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<document id="EDBBE5B1F3DF6F0D779BA4594FEA8C4B" ID-CLB-Dataset="5604" ID-DOI="10.11646/zootaxa.3795.5.10" ID-GBIF-Dataset="ffb607f0-c1ab-4398-acf2-cb84fbbf9c37" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915107" ID-ZooBank="0315DE1C-6668-4453-8BA2-876CF7E25D30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623198254184" checkinUser="felipe" docAuthor="Génier, François &amp; Howden, Henry F." docDate="2014" docId="993DB83CFF9E4A5301EE7F44E658BC49" docLanguage="en" docName="zt03795p599.pdf" docOrigin="Zootaxa 3795 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus fragosus Genier &amp; Howden 2014, new species" docType="treatment" docUuid="098EC3C4-0A75-4D71-8BE0-8C4D56B1A454" docUuidSource="ZooBank" docVersion="8" lastPageNumber="599" masterDocId="6504C044FF9E4A5101797A03E42DB830" masterDocTitle="Onthophagus fragosus n. sp. a second endemic species of Onthophagus Latreille from Cuba (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="599" masterPageNumber="597" pageNumber="597" updateTime="1699006217642" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0326ED45775E25FBD40A8AF3193D469E">Onthophagus fragosus n. sp. a second endemic species of Onthophagus Latreille from Cuba (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:namePart id="A5176691CF7E60141D89F8131A083961">Génier, François</mods:namePart>
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<mods:namePart id="B82718B0C0C5A7A73155DC996E659201">Howden, Henry F.</mods:namePart>
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<mods:date id="C255FBB881B1EE8C7B51F01D02BDE34F">2014</mods:date>
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<mods:number id="0C8E9D021AA04360DF84DAAA5F0CA417">2014-05-15</mods:number>
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<mods:number id="70B8D00CAF682931FAD4B5A290B15ED4">3795</mods:number>
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<treatment id="993DB83CFF9E4A5301EE7F44E658BC49" ID-DOI="http://doi.org/10.5281/zenodo.5083020" ID-GBIF-Taxon="183403892" ID-Zenodo-Dep="5083020" LSID="urn:lsid:zoobank.org:act:098EC3C4-0A75-4D71-8BE0-8C4D56B1A454" httpUri="http://treatment.plazi.org/id/993DB83CFF9E4A5301EE7F44E658BC49" lastPageId="2" lastPageNumber="599" pageId="0" pageNumber="597">
<subSubSection id="598E5AA1FF9E4A5101EE7F44E6D4BD6E" box="[151,761,1351,1375]" pageId="0" pageNumber="597" type="nomenclature">
<paragraph id="112B092AFF9E4A5101EE7F44E6D4BD6E" blockId="0.[151,761,1351,1409]" box="[151,761,1351,1375]" pageId="0" pageNumber="597">
<heading id="4A63BE46FF9E4A5101EE7F44E6D4BD6E" bold="true" box="[151,761,1351,1375]" fontSize="10" level="2" pageId="0" pageNumber="597" reason="2">
<emphasis id="23E0D538FF9E4A5101EE7F44E6D4BD6E" bold="true" box="[151,761,1351,1375]" pageId="0" pageNumber="597">
<taxonomicName id="D69472A9FF9E4A5101EE7F44E646BD6E" ID-CoL="7XMWW" authority="Genier &amp; Howden" authorityName="Genier &amp; Howden" authorityYear="2014" box="[151,619,1351,1375]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="fragosus" status="sp. nov.">
<emphasis id="23E0D538FF9E4A5101EE7F44E5BCBD6F" bold="true" box="[151,401,1351,1375]" italics="true" pageId="0" pageNumber="597">Onthophagus fragosus</emphasis>
Génier &amp; Howden
</taxonomicName>
,
<taxonomicNameLabel id="38D36843FF9E4A51030E7F44E6D4BD6E" box="[631,761,1351,1374]" pageId="0" pageNumber="597" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9E4A5101EE7F69E4D7BDB1" box="[151,250,1386,1409]" pageId="0" pageNumber="597" type="description">
<paragraph id="112B092AFF9E4A5101EE7F69E4D7BDB1" blockId="0.[151,761,1351,1409]" box="[151,250,1386,1409]" pageId="0" pageNumber="597">
