200 lines
21 KiB
XML
200 lines
21 KiB
XML
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<document id="D8E58185DD782ED0A605737186589A56" ID-CLB-Dataset="31534" ID-DOI="http://dx.doi.org/10.3897/zookeys.715.20497" ID-GBIF-Dataset="138d6982-b157-4301-a3a0-72029f9044ba" ID-PMC="PMC5740420" ID-Pensoft-Pub="1313-2970-715-69" ID-PubMed="29302233" ID-ZooBank="763EF053AFAA41119CA96060F9E09E70" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-715-69" ModsDocOrigin="ZooKeys 715" ModsDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" checkinTime="1510736981334" checkinUser="pensoft" docAuthor="Wang, Cheng-Bin, Perreau, Michel, Ruzicka, Jan & Nishikawa, Masaaki" docDate="2017" docId="1852CF88F7F144C2672F5F4EE28C9614" docLanguage="en" docName="ZooKeys 715: 69-92" docOrigin="ZooKeys 715" docSource="http://dx.doi.org/10.3897/zookeys.715.20497" docTitle="Ptomaphagus nepalensis Perreau 1988" docType="treatment" docVersion="6" lastPageNumber="75" masterDocId="7A4E6F14FFE5FFD08743FFBA6574A77E" masterDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" masterLastPageNumber="92" masterPageNumber="69" pageNumber="73" updateTime="1701375221787" updateUser="ExternalLinkService">
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<mods:titleInfo id="98C4E2258523BFFEDBAA98C7FB7BC28B">
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<mods:title id="F6284B95ABD1EAF6AC0F319AFB97B35F">Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)</mods:title>
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<mods:namePart id="7D49897450A477F51312DBA3ADAD6F75">Wang, Cheng-Bin</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="67E883DD48B8FB53CAD942C915EEFBF1">Perreau, Michel</mods:namePart>
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<mods:namePart id="4D6803D8FC24E5C23496B7E1E9E91373">Ruzicka, Jan</mods:namePart>
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</mods:role>
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<mods:namePart id="608EFC5EE834C5F707F37539849A99A8">Nishikawa, Masaaki</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B5DCB07495AD25B8B3E41344D311EBCB">text</mods:typeOfResource>
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<mods:title id="D85C0B2B24D652CA2F8A1A901874E585">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="051947DA85D4B0F5D38D56901ABA4643">
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<mods:date id="A07EBD32EF393E77AC4D4C962C80C074">2017</mods:date>
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<mods:detail id="21B0E2324BDAF9677EB25D9D5B47CB9A" type="volume">
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<mods:number id="CDEACA91E7042E5C6A640A08AE6F548E">715</mods:number>
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<mods:start id="82D065608B689E7E4BCFAF52DB7F834B">69</mods:start>
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<mods:url id="78CDBCAEB567F11BE39BC3A88FAB0FCA">http://dx.doi.org/10.3897/zookeys.715.20497</mods:url>
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<mods:classification id="E02D37087372CC376FF0C35C61282220">journal article</mods:classification>
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<mods:identifier id="366BECA1F824450B965500B8E42DD25E" type="DOI">http://dx.doi.org/10.3897/zookeys.715.20497</mods:identifier>
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<mods:identifier id="673555405A94946B67F1706BCA64A57A" type="Pensoft-Pub">1313-2970-715-69</mods:identifier>
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<mods:identifier id="C206E053D7652F0A6166F126BFCCEEFF" type="ZooBank">763EF053AFAA41119CA96060F9E09E70</mods:identifier>
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</mods:mods>
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<treatment id="1852CF88F7F144C2672F5F4EE28C9614" ID-GBIF-Taxon="137306769" LSID="urn:lsid:plazi:treatment:1852CF88F7F144C2672F5F4EE28C9614" httpUri="http://treatment.plazi.org/id/1852CF88F7F144C2672F5F4EE28C9614" lastPageId="6" lastPageNumber="75" pageId="4" pageNumber="73">
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<subSubSection id="C29809FE9A796F592AC57C2F70F838EB" pageId="4" pageNumber="73" type="nomenclature">
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<paragraph id="7437B0651DE8236D30C195EDA156BF59" pageId="4" pageNumber="73">
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<taxonomicName id="9F2196B859B1794485B191609F7BDCDC" ID-CoL="78M2C" authority="Perreau, 1988" authorityName="Perreau" authorityYear="1988" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus nepalensis" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="nepalensis">Ptomaphagus (s. str.) nepalensis Perreau, 1988</taxonomicName>
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Figs 1
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<normalizedToken id="7CBF0F7E03AEFA8D2B43581D6A170DBC" originalValue="A–B">A-B</normalizedToken>
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; 2
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<normalizedToken id="F31AA58F0B9047654BC6F5DA8BC180CB" originalValue="A–J">A-J</normalizedToken>
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; 3
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<normalizedToken id="035D1C0A614D6E50CE90BC69521D0621" originalValue="A–F">A-F</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="7375046D4ACE989343F538DF96D6579E" pageId="4" pageNumber="73" type="reference_group">
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<paragraph id="C5974C2306ED144CEBA21079BF93D592" pageId="4" pageNumber="73">
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<taxonomicName id="D52E0AB816D4D214E457785041470CB9" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus nepalensis" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="nepalensis">Ptomaphagus (s. str.) nepalensis</taxonomicName>
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Perreau, 1988: 1005 (
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<taxonomicName id="E0CA26FE8B2209D4E35D44492927939E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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; type locality:
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<normalizedToken id="3BDE29ADE3718D9270733C47B3EE8EDC" originalValue="Népal">Nepal</normalizedToken>
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, district de Lalitpur, Phulcoki [ca.
