<documentid="B2EE8A206E9BE510786F159FA611219D"ID-CLB-Dataset="50014"ID-DOI="10.5281/zenodo.180786"ID-GBIF-Dataset="1d889f27-71ac-4f60-b3d8-4a9ef3ea3316"ID-ISSN="1175-5326"ID-Zenodo-Dep="180786"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="felipe"checkinTime="1459816584071"checkinUser="plazi"docAuthor="Kizirian, David, Bayefsky-Anand, Sarah, Eriksson, April, Le, Minh & Donnelly, Maureen A."docDate="2008"docId="937987E5FFC9FFBBEA977FA39638B99A"docLanguage="en"docName="zt01700p062.pdf"docOrigin="Zootaxa 1700"docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article"docStyleId="6581673A57F01A3145754A1E615EDFF0"docStyleName="Zootaxa.2007-2008.journal_article"docStyleVersion="4"docTitle="Petracola ventrimaculatus Boulenger 1900"docType="treatment"docVersion="10"lastPageNumber="61"masterDocId="6F40FF9DFFCCFFB3EA007A42925CB838"masterDocTitle="A new Petracola and re-description of P. ventrimaculatus (Squamata: Gymnophthalmidae)"masterLastPageNumber="62"masterPageNumber="53"pageNumber="58"updateTime="1732482410747"updateUser="ExternalLinkService">
<bibRefCitationid="7F414B02FFC9FFB6EBE47FA390F3BDC3"author="Boulenger"box="[484,687,1505,1531]"pageId="5"pageNumber="61"refString="Boulenger, G. A. (1900) Descriptions of new batrachians and reptiles collected by Mr. P. O. Simons in Peru. Annals and Magazine of Natural History, 6, 181 - 186."type="journal article"year="1900">Boulenger, 1900</bibRefCitation>
<figureCitationid="83EB2A76FFC9FFB6EA9F7C469373BE26"box="[159,303,1540,1566]"captionStart-0="FIGURE 3"captionStart-1="FIGURE 4"captionStartId-0="5.[151,255,1401,1425]"captionStartId-1="6.[151,255,1921,1945]"captionTargetBox-0="[340,1246,182,1376]"captionTargetBox-1="[358,1221,211,1875]"captionTargetId-0="figure@5.[340,1248,182,1377]"captionTargetId-1="figure@6.[340,1248,194,1897]"captionTargetPageId-0="5"captionTargetPageId-1="6"captionText-0="FIGURE 3. Distributions of Petracola waka sp. nov. (hexagonals) and P. ventrimaculatus (triangles)."captionText-1="FIGURE 4. Dorsal (A), lateral (B), and ventral (C) views of the head of the holotype of Proctoporus ventrimaculatus Boulenger. Line equals 1 mm."httpUri-0="https://zenodo.org/record/180789/files/figure.png"httpUri-1="https://zenodo.org/record/180790/files/figure.png"pageId="5"pageNumber="58">Figures 3–4</figureCitation>
;
<tableCitationid="56520348FFC9FFB6EB3B7C4693CCBE26"box="[315,400,1540,1566]"captionStart="TABLE 1"captionStartId="2.[151,239,431,455]"captionTargetBox="[159,1421,487,1995]"captionTargetPageId="2"captionText="TABLE 1. Summary of meristic variation in Petracola ventrimaculatus and P. w a k a sp. nov."httpUri="http://table.plazi.org/id/4FAF667BFFCEFFB1EA977BED962EB9FF"pageId="5"pageNumber="58"tableUuid="4FAF667BFFCEFFB1EA977BED962EB9FF">Table 1</tableCitation>
<bibRefCitationid="7F414B02FFC9FFB6EBE27C1690C4BE53"author="Boulenger"box="[482,664,1620,1643]"pageId="5"pageNumber="61"refString="Boulenger, G. A. (1900) Descriptions of new batrachians and reptiles collected by Mr. P. O. Simons in Peru. Annals and Magazine of Natural History, 6, 181 - 186."type="journal article"year="1900">Boulenger 1900</bibRefCitation>
<bibRefCitationid="7F414B02FFC9FFB6EBDC7CF69033BEF3"author="Uzzell"box="[476,623,1716,1739]"pageId="5"pageNumber="62"refString="Uzzell, T. (1970) Teiid lizards of the genus Proctoporus from Bolivia and Peru. Postilla, 142, 1 - 39."type="journal article"year="1970">Uzzell (1970)</bibRefCitation>
<bibRefCitationid="7F414B02FFC9FFB6EBDC7C9690DDBED3"author="Kizirian"box="[476,641,1748,1771]"pageId="5"pageNumber="62"refString="Kizirian, D. A. (1996) A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs, 10, 85 - 155."type="journal article"year="1996">Kizirian (1996)</bibRefCitation>
<taxonomicNameid="DCD04D70FFC9FFB6EA977CB6909BBF33"authority="Kohler & Lehr (2004)"authorityName="Kohler & Lehr"authorityYear="2004"box="[151,711,1780,1803]"class="Reptilia"family="Gymnophthalmidae"genus="Proctoporus"kingdom="Animalia"order="Squamata"pageId="5"pageNumber="58"phylum="Chordata"rank="species"species="ventrimaculatus">
<bibRefCitationid="7F414B02FFC9FFB6EBDC7CB6909BBF33"author="Kohler"box="[476,711,1780,1803]"pageId="5"pageNumber="62"refString="Kohler, G. & Lehr, E. (2004) Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with descriptions of three new species and a key to Peruvian species. Herpetologica, 60, 501 - 518."type="journal article"year="2004">Köhler & Lehr (2004)</bibRefCitation>
</taxonomicName>
[for specimens from Amazonas and Cajamarca provinces].