<figureCitation id="89AF15AFFF9E4A5101EE7F69E4D7BDB1" box="[151,250,1386,1409]" captionStart="FIGURES 1–4" captionStartId="1.[151,269,1899,1922]" captionTargetBox="[237,1325,207,1868]" captionTargetId="figure-82@1.[225,1342,194,1887]" captionTargetPageId="1" captionText="FIGURES 1–4. Onthophagus fragosus n. sp. 1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite)." figureDoi="http://doi.org/10.5281/zenodo.4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="0" pageNumber="597">Figs. 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9E4A5101EE7FACE0D8BE44" pageId="0" pageNumber="597" type="materials_examined">
<paragraph id="112B092AFF9E4A5101EE7FACE5ACBE3C" blockId="0.[151,1436,1455,2033]" pageId="0" pageNumber="597">
<materialsCitation id="A1FC0377FF9E4A5101EE7FACE58EBDD9" ID-GBIF-Occurrence="3319795301" collectingDate="2013-05-20" collectionCode="CMNC, R" country="Cuba" elevation="651" latitude="21.93123" location="Rio Cabagan" longLatPrecision="1" longitude="-80.08461" pageId="0" pageNumber="597" specimenCount="1" stateProvince="Cienfuegos" typeStatus="holotype">
<emphasis id="23E0D538FF9E4A5101EE7FACE4D2BDF6" bold="true" box="[151,255,1455,1478]" pageId="0" pageNumber="597">
<typeStatus id="CE2FB788FF9E4A5101EE7FACE4D2BDF6" box="[151,255,1455,1478]" pageId="0" pageNumber="597" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="778591EFFF9E4A51003E7FACE5B1BDF6" box="[327,412,1455,1478]" pageId="0" pageNumber="597">CMNC</collectionCode>
):
<collectingCountry id="698349BAFF9E4A5100D77FACE5D1BDF6" box="[430,508,1455,1478]" name="Cuba" pageId="0" pageNumber="597">CUBA</collectingCountry>
:
<collectingRegion id="D350C7C8FF9E4A51037F7FACE6AFBDF6" box="[518,642,1455,1478]" country="Cuba" name="Cienfuegos" pageId="0" pageNumber="597">Cienfuegos</collectingRegion>
,
<location id="144B5FF1FF9E4A5103F47FACE737BDF6" LSID="urn:lsid:plazi:treatment:993DB83CFF9E4A5301EE7F44E658BC49:144B5FF1FF9E4A5103F47FACE737BDF6" box="[653,794,1455,1478]" country="Cuba" latitude="21.93123" longLatPrecision="1" longitude="-80.08461" name="Rio Cabagan" pageId="0" pageNumber="597" stateProvince="Cienfuegos">Rio Cabagan</location>
,
<geoCoordinate id="74A06FEDFF9E4A51025D7FACE7A5BDF6" box="[804,904,1455,1478]" degrees="21.93123" direction="north" orientation="latitude" pageId="0" pageNumber="597" precision="1" value="21.93123">21.93123</geoCoordinate>
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,
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<elevation id="9AB9EE19FF9E4A5105707FACE064BDF6" box="[1033,1097,1455,1478]" metricMagnitude="2" metricUnit="m" metricValue="6.51" pageId="0" pageNumber="597" unit="m" value="651.0">651m</elevation>
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,
<date id="652A2FEAFF9E4A51052A7FACE092BDF6" box="[1107,1215,1455,1478]" pageId="0" pageNumber="597" value="2013-05-20">
<collectingDate id="756ED602FF9E4A51052A7FACE092BDF6" box="[1107,1215,1455,1478]" pageId="0" pageNumber="597" value="2013-05-20">20.v.2013</collectingDate>
</date>
,
<collectionCode id="778591EFFF9E4A5105B07FACE0F3BDF6" box="[1225,1246,1455,1478]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="597">R</collectionCode>
. Anderson, 2013- 026, gallery forest litter
</materialsCitation>
/
<typeStatus id="CE2FB788FF9E4A5100CF7FD1E66BBDD9" box="[438,582,1490,1513]" pageId="0" pageNumber="597" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="D69472A9FF9E4A51032F7FD1E760BDD9" authority="Génier &amp; Howden, 2014" authorityName="Génier &amp; Howden" authorityYear="2014" box="[598,845,1490,1513]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="fragosus" status="sp. nov.">
<emphasis id="23E0D538FF9E4A51032F7FD1E760BDD9" box="[598,845,1490,1513]" italics="true" pageId="0" pageNumber="597">Onthophagus fragosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38D36843FF9E4A5102237FD1E7A3BDD9" box="[858,910,1490,1513]" pageId="0" pageNumber="597" rank="species">n.sp.</taxonomicNameLabel>
, des. Génier &amp; Howden, 2014. Aedeagus and internal sac extracted.
