treatments-xml/data/03/C9/76/03C97661FFF2FFEE629DFC7DFDF7F870.xml

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<mods:title id="E45FCF30879B34C586AAA451B6FD7E35">A new species of Dactyloa from eastern Panama, with comments on other Dactyloa species present in the region</mods:title>
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<mods:namePart id="343BBCD162D3109684ED883A67FB3544">Lotzkat, Sebastian</mods:namePart>
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<taxonomicName id="4C60BCF4FFF2FFEB629DFC7DFE9AFC1E" ID-CoL="6BYLB" box="[151,326,943,969]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia" status="sp. nov.">
<emphasis id="B9141B65FFF2FFEB629DFC7DFE9AFC1E" bold="true" box="[151,326,943,969]" italics="true" pageId="8" pageNumber="65">Dactyloa maia</emphasis>
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<figureCitation id="135BDBF2FFF2FFEB629DFC00FEDCFC3C" box="[151,256,978,1003]" captionStart="FIGURE 1" captionStartId="3.[151,250,1642,1664]" captionTargetBox="[165,1423,211,1593]" captionTargetId="figure@3.[150,1437,193,1620]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Dactyloa species formerly referred to as D. chocorum in eastern Panama, based on specimens collected for this study, provided by A. Sosa, literature and online databases (http: // www. vertnet. org /)." httpUri="https://zenodo.org/record/289963/files/figure.png" pageId="8" pageNumber="65">Figures 1</figureCitation>
, 
<figureCitation id="135BDBF2FFF2FFEB6304FC01FEC0FC3C" box="[270,284,979,1003]" captionStart="FIGURE 4" captionStartId="11.[151,250,1913,1935]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@11.[174,1459,193,1889]" captionTargetPageId="11" captionText="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="8" pageNumber="65">4</figureCitation>
E–F, 5E–F, 6D, 8, 9, 10.
</paragraph>
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.
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97268, adult male (
<figureCitation id="135BDBF2FFF2FFEB6022FBC8FDAAFBE4" box="[552,630,1050,1075]" captionStart="FIGURE 4" captionStartId="11.[151,250,1913,1935]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@11.[174,1459,193,1889]" captionTargetPageId="11" captionText="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="8" pageNumber="65">Figs. 4</figureCitation>
E, 5E, 6D, 9), from the ridge of the Serranía de Darién (
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) along the trail that connects the Comarca Wargandí and the Comarca Guna Yala, about 
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northeast of the village Nurra, 
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, 
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, 
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asl., Corregimiento de Nurra, Comarca Wargandí, 
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; collected by Abel Batista and Milan Vesely on 0 
<date id="FFDEE1B7FFF2FFEB63E3FB55FD45FB48" box="[489,665,1158,1183]" pageId="8" pageNumber="65" value="2012-10-03">3 October 2012</date>
; original field number AB 760.
</paragraph>
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.
</emphasis>
All from 
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: 
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97269, a male, from Cerro la Javillosa Ambroya, Torti, Chepo, 
<collectingCountry id="F37787E7FFF2FFEB6749FB79FA40FB15" box="[1347,1436,1195,1218]" name="Panama" pageId="8" pageNumber="65">Panama</collectingCountry>
province, collected on 
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, 19:39 hrs, 
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, 
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, 
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asl, collected by Abel Batista and Milan Vesely; 
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2782, a female, same collecting data as 
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; 
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2781 and 
<collectionCode id="ED715FB2FFF2FFEB6742FB21FA40FADC" box="[1352,1436,1267,1291]" pageId="8" pageNumber="65">MHCH</collectionCode>
2783, females, respectively, from Cerro Pechito Parado, Bajo pequeño, Lajas blancas, Cémaco, Comarca Emberá- Wounáan, on 0 
<date id="FFDEE1B7FFF2FFEB634BFAE9FDCCFA84" box="[321,528,1338,1363]" pageId="8" pageNumber="65" value="2012-11-07">7 November 2012</date>
, 
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, 
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, 
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asl, collected by Abel Batista; 
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97270, a female from la Cascada trail, Burbayar private reservation, Cartí, Narganá, Comarca Guna Yala, on 
<date id="FFDEE1B7FFF2FFEB6708FA8CFF0EFA4C" pageId="8" pageNumber="65" value="2012-11-26">26 November 2012</date>
, 
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, 
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, 
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asl, collected by Abel Batista and Konrad Mebert.
</paragraph>
</subSubSection>
<subSubSection id="C37A94FCFFF2FFEA62CDFA74FAD1FE97" lastPageId="9" lastPageNumber="66" pageId="8" pageNumber="65" type="diagnosis">
<paragraph id="8BDFC777FFF2FFEA62CDFA74FAD1FE97" blockId="8.[151,1436,1050,2011]" lastBlockId="9.[151,1437,151,1977]" lastPageId="9" lastPageNumber="66" pageId="8" pageNumber="65">
<emphasis id="B9141B65FFF2FFEB62CDFA74FE9DFA68" bold="true" box="[199,321,1446,1471]" pageId="8" pageNumber="65">Diagnosis.</emphasis>
A moderate-sized species (SVL 
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) of the genus 
<emphasis id="B9141B65FFF2FFEB61FCFA74FB2FFA69" box="[1014,1267,1446,1470]" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C60BCF4FFF2FFEB61FCFA74FBBCFA69" box="[1014,1120,1446,1470]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="genus">Dactyloa</taxonomicName>
, 
<taxonomicName id="4C60BCF4FFF2FFEB6665FA75FB2FFA69" box="[1135,1267,1446,1470]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="latifrons">D. latifrons</taxonomicName>
</emphasis>
species group (
<emphasis id="B9141B65FFF2FFEB62AAFA19FF02FA35" box="[160,222,1483,1506]" italics="true" pageId="8" pageNumber="65">sensu</emphasis>
<bibRefCitation id="EFF1BA86FFF2FFEB62E2FA18FE39FA34" author="Nicholson" box="[232,485,1482,1507]" pageId="8" pageNumber="65" refString="Nicholson, K. E., Crother, B. I., Guyer, C. &amp; Savage, J. M. (2012) It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa, 3477, 1 - 108." type="journal article" year="2012">
Nicholson 
<emphasis id="B9141B65FFF2FFEB636FFA19FE7CFA34" box="[357,416,1483,1507]" italics="true" pageId="8" pageNumber="65">et al.</emphasis>
