F5FF" pageId="3" pageNumber="152" type="description">
<paragraph id="496D36E93C3D1235C0B6F827FAE0F5FF" blockId="3.[151,1437,153,1079]" pageId="3" pageNumber="152">
The male genitalia illustrations (
<figureCitation id="D1E92A6C3C3D1235C24DF827FDECF5B7" box="[572,638,621,647]" captionStart="FIGURE 1" captionStartId="2.[151,250,1639,1664]" captionTargetBox="[177,1418,249,1576]" captionTargetId="figure@2.[151,1435,234,1611]" captionTargetPageId="2" captionText="FIGURE 1. Hydrochus argutus Knisch, habitus of lectotype, and male genitalia of specimen from Brazil." figureDoi="http://doi.org/10.5281/zenodo.3557929" httpUri="https://zenodo.org/record/3557929/files/figure.png" pageId="3" pageNumber="152">Fig. 1</figureCitation>
) are from a non-type specimen with the following labels: “
<collectingCountry id="31C576793C3D1235C55CF827FA0AF5B7" box="[1325,1432,621,647]" name="Brazil" pageId="3" pageNumber="152">BRAZIL</collectingCountry>
, State of
<collectingRegion id="8B16F80B3C3D1235C084F8DBFE18F59B" box="[245,394,657,683]" country="Brazil" name="Mato Grosso" pageId="3" pageNumber="152">Mato Grosso</collectingRegion>
, Cuiaba, Parque de Exposicao // HABITUS/GENIT. DIGITAL IMAGES captured 2019 P.D. Perkins //
<taxonomicName id="8ED24D6A3C3D1235C178F8FFFDE6F5FF" authority="Knisch 1921" authorityName="Knisch. Labels" authorityYear="1921" box="[265,628,693,719]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
Hydrochus argutus
<bibRefCitation id="2D434B183C3D1235C195F8FFFDE6F5FF" author="Knisch, A." box="[484,628,693,719]" pageId="3" pageNumber="152" pagination="1 - 24" refId="ref5602" refString="Knisch, A. (1921) Hydrophiliden aus Matto Grosso (Erster Teil). Archiv fur Naturegeschichte, 87 A, 1 - 24." type="journal article" year="1921">Knisch 1921</bibRefCitation>
</taxonomicName>
, det. P. D. Perkins 2019.” This specimen is deposited in the NMNH.
</paragraph>
</subSubSection>
<subSubSection id="01C865623C3D1235C0B6F893FE63F307" pageId="3" pageNumber="152" type="discussion">
<paragraph id="496D36E93C3D1235C0B6F893FE63F307" blockId="3.[151,1437,153,1079]" pageId="3" pageNumber="152">
<emphasis id="7BA6EAFB3C3D1235C0B6F893FEA6F5C3" bold="true" box="[199,308,729,755]" pageId="3" pageNumber="152">Remarks</emphasis>
.
<bibRefCitation id="2D434B183C3D1235C130F893FE42F5C3" author="Oliva, A." box="[321,464,729,755]" pageId="3" pageNumber="152" pagination="87 - 104" refId="ref6088" refString="Oliva, A. (1992) The species of Hydrochus (Coleoptera; Hydrophiloidea; Hydrochidae) described from South America. Bulletin et annales de la Societe royale belge d'entomologie, 128, 87 - 104." type="journal article" year="1992">Oliva (1992)</bibRefCitation>
provided a male genitalia drawing supposedly of
<taxonomicName id="8ED24D6A3C3D1235C387F890FBFFF5C3" authorityName="Knisch. Labels" authorityYear="1921" box="[1014,1133,730,755]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3D1235C387F890FBFFF5C3" box="[1014,1133,730,755]" italics="true" pageId="3" pageNumber="152">H. argutus</emphasis>
</taxonomicName>
, but unfortunately does not specify any data of the specimen that was used. The drawing is very different from the male genitalia of specimens examined in this study, or in any other published figures of Neotropical
<taxonomicName id="8ED24D6A3C3D1235C3BCF96BFBD5F40B" authorityName="Leach" authorityYear="1817" box="[973,1095,801,827]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="7BA6EAFB3C3D1235C3BCF96BFBD5F40B" box="[973,1095,801,827]" italics="true" pageId="3" pageNumber="152">Hydrochus</emphasis>