</paragraph>
<paragraph id="112B092AFF9E4A5101BE7C14E0D8BE44" blockId="0.[151,1436,1455,2033]" pageId="0" pageNumber="597">
<materialsCitation id="A1FC0377FF9E4A5101BE7C14E65DBE1E" ID-GBIF-Occurrence="3319795304" box="[199,624,1559,1582]" collectionCode="CMNC" country="Cuba" pageId="0" pageNumber="597" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<emphasis id="23E0D538FF9E4A5101BE7C14E56DBE1E" bold="true" box="[199,320,1559,1582]" inLineHeading="true" pageId="0" pageNumber="597" reason="1">
<typeStatus id="CE2FB788FF9E4A5101BE7C14E516BE1E" box="[199,315,1559,1582]" pageId="0" pageNumber="597" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="0792C2A3FF9E4A51003E7C14E5BFBE1E" box="[327,402,1559,1582]" pageId="0" pageNumber="597" type="male">1 male</specimenCount>
,
<specimenCount id="0792C2A3FF9E4A5100E47C14E628BE1E" box="[413,517,1559,1582]" pageId="0" pageNumber="597" type="female">2 females</specimenCount>
(
<collectionCode id="778591EFFF9E4A51036D7C14E645BE1E" box="[532,616,1559,1582]" pageId="0" pageNumber="597">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A1FC0377FF9E4A5103037C14E0A1BE61" ID-GBIF-Occurrence="3319795303" collectingDate="2013-05-19" collectingDateMax="2013-05-20" collectingDateMin="2013-05-19" collectionCode="R" collectorName="P. N. Pico San Juan &amp; Anderson" country="Cuba" elevation="934" latitude="21.93123" location="Rio Cabagan" longLatPrecision="1" longitude="-80.08461" pageId="0" pageNumber="597" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Cienfuegos" typeStatus="paratype">
<emphasis id="23E0D538FF9E4A5103037C14E6E8BE1E" bold="true" box="[634,709,1559,1582]" pageId="0" pageNumber="597">
<collectingCountry id="698349BAFF9E4A5103037C14E6E8BE1E" box="[634,709,1559,1582]" name="Cuba" pageId="0" pageNumber="597">CUBA</collectingCountry>
</emphasis>
:
<collectingRegion id="D350C7C8FF9E4A5103AD7C14E77CBE1E" box="[724,849,1559,1582]" country="Cuba" name="Cienfuegos" pageId="0" pageNumber="597">Cienfuegos</collectingRegion>
:
<collectorName id="BC616CFCFF9E4A5102257C14E004BE1E" box="[860,1065,1559,1582]" pageId="0" pageNumber="597">P.N. Pico San Juan</collectorName>
, road,
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,
<quantity id="D66CA4CFFF9E4A5104217C14E1B5BE1E" box="[1368,1432,1559,1582]" metricMagnitude="2" metricUnit="m" metricValue="9.34" pageId="0" pageNumber="597" unit="m" value="934.0">
<elevation id="9AB9EE19FF9E4A5104217C14E1B5BE1E" box="[1368,1432,1559,1582]" metricMagnitude="2" metricUnit="m" metricValue="9.34" pageId="0" pageNumber="597" unit="m" value="934.0">934m</elevation>
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,
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<collectingDate id="756ED602FF9E4A5101EE7C39E52EBE61" box="[151,259,1594,1617]" pageId="0" pageNumber="597" value="2013-05-19">19.v.2013</collectingDate>
</date>
.
<collectionCode id="778591EFFF9E4A5100757C39E50CBE61" box="[268,289,1594,1617]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="597">R</collectionCode>
.