2012
</bibRefCitation>
), that is most similar in external morphology to 
<taxonomicName id="4C60BCF4FFF2FFEB6613FA19FB14FA35" box="[1049,1224,1483,1506]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pupurescens">
<emphasis id="B9141B65FFF2FFEB6613FA19FB14FA35" box="[1049,1224,1483,1506]" italics="true" pageId="8" pageNumber="65">D. pupurescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C60BCF4FFF2FFEB66D3FA19FAE2FA34" box="[1241,1342,1483,1507]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF2FFEB66D3FA19FAE2FA34" box="[1241,1342,1483,1507]" italics="true" pageId="8" pageNumber="65">D. limon</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C60BCF4FFF2FFEB678BFA19FF34F9D1" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF2FFEB678BFA19FF34F9D1" italics="true" pageId="8" pageNumber="65">D. ibanezi</emphasis>
</taxonomicName>
, and according to molecular evidence is most closely related to 
<taxonomicName id="4C60BCF4FFF2FFEB61D9FA3DFB5FF9D1" box="[979,1155,1519,1542]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pupurescens">
<emphasis id="B9141B65FFF2FFEB61D9FA3DFB5FF9D1" box="[979,1155,1519,1542]" italics="true" pageId="8" pageNumber="65">D. pupurescens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C60BCF4FFF2FFEB66B5FA3DFAE9F9D1" box="[1215,1333,1518,1542]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF2FFEB66B5FA3DFAE9F9D1" box="[1215,1333,1518,1542]" italics="true" pageId="8" pageNumber="65">D. ibanezi</emphasis>
</taxonomicName>
(
<figureCitation id="135BDBF2FFF2FFEB674CFA3CFA51F9D0" box="[1350,1421,1518,1543]" captionStart="FIGURE 7" captionStartId="14.[151,250,1725,1747]" captionTargetBox="[185,1338,233,1673]" captionTargetId="figure@14.[185,1383,225,1677]" captionTargetPageId="14" captionText="FIGURE 7. Bayesian consensus tree of the genus Dactyloa based on 16 S and COI mtDNA. Outgroups: Norops capito, Norops limifrons, Norops tropidogaster, Norops poecilopus and Polychrus marmoratus. Asterisks on nodes indicate estimated posterior probabilities P ≥ 0.90. Scale bar refers to substitutions per site." httpUri="https://zenodo.org/record/289969/files/figure.png" pageId="8" pageNumber="65">Fig. 7</figureCitation>
). These four species share a moderate adult size (SVL 
<quantity id="4C986A92FFF2FFEB60E0F9C0FCBBF9FC" box="[746,871,1554,1579]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.8" metricValueMin="6.2" pageId="8" pageNumber="65" unit="mm" value="75.0" valueMax="88.0" valueMin="62.0">62–88 mm</quantity>
); a large dewlap; a peculiar flank pattern in males, which is green with dark oblique bands, or blotches, or ocelli always arranged in oblique rows (
<figureCitation id="135BDBF2FFF2FFEB66D5F9E4FAF8F998" box="[1247,1316,1590,1615]" captionStart="FIGURE 5" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[166,1436,193,1888]" captionTargetId="figure@12.[166,1451,193,1889]" captionTargetPageId="12" captionText="FIGURE 5. Flank pattern in species of Dactyloa related to D. purpurescens. Arrows indicate capitad direction for all males (left column) and females (right column). A) D. limon holotype MHUA 11760; B) D. limon MHUA 11248; C) D. purpurescens (MHCH 2636); D) D. purpurescens (SMF 91475); E) D. maia (SMF 97268); F) D. maia (MHCH 2782); G) D. ibanezi, MHCH 2184; H) D. ibanezi, SMF 91475. Scale bars equal 10 mm." httpUri="https://zenodo.org/record/289967/files/figure.png" pageId="8" pageNumber="65">Fig. 5</figureCitation>
); enlarged postcloacal scales in males; and smooth ventrals. 
<taxonomicName id="4C60BCF4FFF2FFEB60D6F989FC59F9A5" box="[732,901,1627,1651]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF2FFEB60D6F989FC59F9A5" box="[732,901,1627,1651]" italics="true" pageId="8" pageNumber="65">Dactyloa maia</emphasis>
</taxonomicName>
can readily be distinguished from these three species by its color pattern and morphology (
<figureCitation id="135BDBF2FFF2FFEB6094F9ACFCD1F940" box="[670,781,1662,1687]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="10.[151,250,1925,1947]" captionStartId-1="11.[151,250,1913,1935]" captionStartId-2="12.[151,250,1919,1941]" captionStartId-3="13.[151,250,973,995]" captionTargetBox-0="[174,1428,193,1888]" captionTargetBox-1="[174,1428,193,1888]" captionTargetBox-2="[166,1436,193,1888]" captionTargetBox-3="[217,1369,193,951]" captionTargetId-0="figure@10.[174,1459,193,1889]" captionTargetId-1="figure@11.[174,1459,193,1889]" captionTargetId-2="figure@12.[166,1451,193,1889]" captionTargetId-3="figure@13.[217,1369,193,952]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 3. Some species of Dactyloa found in Eastern Panama. A – B) D. purpurescens (female and male, MHCH 2635 – 36); C – D) D. kunayalae (male, SMF 97266); E) D. frenata (male, MHCH 2785); F) D. latifrons (young male, SMF 96575); G – H) D. chloris (female, SMF 97096)." captionText-1="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." captionText-2="FIGURE 5. Flank pattern in species of Dactyloa related to D. purpurescens. Arrows indicate capitad direction for all males (left column) and females (right column). A) D. limon holotype MHUA 11760; B) D. limon MHUA 11248; C) D. purpurescens (MHCH 2636); D) D. purpurescens (SMF 91475); E) D. maia (SMF 97268); F) D. maia (MHCH 2782); G) D. ibanezi, MHCH 2184; H) D. ibanezi, SMF 91475. Scale bars equal 10 mm." captionText-3="FIGURE 6. Lateral head views of the species of Dactyloa related to D. purpurescens. A) D. limon holotype MHUA 11760; B) D. ibanezi MHCH 2184; C) D. purpurescens SMF 97271; D) D. maia sp. nov. holotype SMF 97268. Scale bars equal 10 mm" httpUri-0="https://zenodo.org/record/289965/files/figure.png" httpUri-1="https://zenodo.org/record/289966/files/figure.png" httpUri-2="https://zenodo.org/record/289967/files/figure.png" httpUri-3="https://zenodo.org/record/289968/files/figure.png" pageId="8" pageNumber="65">Figs. 3–6</figureCitation>
; Table 1–2), and from the remaining species of 
<taxonomicName id="4C60BCF4FFF2FFEB673FF9ACFA40F941" box="[1333,1436,1662,1686]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="B9141B65FFF2FFEB673FF9ACFA40F941" box="[1333,1436,1662,1686]" italics="true" pageId="8" pageNumber="65">Dactyloa</emphasis>
</taxonomicName>
within the 
<taxonomicName id="4C60BCF4FFF2FFEB6311F971FE7DF96C" box="[283,417,1699,1723]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B9141B65FFF2FFEB6311F971FE7DF96C" box="[283,417,1699,1723]" italics="true" pageId="8" pageNumber="65">D. latifrons</emphasis>
</taxonomicName>
group by its moderate size (SVL &lt;
<quantity id="4C986A92FFF2FFEB6169F971FC16F96C" box="[867,970,1699,1723]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="65" unit="mm" value="100.0">100 mm</quantity>
); and the orange male dewlap with an uninterrupted white margin. 