</taxonomicName>
male genitalia: the parameres in the drawing are very short, only about ½ the length of the aedeagus. I conclude that this drawing was made with inadequate magnification, using the damaged specimen noted above, which has the distal portions of both parameres sheared-off. I have examined the
<typeStatus id="9669884B3C3D1235C233F9C7FD35F497" box="[578,679,909,935]" pageId="3" pageNumber="152" type="holotype">holotype</typeStatus>
, and the
<typeStatus id="9669884B3C3D1235C37AF9C7FCFFF497" box="[779,877,909,935]" pageId="3" pageNumber="152" type="holotype">holotype</typeStatus>
male genitalia, of
<taxonomicName id="8ED24D6A3C3D1235C44FF9C7FEA4F4FB" authority="Makhan 1995" authorityName="Makhan" authorityYear="1995" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="johannapietersenae">
<emphasis id="7BA6EAFB3C3D1235C44FF9C7FA0EF497" box="[1086,1436,909,935]" italics="true" pageId="3" pageNumber="152">Hydrochus johannapietersenae</emphasis>
Makhan 1995
</taxonomicName>
, deposited in the NMW, and concur with
<bibRefCitation id="2D434B183C3D1235C37EF9FBFC32F4FC" author="Oliva, A." box="[783,928,945,972]" pageId="3" pageNumber="152" pagination="301 - 341" refId="ref6127" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
that it is a junior synonym of
<taxonomicName id="8ED24D6A3C3D1235C488F9FBFE13F4DF" authority="Knisch. Labels" authorityName="Knisch. Labels" authorityYear="1921" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
<emphasis id="7BA6EAFB3C3D1235C488F9FBFF46F4DF" italics="true" pageId="3" pageNumber="152">Hydrochus argutus</emphasis>
Knisch. Labels
</taxonomicName>
on
<typeStatus id="9669884B3C3D1235C1D8F99FFD99F4DF" box="[425,523,981,1007]" pageId="3" pageNumber="152" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8ED24D6A3C3D1235C25FF99CFCBEF4DF" authorityName="Makhan" authorityYear="1995" box="[558,812,981,1007]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="johannapietersenae">
<emphasis id="7BA6EAFB3C3D1235C25FF99CFCBEF4DF" box="[558,812,981,1007]" italics="true" pageId="3" pageNumber="152">H. johannapietersenae</emphasis>
</taxonomicName>
: “
<collectingCountry id="31C576793C3D1235C332F99FFC48F4DF" box="[835,986,981,1007]" name="Paraguay" pageId="3" pageNumber="152">PARAGUAY</collectingCountry>
, 16.20.10
<collectingRegion id="8B16F80B3C3D1235C43FF99FFB4BF4DF" box="[1102,1241,981,1007]" country="Paraguay" name="Guaira" pageId="3" pageNumber="152">Dep. Guaira</collectingRegion>
, 1992 Zorilla leg. U. Drechsel //
<taxonomicName id="8ED24D6A3C3D1235C14AF9B3FD06F323" authorityName="Makhan" authorityYear="1995" box="[315,660,1017,1043]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="johannapietersenae">Hydrochus johannapietersenae</taxonomicName>
det. D. Makhan 1995 //
<typeStatus id="9669884B3C3D1235C3D6F9B3FB9DF323" box="[935,1039,1017,1043]" pageId="3" pageNumber="152" type="holotype">Holotype</typeStatus>
[red] //
<taxonomicName id="8ED24D6A3C3D1235C41AF9B3FF5DF307" authority="Knisch, 1921" authorityName="Knisch. Labels" authorityYear="1921" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="argutus">
Hydrochus argutus
<bibRefCitation id="2D434B183C3D1235C537F9B3FF5DF307" author="Knisch, A." pageId="3" pageNumber="152" pagination="1 - 24" refId="ref5602" refString="Knisch, A. (1921) Hydrophiliden aus Matto Grosso (Erster Teil). Archiv fur Naturegeschichte, 87 A, 1 - 24." type="journal article" year="1921">Knisch, 1921</bibRefCitation>
</taxonomicName>
det. P. D. Perkins, 2019”.
</paragraph>
</subSubSection>
</treatment>
</document>