<collectorName id="BC616CFCFF9E4A5100527C39E5B8BE61" box="[299,405,1594,1617]" pageId="0" pageNumber="597">Anderson</collectorName>
, 2013-21, hardwood forest litter (
<specimenCount id="0792C2A3FF9E4A5103857C39E77ABE61" box="[764,855,1594,1617]" pageId="0" pageNumber="597" type="female">1 female</specimenCount>
<typeStatus id="CE2FB788FF9E4A5102257C39E790BE61" box="[860,957,1594,1617]" pageId="0" pageNumber="597" type="paratype">paratype</typeStatus>
); same data except:
</materialsCitation>
<materialsCitation id="A1FC0377FF9E4A5105E87C39E0DCBE44" ID-GBIF-Occurrence="3319795302" collectingDate="2013-05-19" collectingDateMax="2013-05-20" collectingDateMin="2013-05-19" collectionCode="R" collectorName="P. N. Pico San Juan &amp; Anderson" country="Cuba" elevation="934" latitude="21.93123" location="Rio Cabagan" longLatPrecision="1" longitude="-80.08461" pageId="0" pageNumber="597" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Cienfuegos" typeStatus="paratype">
<location id="144B5FF1FF9E4A5105E87C39E133BE61" LSID="urn:lsid:plazi:treatment:993DB83CFF9E4A5301EE7F44E658BC49:144B5FF1FF9E4A5105E87C39E133BE61" box="[1169,1310,1594,1617]" country="Cuba" latitude="21.93123" longLatPrecision="1" longitude="-80.08461" name="Rio Cabagan" pageId="0" pageNumber="597" stateProvince="Cienfuegos">Rio Cabagan</location>
,
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<geoCoordinate id="74A06FEDFF9E4A5104EB7C39E4D3BE44" degrees="80.08461" direction="west" orientation="longitude" pageId="0" pageNumber="597" precision="1" value="-80.08461">- 80.08461</geoCoordinate>
,
<quantity id="D66CA4CFFF9E4A5100717C5EE565BE44" box="[264,328,1629,1652]" metricMagnitude="2" metricUnit="m" metricValue="6.51" pageId="0" pageNumber="597" unit="m" value="651.0">
<elevation id="9AB9EE19FF9E4A5100717C5EE565BE44" box="[264,328,1629,1652]" metricMagnitude="2" metricUnit="m" metricValue="6.51" pageId="0" pageNumber="597" unit="m" value="651.0">651m</elevation>
</quantity>
,
<date id="652A2FEAFF9E4A51002B7C5EE593BE44" box="[338,446,1629,1652]" pageId="0" pageNumber="597" value="2013-05-20">
<collectingDate id="756ED602FF9E4A51002B7C5EE593BE44" box="[338,446,1629,1652]" pageId="0" pageNumber="597" value="2013-05-20">20.v.2013</collectingDate>
</date>
,
<collectionCode id="778591EFFF9E4A5100B17C5EE5F0BE44" box="[456,477,1629,1652]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="597">R</collectionCode>
. Anderson, 2013-026, gallery forest litter. (
<specimenCount id="0792C2A3FF9E4A5102D57C5EE7DEBE44" box="[940,1011,1629,1652]" pageId="0" pageNumber="597" type="male">1 male</specimenCount>
and
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<typeStatus id="CE2FB788FF9E4A5105F07C5EE0C7BE44" box="[1161,1258,1629,1652]" pageId="0" pageNumber="597" type="paratype">paratype</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9E4A5101BE7C7CE05BBEEC" pageId="0" pageNumber="597" type="diagnosis">
<paragraph id="112B092AFF9E4A5101BE7C7CE05BBEEC" blockId="0.[151,1436,1455,2033]" pageId="0" pageNumber="597">
<emphasis id="23E0D538FF9E4A5101BE7C7CE516BEA6" box="[199,315,1663,1686]" inLineHeading="true" pageId="0" pageNumber="597" reason="1">
<emphasis id="23E0D538FF9E4A5101BE7C7CE519BEA6" bold="true" box="[199,308,1663,1686]" pageId="0" pageNumber="597">Diagnosis</emphasis>
.
</emphasis>
Readily separated from all other New World
<taxonomicName id="D69472A9FF9E4A5102387C7CE004BEA6" box="[833,1065,1663,1686]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="23E0D538FF9E4A5102387C7CE7FDBEA6" box="[833,976,1663,1686]" italics="true" pageId="0" pageNumber="597">Onthophagus</emphasis>
species
</taxonomicName>
by the following combination of characters: pronotal and pygidial surface covered with large and coarse umbilicate punctures; elytral striae each with a single row of coarse setiferous punctures, each puncture preceded by a small coarse tubercle.