<taxonomicName id="4C60BCF4FFF2FFEB63E8F914FD50F909" box="[482,652,1734,1758]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF2FFEB63E8F914FD50F909" box="[482,652,1734,1758]" italics="true" pageId="8" pageNumber="65">Dactyloa maia</emphasis>
</taxonomicName>
can be differentiated from 
<taxonomicName id="4C60BCF4FFF2FFEB61DAF915FBEAF909" box="[976,1078,1734,1758]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF2FFEB61DAF915FBEAF909" box="[976,1078,1734,1758]" italics="true" pageId="8" pageNumber="65">D. limon</emphasis>
</taxonomicName>
by its male dewlap coloration which is orange with an uninterrupted white margin (vs. yellowish near the throat and tan on distal portion, or uniformly light tan in 
<emphasis id="B9141B65FFF2FFEB6396F8DDFDD6F8F1" box="[412,522,1806,1830]" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C60BCF4FFF2FFEB6396F8DDFDDFF8F1" box="[412,515,1806,1830]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="limon">D. limon</taxonomicName>
;
</emphasis>
<figureCitation id="135BDBF2FFF2FFEB601EF8DCFD80F8F0" box="[532,604,1806,1831]" captionStart="FIGURE 4" captionStartId="11.[151,250,1913,1935]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@11.[174,1459,193,1889]" captionTargetPageId="11" captionText="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="8" pageNumber="65">Fig. 4</figureCitation>
). 
<taxonomicName id="4C60BCF4FFF2FFEB607EF8DCFCC1F8F1" box="[628,797,1806,1830]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF2FFEB607EF8DCFCC1F8F1" box="[628,797,1806,1830]" italics="true" pageId="8" pageNumber="65">Dactyloa maia</emphasis>
</taxonomicName>
further differs from 
<emphasis id="B9141B65FFF2FFEB661BF8DDFA98F8F1" box="[1041,1348,1806,1830]" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C60BCF4FFF2FFEB661BF8DDFB0CF8F1" box="[1041,1232,1807,1830]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="purpurescens">D. purpurescens</taxonomicName>
, 
<taxonomicName id="4C60BCF4FFF2FFEB66D4F8DDFA98F8F1" box="[1246,1348,1806,1830]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="limon">D. limon</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C60BCF4FFF2FFEB678BF8DDFF34F89C" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF2FFEB678BF8DDFF34F89C" italics="true" pageId="8" pageNumber="65">D. ibanezi</emphasis>
</taxonomicName>
in the color pattern of the flanks as follows: 
<taxonomicName id="4C60BCF4FFF2FFEB60E8F8E1FC54F89D" box="[738,904,1843,1867]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF2FFEB60E8F8E1FC54F89D" box="[738,904,1843,1867]" italics="true" pageId="8" pageNumber="65">Dactyloa maia</emphasis>
</taxonomicName>
has oblique rows of turquoise ocelli or oblique dark green bands without sexual dimorphism (
<figureCitation id="135BDBF2FFF2FFEB6094F884FD3DF8B8" box="[670,737,1878,1903]" captionStart="FIGURE 8" captionStartId="15.[151,250,1903,1925]" captionTargetBox="[151,1436,197,1891]" captionTargetId="figure@15.[151,1436,197,1893]" captionTargetPageId="15" captionText="FIGURE 8. Color variation of Dactyloa maia sp. nov. in life. A – B) female MHCH 2782; C – D) female MHCH 2781; E – F) male SMF 97269; G) female SMF 97270." httpUri="https://zenodo.org/record/289970/files/figure.png" pageId="8" pageNumber="65">Fig. 8</figureCitation>
); in 
<taxonomicName id="4C60BCF4FFF2FFEB611FF885FCABF8B9" box="[789,887,1878,1902]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF2FFEB611FF885FCABF8B9" box="[789,887,1878,1902]" italics="true" pageId="8" pageNumber="65">D. limon</emphasis>
</taxonomicName>
, males have wide dark green bands on the flanks whereas females have diffuse dark green spots distributed evenly or randomly; males of 
<taxonomicName id="4C60BCF4FFF2FFEB6686F8A9FA99F845" box="[1164,1349,1915,1938]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF2FFEB6686F8A9FA99F845" box="[1164,1349,1915,1938]" italics="true" pageId="8" pageNumber="65">D. purpurescens</emphasis>
</taxonomicName>
exhibit oblique rows of ocelli or blotches whereas females have dark green spots arranged in oblique rows; in 
<taxonomicName id="4C60BCF4FFF2FFEB6722F84DFA40F861" box="[1320,1436,1950,1974]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF2FFEB6722F84DFA40F861" box="[1320,1436,1950,1974]" italics="true" pageId="8" pageNumber="65">D. ibanezi</emphasis>
</taxonomicName>
both sexes exhibit oblique thin black lines. The hemipenis of 
<taxonomicName id="4C60BCF4FFF2FFEB6141F811FC7AF80D" box="[843,934,1987,2010]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF2FFEB6141F811FC7AF80D" box="[843,934,1987,2010]" italics="true" pageId="8" pageNumber="65">D. maia</emphasis>
</taxonomicName>
is a small, unilobate organ (slightly bilobate in 
<emphasis id="B9141B65FFF3FFEA62BCFF4BFEE8FF67" box="[182,308,152,176]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C60BCF4FFF3FFEA62BCFF4BFEF1FF67" box="[182,301,152,176]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="ibanezi">D. ibanezi</taxonomicName>
;
</emphasis>
no information available for 
<taxonomicName id="4C60BCF4FFF3FFEA608DFF4BFD30FF67" box="[647,748,152,176]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF3FFEA608DFF4BFD30FF67" box="[647,748,152,176]" italics="true" pageId="9" pageNumber="66">D. limon</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C60BCF4FFF3FFEA612CFF4BFC3DFF67" box="[806,993,153,176]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF3FFEA612CFF4BFC3DFF67" box="[806,993,153,176]" italics="true" pageId="9" pageNumber="66">D. purpurescens</emphasis>
</taxonomicName>
). Also, 
<taxonomicName id="4C60BCF4FFF3FFEA6635FF4AFB3AFF78" box="[1087,1254,152,176]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF3FFEA6635FF4AFB3AFF78" box="[1087,1254,152,176]" italics="true" pageId="9" pageNumber="66">Dactyloa maia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C60BCF4FFF3FFEA678BFF4BFEF1FF03" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF3FFEA678BFF4BFEF1FF03" italics="true" pageId="9" pageNumber="66">D. purpurescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C60BCF4FFF3FFEA6335FF6FFE7AFF03" box="[319,422,188,212]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF3FFEA6335FF6FFE7AFF03" box="[319,422,188,212]" italics="true" pageId="9" pageNumber="66">D. limon</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C60BCF4FFF3FFEA63E6FF6FFDBFFF03" box="[492,611,188,212]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF3FFEA63E6FF6FFDBFFF03" box="[492,611,188,212]" italics="true" pageId="9" pageNumber="66">D. ibanezi</emphasis>
</taxonomicName>
in mean values of several morphological characters as follows (mean values for 
<taxonomicName id="4C60BCF4FFF3FFEA6305FF33FE14FF2F" box="[271,456,225,248]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF3FFEA6305FF33FE14FF2F" box="[271,456,225,248]" italics="true" pageId="9" pageNumber="66">D. purpurescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C60BCF4FFF3FFEA63DDFF33FDE6FF2F" box="[471,570,224,248]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF3FFEA63DDFF33FDE6FF2F" box="[471,570,224,248]" italics="true" pageId="9" pageNumber="66">D. limon</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C60BCF4FFF3FFEA6073FF33FD31FF2F" box="[633,749,224,248]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF3FFEA6073FF33FD31FF2F" box="[633,749,224,248]" italics="true" pageId="9" pageNumber="66">D. ibanezi</emphasis>
</taxonomicName>
presented in that order): HL/SVL 
<quantity id="4C986A92FFF3FFEA667DFF32FB1AFF2F" box="[1143,1222,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="6.858" pageId="9" pageNumber="66" unit="in" value="0.27">0.27 in</quantity>
<taxonomicName id="4C60BCF4FFF3FFEA66C7FF32FAA8FF20" box="[1229,1396,224,248]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF3FFEA66C7FF32FAA8FF20" box="[1229,1396,224,248]" italics="true" pageId="9" pageNumber="66">Dactyloa maia</emphasis>
</taxonomicName>
vs. 0.25, 0.23 and 0.26,.LST 
<quantity id="4C986A92FFF3FFEA63B9FED6FDDDFECA" box="[435,513,260,285]" metricMagnitude="0" metricUnit="m" metricValue="1.64338" pageId="9" pageNumber="66" unit="in" value="64.7">64.7 in</quantity>
<taxonomicName id="4C60BCF4FFF3FFEA600DFED6FD70FECB" box="[519,684,260,284]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF3FFEA600DFED6FD70FECB" box="[519,684,260,284]" italics="true" pageId="9" pageNumber="66">Dactyloa maia</emphasis>
</taxonomicName>
vs. 62.8, 43.7, and 44.5; number of scales between first and second canthals 14.5/12.3 (first canthals/second canthals) in 
<taxonomicName id="4C60BCF4FFF3FFEA60E3FEFAFC52FEE8" box="[745,910,296,320]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF3FFEA60E3FEFAFC52FEE8" box="[745,910,296,320]" italics="true" pageId="9" pageNumber="66">Dactyloa maia</emphasis>
</taxonomicName>
vs 12.9/10.2, 10/8.6, and 11.5/10.