<taxonomicNameid="DCD04D70FFC9FFB6EA977D5690E8BF13"authority="Doan & Castoe (2005)"authorityName="Doan & Castoe"authorityYear="2005"box="[151,692,1812,1835]"class="Reptilia"family="Gymnophthalmidae"genus="Petracola"kingdom="Animalia"order="Squamata"pageId="5"pageNumber="58"phylum="Chordata"rank="species"species="ventrimaculatus">
<bibRefCitationid="7F414B02FFC9FFB6EBC27D5690E8BF13"author="Doan"box="[450,692,1812,1835]"pageId="5"pageNumber="61"refString="Doan, T. M. & Castoe, T. A. (2005) Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143, 405 - 416."type="journal article"year="2005">Doan & Castoe (2005)</bibRefCitation>
Scales smooth, juxtaposed; prefrontals absent; supraoculars two; anteriormost superciliary extends onto dorsal surface; superciliary row usually incomplete; loreal usually absent; genials usually two; transverse dorsal scale rows 29–38; transverse ventral scale rows 17–23; palpebral disk nearly transparent; femoral pores 1–5, present in females; dark brown with pale stripes; venter pale yellow with wide black bands.
Rostral scale wider than long, taller than adjacent supralabials, in contact with frontonasal, nasals, and anteriormost supralabials. Frontonasal longer than wide, widest at posterior suture of nasal, distinctly larger than frontal. Frontal longer than wide, widest at anterior suture, with distinct extension between frontoparietals. Frontoparietals subhexagonal, in contact medially, in contact with supraoculars. Interparietal triangular. Parietals about as broad as long. Postparietals three, subequal, posterior sutures form point at middorsal line. Suproculars two, subequal in size. Superciliary series discontinuous. Anterior superciliary undivided. Anterior sutures of frontal and anteriormost superciliary form a curved line. Medial scales of anteriormost dorsal scale row much larger than adjacent posterior dorsals (
<figureCitationid="83EB2A76FFCBFFB4EE207BEF9635B9FF"box="[1056,1129,429,455]"captionStart="FIGURE 4"captionStartId="6.[151,255,1921,1945]"captionTargetBox="[358,1221,211,1875]"captionTargetId="figure@6.[340,1248,194,1897]"captionTargetPageId="6"captionText="FIGURE 4. Dorsal (A), lateral (B), and ventral (C) views of the head of the holotype of Proctoporus ventrimaculatus Boulenger. Line equals 1 mm."httpUri="https://zenodo.org/record/180790/files/figure.png"pageId="7"pageNumber="60">Fig. 4</figureCitation>
Nasal pentagonal, pierced anteriorly by nostral. Loreal absent. Frenocular quadrangular. Central palbebral scale nearly transparent. Supralabials seven. Suboculars three. Postoculars two. Supratympanic temporals three. Tympanum mostly transparent, with some pigmentation posteriorly (
<figureCitationid="83EB2A76FFCBFFB4EE0678619612BA05"box="[1030,1102,547,573]"captionStart="FIGURE 4"captionStartId="6.[151,255,1921,1945]"captionTargetBox="[358,1221,211,1875]"captionTargetId="figure@6.[340,1248,194,1897]"captionTargetPageId="6"captionText="FIGURE 4. Dorsal (A), lateral (B), and ventral (C) views of the head of the holotype of Proctoporus ventrimaculatus Boulenger. Line equals 1 mm."httpUri="https://zenodo.org/record/180790/files/figure.png"pageId="7"pageNumber="60">Fig. 4</figureCitation>
Infralabials six. Genials in two symmetrical pairs, meeting at broad midventral sutures. Pregulars subequal in size, in three regular rows. Gulars in seven rows (
<figureCitationid="83EB2A76FFCBFFB4E940783091D5BAB4"box="[832,905,626,652]"captionStart="FIGURE 4"captionStartId="6.[151,255,1921,1945]"captionTargetBox="[358,1221,211,1875]"captionTargetId="figure@6.[340,1248,194,1897]"captionTargetPageId="6"captionText="FIGURE 4. Dorsal (A), lateral (B), and ventral (C) views of the head of the holotype of Proctoporus ventrimaculatus Boulenger. Line equals 1 mm."httpUri="https://zenodo.org/record/180790/files/figure.png"pageId="7"pageNumber="60">Fig. 4</figureCitation>
).