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9E4A5301BE7CE4E042BAA4" lastPageId="2" lastPageNumber="599" pageId="0" pageNumber="597" type="description">
<paragraph id="112B092AFF9E4A5101BE7CE4E1BBBFC1" blockId="0.[151,1436,1455,2033]" pageId="0" pageNumber="597">
<emphasis id="23E0D538FF9E4A5101BE7CE4E57FBECE" box="[199,338,1767,1790]" inLineHeading="true" pageId="0" pageNumber="597" reason="1">
<emphasis id="23E0D538FF9E4A5101BE7CE4E566BECE" bold="true" box="[199,331,1767,1790]" pageId="0" pageNumber="597">Description</emphasis>
.
</emphasis>
<typeStatus id="CE2FB788FF9E4A51002E7CE4E597BECE" box="[343,442,1767,1790]" pageId="0" pageNumber="597" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="89AF15AFFF9E4A51037B7CE4E66CBECE" box="[514,577,1767,1790]" captionStart="FIGURES 1–4" captionStartId="1.[151,269,1899,1922]" captionTargetBox="[237,1325,207,1868]" captionTargetId="figure-82@1.[225,1342,194,1887]" captionTargetPageId="1" captionText="FIGURES 1–4. Onthophagus fragosus n. sp. 1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite)." figureDoi="http://doi.org/10.5281/zenodo.4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="0" pageNumber="597">Fig. 1</figureCitation>
).
<emphasis id="23E0D538FF9E4A51032E7CE4E6A8BECE" bold="true" box="[599,645,1767,1790]" pageId="0" pageNumber="597">Size</emphasis>
. Length
<quantity id="D66CA4CFFF9E4A51039A7CE4E71BBECE" box="[739,822,1767,1790]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="0" pageNumber="597" unit="mm" value="5.5">5.5 mm</quantity>
, maximum width 3.0 mm.
<emphasis id="23E0D538FF9E4A51052D7CE4E0B7BECE" bold="true" box="[1108,1178,1767,1790]" pageId="0" pageNumber="597">Color.</emphasis>
Black, with legs, elytral apex, lateral portion of visible abdominal sternites and pygidium brown; surface finely microsculptured, appearing more or less sericeous.
<emphasis id="23E0D538FF9E4A5100197D2EE5B0BF74" bold="true" box="[352,413,1837,1860]" pageId="0" pageNumber="597">Head</emphasis>
. Clypeal anterior edge bidentate, clypeal surface coarsely transversally rugose; clypeogenal junction slightly but distinctly emarginate; vertex with a short transverse carina occupying one-third width between eyes and with small tubercle adjacent to eye in line with carina; dorsal portion of eyes large, interocular ratio 2.7; antennal club yellowish, segments 1-6 slightly darker.
<emphasis id="23E0D538FF9E4A5103F47D96E72CBF9C" bold="true" box="[653,769,1941,1964]" pageId="0" pageNumber="597">Pronotum</emphasis>
. Simply convex, lacking tubercle or carina; surface covered with large and coarse umbilicate punctures varying in size, largest punctures three time as large as smallest, each puncture with moderately long erect yellowish seta; surface between punctures more coarsely microsculptured than elytra.
</paragraph>
<caption id="45EB59A2FF9F4A5001EE7D68E7A8BFDA" ID-DOI="http://doi.org/10.5281/zenodo.4915109" ID-Zenodo-Dep="4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="1" pageNumber="598" startId="1.[151,269,1899,1922]" targetBox="[237,1325,207,1868]" targetPageId="1">
<paragraph id="112B092AFF9F4A5001EE7D68E7A8BFDA" blockId="1.[151,1435,1899,2026]" pageId="1" pageNumber="598">
<emphasis id="23E0D538FF9F4A5001EE7D68E568BFB2" bold="true" box="[151,325,1899,1922]" pageId="1" pageNumber="598">FIGURES 1–4.</emphasis>
<taxonomicName id="D69472A9FF9F4A5000367D68E668BFB2" authority="Génier &amp; Howden, 2014" authorityName="Génier &amp; Howden" authorityYear="2014" box="[335,581,1899,1922]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="fragosus" status="sp. nov.">
<emphasis id="23E0D538FF9F4A5000367D68E668BFB2" box="[335,581,1899,1922]" italics="true" pageId="1" pageNumber="598">Onthophagus fragosus</emphasis>
</taxonomicName>
<emphasis id="23E0D538FF9F4A5003377D68E6A0BFB2" bold="true" box="[590,653,1899,1922]" pageId="1" pageNumber="598">
<taxonomicNameLabel id="38D36843FF9F4A5003377D68E6A0BFB2" box="[590,653,1899,1922]" pageId="1" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite).