</paragraph>
</subSubSection>
<subSubSection id="C37A94FCFFF3FFEE62CDFE9EFD8AF9CC" lastPageId="13" lastPageNumber="70" pageId="9" pageNumber="66" type="description">
<paragraph id="8BDFC777FFF3FFEA62CDFE9EFC24FA4A" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="66">
<emphasis id="B9141B65FFF3FFEA62CDFE9EFDC5FEB2" bold="true" box="[199,537,332,357]" pageId="9" pageNumber="66">
Description of the 
<typeStatus id="54DB79D5FFF3FFEA63A1FE9EFDC8FEB2" box="[427,532,332,357]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male as indicated by everted hemipenes, a pair of enlarged postcloacal scales, and presence of a large dewlap (
<figureCitation id="135BDBF2FFF3FFEA605CFEBDFD7FFE5F" box="[598,675,367,392]" captionStart="FIGURE 3" captionStartId="10.[151,250,1925,1947]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@10.[174,1459,193,1889]" captionTargetPageId="10" captionText="FIGURE 3. Some species of Dactyloa found in Eastern Panama. A – B) D. purpurescens (female and male, MHCH 2635 – 36); C – D) D. kunayalae (male, SMF 97266); E) D. frenata (male, MHCH 2785); F) D. latifrons (young male, SMF 96575); G – H) D. chloris (female, SMF 97096)." httpUri="https://zenodo.org/record/289965/files/figure.png" pageId="9" pageNumber="66">Figs. 3</figureCitation>
E, 4E, 7D, 9); SVL 
<quantity id="4C986A92FFF3FFEA618DFEBDFC06FE50" box="[903,986,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="66" unit="mm" value="76.0">76 mm</quantity>
; tail length 
<quantity id="4C986A92FFF3FFEA6655FEA2FB61FE50" box="[1119,1213,368,392]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="66" unit="mm" value="178.0">178 mm</quantity>
(tail complete), tail length/SVL ratio 2.34; tail laterally compressed in cross section, tail height 
<quantity id="4C986A92FFF3FFEA61F0FE46FB88FE7B" box="[1018,1108,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="66" unit="mm" value="2.2">2.2 mm</quantity>
, tail width 2.0 mm; axilla to groin distance 
<quantity id="4C986A92FFF3FFEA6348FE65FE70FE18" box="[322,428,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="9" pageNumber="66" unit="mm" value="35.6">35.6 mm</quantity>
; head length 
<quantity id="4C986A92FFF3FFEA6041FE6AFD68FE18" box="[587,692,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="9" pageNumber="66" unit="mm" value="19.2">19.2 mm</quantity>
, HL/SVL ratio 0.25; snout length 
<quantity id="4C986A92FFF3FFEA664DFE6AFB78FE18" box="[1095,1188,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="66" unit="mm" value="9.3">9.3 mm</quantity>
; head width 
<quantity id="4C986A92FFF3FFEA6731FE6AFA44FE18" box="[1339,1432,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="9" pageNumber="66" unit="mm" value="9.7">9.7 mm</quantity>
; longest toe of adpressed hind limb reaching posterior margin of orbit; shank length 
<quantity id="4C986A92FFF3FFEA6653FE0EFB1DFE23" box="[1113,1217,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="9" pageNumber="66" unit="mm" value="20.7">20.7 mm</quantity>
, shank length/SVL ratio 0.27, shank length/HL ratio 1.08; tip of longest finger of extended forelimb reaching tip of snout; longest finger of adpressed forelimb not reaching to anterior insertion of hind limbs; prefrontal ridges distinct, parietal ridges conspicuous; scales on snout mostly keeled; 6 postrostrals; 7 scales between nasals; scales in distinct prefrontal depression smooth; supraorbital semicircles differentiated, composed of smooth scales, separated by a minimum of 2 scales; supraorbital disc composed of 7 enlarged smooth scales; two elongated, smooth anterior superciliaries, followed posteriorly by a much smaller, elongate scale; about 5 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; interparietal plate distinct, parietal eye visible; canthal ridge distinct, composed of 3 large (posterior) and 7 small (anterior) canthal scales; 11 scales present between second canthals; 13 scales present between posterior canthals; 78 loreal scales arranged in 7 horizontal rows; subocular scales flat, subocular row well-defined; 9 supralabials to level below center of eye; ear opening 0.73 x 
<quantity id="4C986A92FFF3FFEA629DFCB5FF2FFCA8" box="[151,243,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="66" unit="mm" value="1.6">1.6 mm</quantity>
(length x height); mental distinctly wider than long, almost completely divided medially, bordered posteriorly by 6 postmentals; 10 infralabials to level below center of eye; third and fourth sublabials posterior to mental slightly enlarged; keeled granular to elongate scales present on chin and throat; dewlap large, extending well onto body, anterior insertion is about halfway between nose and orbit, posterior insertion at a level between one-third and one half of the distance between axilla and groin, with about 4 gorgetal-sternal rows, each 2–3 scales wide, becoming less regular posteriorly; low nuchal crest present, dorsal crest barely visible; dorsum of body with keeled scales, 1–2 middorsal rows of prominently keeled, but not otherwise enlarged scales; about 50 medial dorsal scales in one HL; about 125 medial dorsal scales between levels of axilla and groin; lateral scales small, long and keeled; ventrals at midbody smooth, subimbricate; about 66 ventral scales in one HL; about 87 ventral scales between axilla and groin; about 164 scales around midbody; caudal scales strongly keeled, without whorls of enlarged scales, subcaudal scales with a single prominent keel; a pair of greatly enlarged postcloacal scales, larger one about 0.88 x 
<quantity id="4C986A92FFF3FFEA6353FB26FE61FADB" box="[345,445,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="66" unit="mm" value="1.9">1.90 mm</quantity>
(length x width); tube-like axillary pocket not developed; scales on anterodorsal surface of thigh and on dorsal surface of forearm keeled; digital pads dilated, dilated pad about 3 times width of nondilated scales under distal phalanx; distal phalanx narrower than and raised from dilated pad; 31/31 (left/right) lamellae under phalanges ii to iv of 4th toe; 11/12 scales under distal phalanx of 4th toe; 21/19 lamellae under phalanges ii to iv of 4th finger; 10/9 scales under distal phalanx of 4th finger.