</paragraph>
<paragraphid="1B6F36F3FFCBFFB4EAC678D890EEBB6A"blockId="7.[151,1437,151,2038]"pageId="7"pageNumber="60">Dorsals smooth, rectangular, in 35 transverse, 18 longitudinal rows (at tenth transverse ventral scale row). Ventrals smooth, in 22 transverse, 10 longitudinal rows. Dorsals and ventrals separated by approximately three rows of small to granular scales, continuous with small to granular scales on body at insertion of limbs. Cloacal plate with two large anterior and four large posterior scales. Distal portion of tail lost; dorsals rectangular, smooth, glossy, in [36] transverse rows.</paragraph>
Limbs pentadactyl, with claws. Anterodorsal scales smooth, glossy, polygonal, distinctly larger than those on posterior side. Dorsal scales on brachium much larger than adjacent scales. Ventral scales on brachium small to subgranular. Anterior scales on antebrachium polygonal, smooth, glossy. Posteroventral scales on antebrachium roundish, smooth, glossy. Palmar scales granular. Dorsal scales on
/6. Scales on anterodorsal surface of thigh polygonal, smooth, glossy, distinctly larger than adjacent scales. Scales on anteroventral surface of thigh polygonal to round, smooth, glossy. Femoral pores 3/3. Scales on posterior surface of thigh small to granular. Scales on dorsal surface of crus rounded diamonds, subimbricate, smooth, glossy. Scales on anteroventral surface of crus distinctly larger than adjacent scales. Scales on dorsal surface of foot polygonal, irregularly arranged, subimbricate, of varying sizes. Dorsal scales on
<paragraphid="1B6F36F3FFCBFFB4EAC67F799017BE23"blockId="7.[151,1437,151,2038]"pageId="7"pageNumber="60">Overall color pattern faded, dark brown. Pale dorsolateral stripe bordered by black, especially distinct anteriorly. Dorsal surface of head brown, without distinct markings. Large irregular black markings laterally on head and neck. Weakly organized ocelli near insertion of forelimbs. Ventral surface of head yellow with scattered, large, dark brown markings. Venter yellow with thick, dark brown bands with irregular posterior borders. Subcaudal pattern consisting of black spots proximally, ocelli distally. Irregularly arranged yellow, brown, and black markings on limbs.</paragraph>
“Dorsum dull brown with tan stripes, becoming dull reddish tan on tail. Throat cream with black spots; belly and underside of tail black with cream borders to scales. In adult males cream areas dull red (William E. Duellman field notes for KU 181966–76)...Labial region dull yellowish tan....Juveniles with reddish tint on tail and creamy tan venters; adults with black venter. Iris red-brown” (William E. Duellman field notes for KU 181943–76). “Dorsum black or dark brown with two orange-brown dorsal lateral stripes. Flanks same with three longitudinal lines of lighter orange-brown spots. Dorsum of head is mottled black and brown. Venter is gray with few orange-brown spots. Tail dark, with five longitudinal orange-brown stripes. Iris intense red” (Erik R. Wild field notes for KU 219835). Dorsum reddish brown with pinkish orange stripes, throat pale orange (William E. Duellman field notes for KU 212689–707). Whereas specimens from west of the
<figureCitationid="83EB2A76FFC4FFBBEE7A7A82969FB8E2"box="[1146,1219,192,218]"captionStart="FIGURE 3"captionStartId="5.[151,255,1401,1425]"captionTargetBox="[340,1246,182,1376]"captionTargetId="figure@5.[340,1248,182,1377]"captionTargetPageId="5"captionText="FIGURE 3. Distributions of Petracola waka sp. nov. (hexagonals) and P. ventrimaculatus (triangles)."httpUri="https://zenodo.org/record/180789/files/figure.png"pageId="8"pageNumber="61">Fig. 3</figureCitation>
). Adults and eggs have been found under rocks in cloud forest or along roads (Erik R. Wild field notes for KU 219835–38), under rocks in bunch grass-
association (William E. Duellman field notes for KU 181943–76), and under rocks in “wet paramo” and plowed fields (William E. Duellman field notes for KU 212689–707).