</paragraph>
</caption>
<paragraph id="112B092AFF9C4A5301EF7A9BE759BA39" blockId="2.[150,1436,152,1145]" pageId="2" pageNumber="599">
<emphasis id="23E0D538FF9C4A5301EF7A9BE4CBB89F" box="[150,230,152,175]" inLineHeading="true" pageId="2" pageNumber="599" reason="1">
<emphasis id="23E0D538FF9C4A5301EF7A9BE4F2B89F" bold="true" box="[150,223,152,175]" pageId="2" pageNumber="599">Elytra</emphasis>
.
</emphasis>
Interstriae uneven, interstriae 1-7 with single more or less regular row of coarse setiferous punctures, each puncture preceded by small coarse tubercle, interstriae 8 with 2 more or less regular rows of coarse setiferous punctures on basal half; striae moderately wide, with row of well-defined punctures, lateral edge of striae bordered by fine sharp carina interrupted at each puncture, punctures deeper on apical declivity.
<emphasis id="23E0D538FF9C4A5302AD7B03E024B927" bold="true" box="[980,1033,256,279]" pageId="2" pageNumber="599">Legs</emphasis>
. Protibia unmodified, rather robust, lacking apical setal brush, apical tooth obliquely oriented, less developed than penultimate; median and posterior legs unmodified.
<emphasis id="23E0D538FF9C4A53005B7B46E5F4B96C" bold="true" box="[290,473,325,348]" pageId="2" pageNumber="599">Thoracic sterna</emphasis>
. Median lobe of metasternum with coarse setiferous punctures along mesocoxal edges; lateral lobes surface coarsely microsculptured, with coarse setiferous umbilicate punctures separated by approximately one diameter.
<emphasis id="23E0D538FF9C4A53004F7B89E60CB991" bold="true" box="[310,545,394,417]" pageId="2" pageNumber="599">Abdominal sternites</emphasis>
. Segments 4-8 each with single row of large and very deep punctures basally; pygidium wider than long (ratio 1.4), surface covered with moderate to large setiferous umbilicate punctures separated by less than one diameter.
<emphasis id="23E0D538FF9C4A5300CE7BD3E609B9D7" bold="true" box="[439,548,464,487]" pageId="2" pageNumber="599">Aedeagus</emphasis>
. Parameres (
<figureCitation id="89AF15AFFF9C4A5303CB7BD3E6DFB9D7" box="[690,754,464,487]" captionStart="FIGURES 1–4" captionStartId="1.[151,269,1899,1922]" captionTargetBox="[237,1325,207,1868]" captionTargetId="figure-82@1.[225,1342,194,1887]" captionTargetPageId="1" captionText="FIGURES 1–4. Onthophagus fragosus n. sp. 1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite)." figureDoi="http://doi.org/10.5281/zenodo.4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="2" pageNumber="599">Fig. 3</figureCitation>
) tapering to apex in lateral view, slightly divergent and simply rounded apically in frontal view; sclerites of internal sac as in
<figureCitation id="89AF15AFFF9C4A5302577BF1E742BA39" box="[814,879,498,521]" captionStart="FIGURES 1–4" captionStartId="1.[151,269,1899,1922]" captionTargetBox="[237,1325,207,1868]" captionTargetId="figure-82@1.[225,1342,194,1887]" captionTargetPageId="1" captionText="FIGURES 1–4. Onthophagus fragosus n. sp. 1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite)." figureDoi="http://doi.org/10.5281/zenodo.4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="2" pageNumber="599">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="112B092AFF9C4A5301BE7816E042BAA4" blockId="2.[150,1436,152,1145]" pageId="2" pageNumber="599">
<emphasis id="23E0D538FF9C4A5301BE7816E514BA1C" box="[199,313,533,556]" inLineHeading="true" pageId="2" pageNumber="599" reason="1">
<emphasis id="23E0D538FF9C4A5301BE7816E51FBA1C" bold="true" box="[199,306,533,556]" pageId="2" pageNumber="599">Variation</emphasis>
.