</paragraph>
<paragraph id="8BDFC777FFF3FFEA62CDFA75FB16F9FA" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="66">
<emphasis id="B9141B65FFF3FFEA62CDFA75FE3DFA17" bold="true" box="[199,481,1447,1472]" pageId="9" pageNumber="66">Hemipenis description:</emphasis>
The completely everted hemipenis (
<figureCitation id="135BDBF2FFF3FFEA6185FA75FC39FA17" box="[911,997,1447,1472]" captionStart="FIGURE 10" captionStartId="17.[151,250,790,812]" captionTargetBox="[151,1435,176,768]" captionTargetId="figure@17.[151,1435,159,769]" captionTargetPageId="17" captionText="FIGURE 10. Hemipenis of the Paratype (SMF 97269) of Dactyloa maia sp. nov. A) Sulcate view; B) asulcate view; C) lateral view." httpUri="https://zenodo.org/record/289972/files/figure.png" pageId="9" pageNumber="66">Fig. 10</figureCitation>
) of SMF 97269 is a small, unilobate organ; sulcus spermaticus bordered by well-developed sulcal lips, opening at base of apex into a small concave area; large asulcate processus and ridge present; a prominent fleshy fringe present on each lateral side of truncus; most of apex on asulcate side and distal portion finely calyculate, truncus with transverse folds..
</paragraph>
<paragraph id="8BDFC777FFF3FFEA62CDF9E5FB59F8D2" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="66">
<emphasis id="B9141B65FFF3FFEA62CDF9E5FD86F987" bold="true" box="[199,602,1591,1616]" pageId="9" pageNumber="66">
Coloration in life of the 
<typeStatus id="54DB79D5FFF3FFEA63E6F9E5FD89F987" box="[492,597,1591,1616]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
.
</emphasis>
(
<figureCitation id="135BDBF2FFF3FFEA6061F9E5FD72F987" box="[619,686,1591,1616]" captionStart="FIGURE 4" captionStartId="11.[151,250,1913,1935]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@11.[174,1459,193,1889]" captionTargetPageId="11" captionText="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="9" pageNumber="66">Fig. 4</figureCitation>
E) Dorsal and lateral ground color of body and limbs Light Grass Green (109); lateral surface of body with three Parrot Green (121) oblique bands directed backwards as well as downwards, each with six to seven Medium Blue (168) small oval spots; tail with Dark Green (136) transverse bands; eyelids Cream Yellow (82); iris Chrome Orange (74); ventral surfaces dirty white, suffused with Pale Cyan (157); dewlap Light Pratt’s Rufous (71) with three well defined longitudinal series of Emerald Green (143) scales, free margin of dewlap dirty white, anterior insertion of dewlap suffused with Beige (254).
</paragraph>
<paragraph id="8BDFC777FFF3FFEA62CDF8DDFD3DF86E" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="66">
<emphasis id="B9141B65FFF3FFEA62CDF8DDFB33F8FF" bold="true" box="[199,1263,1807,1832]" pageId="9" pageNumber="66">
Coloration of the 
<typeStatus id="54DB79D5FFF3FFEA6390F8DDFDDDF8FF" box="[410,513,1807,1832]" pageId="9" pageNumber="66" type="holotype">holotype</typeStatus>
after approximately two years of preservation in 70% ethanol.
</emphasis>
(
<figureCitation id="135BDBF2FFF3FFEA66F4F8DDFA9EF8FF" box="[1278,1346,1807,1832]" captionStart="FIGURE 9" captionStartId="16.[151,250,1927,1949]" captionTargetBox="[166,1436,193,1888]" captionTargetId="figure@16.[166,1451,193,1889]" captionTargetPageId="16" captionText="FIGURE 9. Dactyloa maia sp. nov. holotype (SMF 97268), preserved specimen. A) head, laterally; B) head, dorsally; C) head, ventrally; D) venter; E) dorsum; F) flank; G) left hand; H) left foot." httpUri="https://zenodo.org/record/289971/files/figure.png" pageId="9" pageNumber="66">Fig. 9</figureCitation>
) Dorsal and lateral ground color of body and limbs Lavender (195); lateral surfaces of body with three oblique rows of small oval Medium Water Blue (182) spots directed backwards as well as downwards; tail with Plumbeous (295) transverse bands; ventral surfaces dirty white, suffused with Medium Blue Gray (194); dewlap Pale Sulfur Yellow (92), its base suffused with Pale Neutral Gray (296).
</paragraph>
<caption id="DF1F97FFFFF0FFE9629DF857FE38F80F" httpUri="https://zenodo.org/record/289965/files/figure.png" pageId="10" pageNumber="67" targetBox="[174,1428,193,1888]" targetPageId="10">
<paragraph id="8BDFC777FFF0FFE9629DF857FE38F80F" blockId="10.[151,1436,1925,2008]" pageId="10" pageNumber="67">
<emphasis id="B9141B65FFF0FFE9629DF857FECFF84D" bold="true" box="[151,275,1925,1947]" pageId="10" pageNumber="67">FIGURE 3.</emphasis>
Some species of 
<taxonomicName id="4C60BCF4FFF0FFE963CCF857FDFFF84C" box="[454,547,1925,1947]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="genus">
<emphasis id="B9141B65FFF0FFE963CCF857FDFFF84C" box="[454,547,1925,1947]" italics="true" pageId="10" pageNumber="67">Dactyloa</emphasis>
</taxonomicName>
found in Eastern Panama. A–B) 
<taxonomicName id="4C60BCF4FFF0FFE9617FF854FBC1F84C" box="[885,1053,1926,1947]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF0FFE9617FF854FBC1F84C" box="[885,1053,1926,1947]" italics="true" pageId="10" pageNumber="67">D. purpurescens</emphasis>
</taxonomicName>
(female and male, MHCH 2635–36); C–D) 
<taxonomicName id="4C60BCF4FFF0FFE962DCF877FE82F86D" box="[214,350,1956,1978]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="kunayalae">
<emphasis id="B9141B65FFF0FFE962DCF877FE82F86D" box="[214,350,1956,1978]" italics="true" pageId="10" pageNumber="67">D. kunayalae</emphasis>
</taxonomicName>
(male, SMF 97266); E) 
<taxonomicName id="4C60BCF4FFF0FFE96050F877FD1EF86D" box="[602,706,1956,1978]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="frenata">
<emphasis id="B9141B65FFF0FFE96050F877FD1EF86D" box="[602,706,1956,1978]" italics="true" pageId="10" pageNumber="67">D. frenata</emphasis>
</taxonomicName>
(male, MHCH 2785); F
<emphasis id="B9141B65FFF0FFE961B0F876FBE3F86D" box="[954,1087,1956,1978]" italics="true" pageId="10" pageNumber="67">
) 
<taxonomicName id="4C60BCF4FFF0FFE961C0F877FBE3F86D" box="[970,1087,1956,1978]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="latifrons">D. latifrons</taxonomicName>
</emphasis>
(young male, SMF 96575); G–H) 
<taxonomicName id="4C60BCF4FFF0FFE9629DF811FF20F80F" box="[151,252,1986,2008]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="67" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B9141B65FFF0FFE9629DF811FF20F80F" box="[151,252,1986,2008]" italics="true" pageId="10" pageNumber="67">D. chloris</emphasis>
</taxonomicName>
(female, SMF 97096).
</paragraph>
</caption>
<caption id="DF1F97FFFFF1FFE8629DF8ABFE6EF83C" httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="11" pageNumber="68" targetBox="[174,1428,193,1888]" targetPageId="11">
<paragraph id="8BDFC777FFF1FFE8629DF8ABFE6EF83C" blockId="11.[151,1437,1913,2027]" pageId="11" pageNumber="68">
<emphasis id="B9141B65FFF1FFE8629DF8ABFEC9F859" bold="true" box="[151,277,1913,1935]" pageId="11" pageNumber="68">FIGURE 4.</emphasis>
Dewlaps of the 
<taxonomicName id="4C60BCF4FFF1FFE863CCF8ABFDFFF858" box="[454,547,1913,1935]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="B9141B65FFF1FFE863CCF8ABFDFFF858" box="[454,547,1913,1935]" italics="true" pageId="11" pageNumber="68">Dactyloa</emphasis>
</taxonomicName>
spp. related to 
<taxonomicName id="4C60BCF4FFF1FFE860C3F8A8FCAEF858" box="[713,882,1914,1935]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF1FFE860C3F8A8FCAEF858" box="[713,882,1914,1935]" italics="true" pageId="11" pageNumber="68">D. purpurescens</emphasis>
</taxonomicName>
(left column males, right column females). A–B) 
<taxonomicName id="4C60BCF4FFF1FFE86789F8A8FF0DF879" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF1FFE86789F8A8FF0DF879" italics="true" pageId="11" pageNumber="68">D. limon</emphasis>
</taxonomicName>
(MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C– D) 
<taxonomicName id="4C60BCF4FFF1FFE862B6F86AFEB9F81A" box="[188,357,1976,1997]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF1FFE862B6F86AFEB9F81A" box="[188,357,1976,1997]" italics="true" pageId="11" pageNumber="68">D. purpurescens</emphasis>
</taxonomicName>
, (MHCH 2636, MHCH 2635); E–F) 
<taxonomicName id="4C60BCF4FFF1FFE860F2F86AFC91F81A" box="[760,845,1976,1997]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="maia" status="sp. nov.">
<emphasis id="B9141B65FFF1FFE860F2F86AFC91F81A" box="[760,845,1976,1997]" italics="true" pageId="11" pageNumber="68">D. maia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227A61EFFF1FFE8615DF865FC76F81B" box="[855,938,1975,1996]" pageId="11" pageNumber="68" rank="species">
<emphasis id="B9141B65FFF1FFE8615DF865FC76F81B" bold="true" box="[855,938,1975,1996]" pageId="11" pageNumber="68">sp. nov.</emphasis>
</taxonomicNameLabel>
(SMF 97268, MHCH 2782); G–H) 
<taxonomicName id="4C60BCF4FFF1FFE8673AF86AFA40F81B" box="[1328,1436,1974,1997]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="68" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF1FFE8673AF86AFA40F81B" box="[1328,1436,1974,1997]" italics="true" pageId="11" pageNumber="68">D. ibanezi</emphasis>
</taxonomicName>
(MHCH2019, SMF 91475).
</paragraph>
</caption>
<caption id="DF1F97FFFFF6FFEF629DF8ADFD36F826" httpUri="https://zenodo.org/record/289967/files/figure.png" pageId="12" pageNumber="69" targetBox="[166,1436,193,1888]" targetPageId="12">
<paragraph id="8BDFC777FFF6FFEF629DF8ADFD36F826" blockId="12.[151,1436,1919,2033]" pageId="12" pageNumber="69">
<emphasis id="B9141B65FFF6FFEF629DF8ADFEC8F843" bold="true" box="[151,276,1919,1941]" pageId="12" pageNumber="69">FIGURE 5.</emphasis>
Flank pattern in species of 
<taxonomicName id="4C60BCF4FFF6FFEF603CF8ADFD4FF842" box="[566,659,1919,1941]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="genus">
<emphasis id="B9141B65FFF6FFEF603CF8ADFD4FF842" box="[566,659,1919,1941]" italics="true" pageId="12" pageNumber="69">Dactyloa</emphasis>
</taxonomicName>
related to 
<taxonomicName id="4C60BCF4FFF6FFEF610EF852FC77F842" box="[772,939,1920,1941]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF6FFEF610EF852FC77F842" box="[772,939,1920,1941]" italics="true" pageId="12" pageNumber="69">D. purpurescens</emphasis>
</taxonomicName>
. Arrows indicate capitad direction for all males (left column) and females (right column). A) 
<taxonomicName id="4C60BCF4FFF6FFEF606AF84DFD64F863" box="[608,696,1950,1972]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF6FFEF606AF84DFD64F863" box="[608,696,1950,1972]" italics="true" pageId="12" pageNumber="69">D. limon</emphasis>
</taxonomicName>
holotype MHUA 11760; B) D. 
<taxonomicName id="4C60BCF4FFF6FFEF61F3F84CFBEFF863" box="[1017,1075,1950,1972]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF6FFEF61F3F84CFBEFF863" box="[1017,1075,1950,1972]" italics="true" pageId="12" pageNumber="69">limon</emphasis>
</taxonomicName>
MHUA 11248; C) 
<taxonomicName id="4C60BCF4FFF6FFEF66FCF84DFA47F863" box="[1270,1435,1951,1972]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF6FFEF66FCF84DFA47F863" box="[1270,1435,1951,1972]" italics="true" pageId="12" pageNumber="69">D. purpurescens</emphasis>
</taxonomicName>
(MHCH 2636); D) 
<taxonomicName id="4C60BCF4FFF6FFEF6350F86FFE23F805" box="[346,511,1981,2002]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF6FFEF6350F86FFE23F805" box="[346,511,1981,2002]" italics="true" pageId="12" pageNumber="69">D. purpurescens</emphasis>
</taxonomicName>
(SMF 91475); E) 
<taxonomicName id="4C60BCF4FFF6FFEF60B3F86FFCD6F805" box="[697,778,1981,2002]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF6FFEF60B3F86FFCD6F805" box="[697,778,1981,2002]" italics="true" pageId="12" pageNumber="69">D. maia</emphasis>
</taxonomicName>
(SMF 97268); F) 
<taxonomicName id="4C60BCF4FFF6FFEF61C8F86FFBCFF805" box="[962,1043,1981,2002]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF6FFEF61C8F86FFBCFF805" box="[962,1043,1981,2002]" italics="true" pageId="12" pageNumber="69">D. maia</emphasis>
</taxonomicName>
(MHCH 2782); G) 
<taxonomicName id="4C60BCF4FFF6FFEF66D6F86FFA98F805" box="[1244,1348,1980,2002]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF6FFEF66D6F86FFA98F805" box="[1244,1348,1980,2002]" italics="true" pageId="12" pageNumber="69">D. ibanezi</emphasis>
</taxonomicName>
, MHCH 2184; H) 
<taxonomicName id="4C60BCF4FFF6FFEF62F2F80EFEBCF826" box="[248,352,2011,2033]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="69" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF6FFEF62F2F80EFEBCF826" box="[248,352,2011,2033]" italics="true" pageId="12" pageNumber="69">D. ibanezi</emphasis>
</taxonomicName>
, SMF 91475. Scale bars equal 10 mm.