</emphasis>
Size. 5.0-7.0 mm. Females (
<figureCitation id="89AF15AFFF9C4A5303147816E683BA1C" box="[621,686,533,556]" captionStart="FIGURES 1–4" captionStartId="1.[151,269,1899,1922]" captionTargetBox="[237,1325,207,1868]" captionTargetId="figure-82@1.[225,1342,194,1887]" captionTargetPageId="1" captionText="FIGURES 1–4. Onthophagus fragosus n. sp. 1, habitus dorsal, holotype male; 2, habitus dorsal, allotype female; 3, aedeagus, lateral view; 4, sclerites of the internal sac (A: axial sclerite; AIS: additional inferior sclerite; FLP: frontolateral peripheral sclerite; IL: inferior left lobe of lamella copulatrix; LC: lamella copulatrix; MP: medial peripheral sclerite; SA: subaxial sclerite; SRP: Superior right peripheral sclerite)." figureDoi="http://doi.org/10.5281/zenodo.4915109" httpUri="https://zenodo.org/record/4915109/files/figure.png" pageId="2" pageNumber="599">Fig. 2</figureCitation>
) only slightly differ from males, in having more robust anterior tibiae and the carina of the vertex is slightly wider and more produced laterally, in addition to the ocular tubercle being slightly more developed. However, as usual they can easily be separated by the shape of abdominal segment 8, which is not reduced medially. The male
<typeStatus id="CE2FB788FF9C4A5300BC787EE60CBAA4" box="[453,545,637,660]" pageId="2" pageNumber="599" type="paratype">paratype</typeStatus>
differs in having the elytral apex much lighter in color.
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9C4A5301BE78A3E5D1BAE9" pageId="2" pageNumber="599" type="etymology">
<paragraph id="112B092AFF9C4A5301BE78A3E5D1BAE9" blockId="2.[150,1436,152,1145]" pageId="2" pageNumber="599">
<emphasis id="23E0D538FF9C4A5301BE78A3E56ABA87" box="[199,327,672,695]" inLineHeading="true" pageId="2" pageNumber="599" reason="1">
<emphasis id="23E0D538FF9C4A5301BE78A3E56DBA87" bold="true" box="[199,320,672,695]" pageId="2" pageNumber="599">Etymology</emphasis>
.
</emphasis>
<emphasis id="23E0D538FF9C4A53002A78A3E595BA87" box="[339,440,672,695]" italics="true" pageId="2" pageNumber="599">Fragosus</emphasis>
(broken, rough, uneven) a Latin adjective in apposition pertaining to the aspect of the tegument of this peculiar species.
</paragraph>
</subSubSection>
<subSubSection id="598E5AA1FF9C4A5301BE78E6E658BC49" pageId="2" pageNumber="599" type="discussion">
<paragraph id="112B092AFF9C4A5301BE78E6E658BC49" blockId="2.[150,1436,152,1145]" pageId="2" pageNumber="599">
<emphasis id="23E0D538FF9C4A5301BE78E6E518BACC" box="[199,309,741,764]" inLineHeading="true" pageId="2" pageNumber="599" reason="1">
<emphasis id="23E0D538FF9C4A5301BE78E6E502BACC" bold="true" box="[199,303,741,764]" pageId="2" pageNumber="599">Remarks</emphasis>
.