</paragraph>
</caption>
<caption id="DF1F97FFFFF7FFEE629DFC1FFA53FBD5" httpUri="https://zenodo.org/record/289968/files/figure.png" pageId="13" pageNumber="70" targetBox="[217,1369,193,951]" targetPageId="13">
<paragraph id="8BDFC777FFF7FFEE629DFC1FFA53FBD5" blockId="13.[151,1436,973,1026]" pageId="13" pageNumber="70">
<emphasis id="B9141B65FFF7FFEE629DFC1FFECEFC34" bold="true" box="[151,274,973,995]" pageId="13" pageNumber="70">FIGURE 6.</emphasis>
Lateral head views of the species of 
<taxonomicName id="4C60BCF4FFF7FFEE6083FC1FFD3AFC34" box="[649,742,973,995]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis id="B9141B65FFF7FFEE6083FC1FFD3AFC34" box="[649,742,973,995]" italics="true" pageId="13" pageNumber="70">Dactyloa</emphasis>
</taxonomicName>
related to 
<emphasis id="B9141B65FFF7FFEE6158FC1DFC23FC33" box="[850,1023,975,996]" italics="true" pageId="13" pageNumber="70">
<taxonomicName id="4C60BCF4FFF7FFEE6158FC1DFC20FC33" box="[850,1020,975,996]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="purpurescens">D. purpurescens</taxonomicName>
.
</emphasis>
A) 
<taxonomicName id="4C60BCF4FFF7FFEE662FFC1DFBA2FC34" box="[1061,1150,973,996]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="limon">
<emphasis id="B9141B65FFF7FFEE662FFC1DFBA2FC34" box="[1061,1150,973,996]" italics="true" pageId="13" pageNumber="70">D. limon</emphasis>
</taxonomicName>
holotype MHUA 11760; B) 
<taxonomicName id="4C60BCF4FFF7FFEE629DFC3FFF23FBD5" box="[151,255,1004,1026]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B9141B65FFF7FFEE629DFC3FFF23FBD5" box="[151,255,1004,1026]" italics="true" pageId="13" pageNumber="70">D. ibanezi</emphasis>
</taxonomicName>
MHCH 2184; C) 
<taxonomicName id="4C60BCF4FFF7FFEE63BDFC3FFD81FBD5" box="[439,605,1005,1026]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF7FFEE63BDFC3FFE15FBD5" box="[439,457,1005,1026]" italics="true" pageId="13" pageNumber="70">D</emphasis>
. 
<emphasis id="B9141B65FFF7FFEE63DCFC3FFD81FBD5" box="[470,605,1005,1026]" italics="true" pageId="13" pageNumber="70">purpurescens</emphasis>
</taxonomicName>
SMF 97271; D) 
<taxonomicName id="4C60BCF4FFF7FFEE6100FC3FFC80FBD5" box="[778,860,1005,1026]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="maia" status="sp. nov.">
<emphasis id="B9141B65FFF7FFEE6100FC3FFC80FBD5" box="[778,860,1005,1026]" italics="true" pageId="13" pageNumber="70">D. maia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227A61EFFF7FFEE6168FC3EFC6FFBD6" box="[866,947,1004,1025]" pageId="13" pageNumber="70" rank="species">
<emphasis id="B9141B65FFF7FFEE6168FC3EFC6FFBD6" bold="true" box="[866,947,1004,1025]" pageId="13" pageNumber="70">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype SMF 97268. Scale bars equal 10 mm
</paragraph>
</caption>
<paragraph id="8BDFC777FFF7FFEE62CDFBFCFD8AF9CC" blockId="13.[151,1437,1070,1959]" pageId="13" pageNumber="70">
<emphasis id="B9141B65FFF7FFEE62CDFBFCFE56FB90" bold="true" box="[199,394,1070,1095]" pageId="13" pageNumber="70">Color variation.</emphasis>
Another male (SMF 97269, 8E–F), at the moment of encounter, was uniform green with bluish spots on flanks arranged in oblique rows; after collection it exhibited metachrosis, and the coloration recorded was as follows: the dorsal ground color of body, head and limbs Salmon Color (58), grading into Yellow Green (103) toward the flanks; tail with Russet (44) transverse bands; four well-defined Olive Green (123) oblique bands between axilla and groin, each including four to seven irregular Cyan Black (153) blotches; ventral surfaces and dewlap as in 
<typeStatus id="54DB79D5FFF7FFEE6357FB31FE1FFB2B" box="[349,451,1251,1276]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
. A female (MHCH 2782, 
<figureCitation id="135BDBF2FFF7FFEE60E3FB31FCEAFB2C" box="[745,822,1251,1276]" captionStart="FIGURE 4" captionStartId="11.[151,250,1913,1935]" captionTargetBox="[174,1428,193,1888]" captionTargetId="figure@11.[174,1459,193,1889]" captionTargetPageId="11" captionText="FIGURE 4. Dewlaps of the Dactyloa spp. related to D. purpurescens (left column males, right column females). A – B) D. limon (MHUA 11760; holotype; photo by Juan Pablo Hurtado), female not collected (from; photo by Alejandro Montoya); C – D) D. purpurescens, (MHCH 2636, MHCH 2635); E – F) D. maia sp. nov. (SMF 97268, MHCH 2782); G – H) D. ibanezi (MHCH 2019, SMF 91475)." httpUri="https://zenodo.org/record/289966/files/figure.png" pageId="13" pageNumber="70">Figs. 4</figureCitation>
F, 5F, 8A–B) lacked the small oval spots, and only the three Parrot Green (121) transverse bands were present with three small Sulphur Yellow (80) spots between each band; dewlap Medium Chrome Orange (75) with three well defined longitudinal series of Emerald Green (143) scales, free margin of dewlap dirty white, anterior insertion of dewlap suffused with Light Neutral Gray (297). Another female (SMF 97270; 
<figureCitation id="135BDBF2FFF7FFEE63F8FAA1FDE8FA5C" box="[498,564,1395,1420]" captionStart="FIGURE 8" captionStartId="15.[151,250,1903,1925]" captionTargetBox="[151,1436,197,1891]" captionTargetId="figure@15.[151,1436,197,1893]" captionTargetPageId="15" captionText="FIGURE 8. Color variation of Dactyloa maia sp. nov. in life. A – B) female MHCH 2782; C – D) female MHCH 2781; E – F) male SMF 97269; G) female SMF 97270." httpUri="https://zenodo.org/record/289970/files/figure.png" pageId="13" pageNumber="70">Fig. 8</figureCitation>
G) agrees in general color pattern with female MHCH 2782. A third female (MHCH 2781; 
<figureCitation id="135BDBF2FFF7FFEE634FFA44FE5BFA78" box="[325,391,1430,1455]" captionStart="FIGURE 8" captionStartId="15.[151,250,1903,1925]" captionTargetBox="[151,1436,197,1891]" captionTargetId="figure@15.[151,1436,197,1893]" captionTargetPageId="15" captionText="FIGURE 8. Color variation of Dactyloa maia sp. nov. in life. A – B) female MHCH 2782; C – D) female MHCH 2781; E – F) male SMF 97269; G) female SMF 97270." httpUri="https://zenodo.org/record/289970/files/figure.png" pageId="13" pageNumber="70">Fig. 8</figureCitation>
C–D) has the same dorsal ground color as the 
<typeStatus id="54DB79D5FFF7FFEE6195FA44FBD8FA78" box="[927,1028,1430,1455]" pageId="13" pageNumber="70" type="holotype">holotype</typeStatus>
, but with six well defined Greenish Olive (125) oblique bands in the flanks (without spots); dewlap Olive (126), with three well defined longitudinal series of Lime Green (116) scales, anterior and external border of dewlap dirty white, anterior insertion of dewlap suffused with Light Neutral Gray (297).
</paragraph>
</subSubSection>
<subSubSection id="C37A94FCFFF7FFEE62CDF9F4FDECF8C0" pageId="13" pageNumber="70" type="distribution">
<paragraph id="8BDFC777FFF7FFEE62CDF9F4FDECF8C0" blockId="13.[151,1437,1070,1959]" pageId="13" pageNumber="70">
<emphasis id="B9141B65FFF7FFEE62CDF9F4FDBBF9E8" bold="true" box="[199,615,1574,1599]" pageId="13" pageNumber="70">Distribution and Natural history.</emphasis>
<collectingCountry id="F37787E7FFF7FFEE607EF9F5FD4FF9E8" box="[628,659,1575,1599]" name="American Samoa" pageId="13" pageNumber="70">As</collectingCountry>
far as we know, 
<taxonomicName id="4C60BCF4FFF7FFEE617EF9F5FBFCF9E8" box="[884,1056,1575,1599]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF7FFEE617EF9F5FBFCF9E8" box="[884,1056,1575,1599]" italics="true" pageId="13" pageNumber="70">Dactyloa maia</emphasis>
</taxonomicName>
is endemic to eastern 
<collectingCountry id="F37787E7FFF7FFEE6736F9FAFA44F9E8" box="[1340,1432,1576,1599]" name="Panama" pageId="13" pageNumber="70">Panama</collectingCountry>
, inhabiting the foothills and ridges of the Majé, San Blas, Darién (as far south and east as to Río Tuquesa, thereafter apparently replaced by 
<taxonomicName id="4C60BCF4FFF7FFEE63BAF9A2FDB3F950" box="[432,623,1648,1671]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="purpurescens">
<emphasis id="B9141B65FFF7FFEE63BAF9A2FDB3F950" box="[432,623,1648,1671]" italics="true" pageId="13" pageNumber="70">D. purpurescens</emphasis>
</taxonomicName>
), and Piedras-Pacora (two specimens photographed by Angel Sosa- Bartuano, not plotted in 
<figureCitation id="135BDBF2FFF7FFEE63A1F941FE36F97C" box="[427,490,1683,1708]" captionStart="FIGURE 1" captionStartId="3.[151,250,1642,1664]" captionTargetBox="[165,1423,211,1593]" captionTargetId="figure@3.[150,1437,193,1620]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Dactyloa species formerly referred to as D. chocorum in eastern Panama, based on specimens collected for this study, provided by A. Sosa, literature and online databases (http: // www. vertnet. org /)." httpUri="https://zenodo.org/record/289963/files/figure.png" pageId="13" pageNumber="70">Fig. 1</figureCitation>
) mountain ranges, where it occurs in the eastern Panamanian montane forest (Fund 2014) and the Isthmian-Atlantic moist forests (in the Piedras-Pacora and San Blas mountain range, 
<bibRefCitation id="EFF1BA86FFF7FFEE66F9F96AFF0FF924" author="Hogan" pageId="13" pageNumber="70" refString="Hogan, C. &amp; Fund, W. (2014) Isthmian-Atlantic moist forests. Available from: http: // www. eoearth. org / view / article / 153927 (accessed 23 September 2015)" type="journal article" year="2014">Hogan &amp; Fund 2014</bibRefCitation>
), at 
<quantity id="4C986A92FFF7FFEE6309F909FE5FF924" box="[259,387,1755,1780]" metricMagnitude="2" metricUnit="m" metricValue="5.87" metricValueMax="8.52" metricValueMin="3.22" pageId="13" pageNumber="70" unit="m" value="587.0" valueMax="852.0" valueMin="322.0">322–852 m</quantity>
asl. All specimens of 
<taxonomicName id="4C60BCF4FFF7FFEE608AF909FCFBF924" box="[640,807,1755,1779]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF7FFEE608AF909FCFBF924" box="[640,807,1755,1779]" italics="true" pageId="13" pageNumber="70">Dactyloa maia</emphasis>
</taxonomicName>
were found during night searches sleeping on branches or leaves 
<quantity id="4C986A92FFF7FFEE6308F92DFE83F8C0" box="[258,351,1791,1815]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="70" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2 to 3 m</quantity>
above the ground.
</paragraph>
</subSubSection>
<subSubSection id="C37A94FCFFF7FFEE62CDF8F1FDF7F870" pageId="13" pageNumber="70" type="etymology">
<paragraph id="8BDFC777FFF7FFEE62CDF8F1FDF7F870" blockId="13.[151,1437,1070,1959]" pageId="13" pageNumber="70">
<emphasis id="B9141B65FFF7FFEE62CDF8F1FE92F8EB" bold="true" box="[199,334,1827,1852]" pageId="13" pageNumber="70">Etymology.</emphasis>
Abel Batista dedicates this beautiful new species to his recently born daughter, 
<taxonomicName id="4C60BCF4FFF7FFEE66E8F8F1FAC3F8EB" box="[1250,1311,1827,1852]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="82" phylum="Chordata" rank="species" species="maia">Maia</taxonomicName>
. The name also comes from Greek mythology, where it is applied to the eldest of the Pleiades, sometimes called mountain nymphs, and are believed to live on the trees in mountains and groves as the guardians for that habitat, a role 
<taxonomicName id="4C60BCF4FFF7FFEE629DF85DFEE1F870" box="[151,317,1935,1959]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="70" phylum="Chordata" rank="species" species="maia">
<emphasis id="B9141B65FFF7FFEE629DF85DFEE1F870" box="[151,317,1935,1959]" italics="true" pageId="13" pageNumber="70">Dactyloa maia</emphasis>
</taxonomicName>
could also represent.
</paragraph>
</subSubSection>
</treatment>
</document>