</emphasis>
Based on their very small size compared to the females, the two known male specimens are most likely underdeveloped individuals. The large dorsal ocular surface suggests that the species is nocturnal. Because all specimens were collected by sifting leaf litter there is no indication of food preference. The only other native
<taxonomicName id="D69472A9FF9C4A5305DF7929E11BBB71" authorityName="Latreille" authorityYear="1802" box="[1190,1334,810,833]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="genus">
<emphasis id="23E0D538FF9C4A5305DF7929E11BBB71" box="[1190,1334,810,833]" italics="true" pageId="2" pageNumber="599">Onthophagus</emphasis>
</taxonomicName>
on
<collectingCountry id="698349BAFF9C4A5304247929E1B5BB71" box="[1373,1432,810,833]" name="Cuba" pageId="2" pageNumber="599">Cuba</collectingCountry>
,
<taxonomicName id="D69472A9FF9C4A5301EE794EE589BB54" authority="Castelnau" authorityName="Castelnau" authorityYear="1840" box="[151,420,845,868]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="23E0D538FF9C4A5301EE794EE51DBB54" box="[151,304,845,868]" italics="true" pageId="2" pageNumber="599">O. marginatus</emphasis>
Castelnau
</taxonomicName>
, is widely distributed on the island. The new species has been collected from an isolated patch of Cuban moist forest at an elevation of
<quantity id="D66CA4CFFF9C4A53031B7973E6F3BBB7" box="[610,734,880,903]" metricMagnitude="2" metricUnit="m" metricValue="7.925" metricValueMax="9.35" metricValueMin="6.5" pageId="2" pageNumber="599" unit="m" value="792.5" valueMax="935.0" valueMin="650.0">
650-
<specimenCount id="0792C2A3FF9C4A5303EA7973E6F3BBB7" box="[659,734,880,903]" pageId="2" pageNumber="599" type="generic">935 m</specimenCount>
</quantity>
. This ecoregion (ecocode NT0120, see http://worldwildlife.org/ science/wildfinder/) is present in several areas of
<collectingCountry id="698349BAFF9C4A5303CE7991E6DEBB99" box="[695,755,914,937]" name="Cuba" pageId="2" pageNumber="599">Cuba</collectingCountry>
. The possibility that the new species could be found in other disjointed patches must be investigated. The presence of similar distinct umbilicate pygidial punctures combined with the presence of coarse granules on elytral intervals, a unique derived character for the New World
<taxonomicName id="D69472A9FF9C4A5305D679DBE113BBDF" authorityName="Latreille" authorityYear="1802" box="[1199,1342,984,1007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="genus">
<emphasis id="23E0D538FF9C4A5305D679DBE113BBDF" box="[1199,1342,984,1007]" italics="true" pageId="2" pageNumber="599">Onthophagus</emphasis>
</taxonomicName>
, suggest that
<taxonomicName id="D69472A9FF9C4A5301B379F9E567BC21" authorityName="Génier &amp; Howden" authorityYear="2014" box="[202,330,1018,1041]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="fragosus">
<emphasis id="23E0D538FF9C4A5301B379F9E567BC21" box="[202,330,1018,1041]" italics="true" pageId="2" pageNumber="599">O. fragosus</emphasis>
</taxonomicName>
is most closely related to the other Cuban endemic
<taxonomicName id="D69472A9FF9C4A5302E479F9E017BC21" authorityName="Castelnau" authorityYear="1840" box="[925,1082,1018,1041]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="23E0D538FF9C4A5302E479F9E017BC21" box="[925,1082,1018,1041]" italics="true" pageId="2" pageNumber="599">O. marginatus</emphasis>
</taxonomicName>
; however, the
<specimenCount id="0792C2A3FF9C4A53059C79F9E17BBC21" box="[1253,1366,1018,1041]" pageId="2" pageNumber="599" type="male">two males</specimenCount>
of
<taxonomicName id="D69472A9FF9C4A5304FB79F9E4DEBC04" authorityName="Génier &amp; Howden" authorityYear="2014" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="fragosus">
<emphasis id="23E0D538FF9C4A5304FB79F9E4DEBC04" italics="true" pageId="2" pageNumber="599">O. fragosus</emphasis>
</taxonomicName>
studied are lacking the distinctive protibial setal brush, but this might be the result of underdevelopment. Sclerites of the internal sac are very similar to those of
<taxonomicName id="D69472A9FF9C4A5303957E43E7A8BC67" authorityName="Castelnau" authorityYear="1840" box="[748,901,1088,1111]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="23E0D538FF9C4A5303957E43E7A8BC67" box="[748,901,1088,1111]" italics="true" pageId="2" pageNumber="599">O. marginatus</emphasis>
</taxonomicName>
. The parameres are of the same
<typeStatus id="CE2FB788FF9C4A53059F7E43E139BC67" box="[1254,1300,1088,1111]" pageId="2" pageNumber="599">type</typeStatus>
but differ in being less projected ventrally in
<taxonomicName id="D69472A9FF9C4A53008B7E61E642BC49" authorityName="Génier &amp; Howden" authorityYear="2014" box="[498,623,1122,1145]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="fragosus">
<emphasis id="23E0D538FF9C4A53008B7E61E642BC49" box="[498,623,1122,1145]" italics="true" pageId="2" pageNumber="599">O. fragosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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