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<geoCoordinate id="7172EFBEE996DAF341CD29DCBA171E21" direction="north" orientation="latitude" precision="925" value="27.566668">27°34'N</geoCoordinate>
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<geoCoordinate id="8BDDFFBA872DA865EC319D8C193C0E0D" direction="east" orientation="longitude" precision="925" value="85.416664">085°25'E</geoCoordinate>
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], 2600 m; MHNG); Perreau, 2000: 363 (
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<taxonomicName id="4379AE51D15CA3CE2511D490F860CDA4" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.); in catalog); Perreau, 2004: 178 (
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<taxonomicName id="46A9145C4FE2B43F0926DEBE3E0014A8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(
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<taxonomicName id="4565EB8A7960459CFD9EC160D734E1A7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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); in catalog); Perreau, 2015: 249 (
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<taxonomicName id="0297ECB585A7682F769D05BFBAEAB322" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
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; in catalog).
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</paragraph>
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</subSubSection>
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<subSubSection id="D191FD657CF393ED8E55C1340AA1505E" pageId="4" pageNumber="73" type="materials_examined">
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<paragraph id="286FD4A98378F912479018F845AB019A" pageId="4" pageNumber="73">Material examined.</paragraph>
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<paragraph id="2EE44221027CBF2BB523FAACAF1640FA" pageId="4" pageNumber="73">
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Type material. Paratypes: 1♂, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 20.IV.1982 / A. & Z. Smetana // PARATYPE (MHNG); 1♂, NEPAL Lalitpur / Distr. Phulcoki / 2600 m 16.X.[19]83 / Smetana &
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<normalizedToken id="8EA0783D8326A8610703393DDF537DA2" originalValue="Löbl">Loebl</normalizedToken>
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//
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<taxonomicName id="7D1B7AE4D2F40243362775C29B1610C7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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/
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<taxonomicName id="DA7FA956A295D302DB3D45FBB3B747E3" class="Insecta" family="Leiodidae" genus="Ptomaphagina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ptomaphagina nepalensis" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="nepalensis">nepalensis</taxonomicName>
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/
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<bibRefCitation id="C0530493BCD9F7A3676E6FC79ADBB7DC" author="Perreau, M" journalOrPublisher="Revue suisse de Zoologie" pageId="19" pageNumber="88" pagination="1005 - 1018" title="Les Cholevidae himalayiens du Museum d'Histoire naturelle de Geneve." url="https://doi.org/10.5962/bhl.part.81947" volume="95" year="1988">Perreau 1988</bibRefCitation>
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// PARATYPE // MHNG / ENTO / 00003344 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2600 m 14.X.[19]83 / Smetana &
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<normalizedToken id="5EB227CD0F3B11952609EDC6D37C7904" originalValue="Löbl">Loebl</normalizedToken>
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// PARATYPE // MHNG / ENTO / 00003345 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2650 m 15.X.[19]83 / Smetana &
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<normalizedToken id="F4F1FB8107BC231042C33265016AED23" originalValue="Löbl">Loebl</normalizedToken>
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// PARATYPE // MHNG / ENTO / 00003346 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2650 m 14.X.[19]83 / Smetana &
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<normalizedToken id="D91174BDC71ED6D172006DF8E0C0717C" originalValue="Löbl">Loebl</normalizedToken>
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// PTOMAPHAGUS / NEPALENSIS n. sp. / M. PERREAU det. 1987 // PARATYPE // MHNG / ENTO / 00003347 (MHNG); 1♀, NEPAL Lalitpur / Distr. Phulcoki / 2700 m 15.X.[19]83 / Smetana &
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<normalizedToken id="CC0361D18DFD16C21E0E73F3A5B587F5" originalValue="Löbl">Loebl</normalizedToken>
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// PARATYPE (MHNG); 1♂, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 20.IV.1982 / A. & Z. Smetana // PARATYPE // MHNG / ENTO / 00003343 (MHNG); 1♀, NEPAL, Kath- / mandu District // Phulcoki 2600 m / 22.IV.1982 / A. & Z. Smetana // PARATYPE // MHNG / ENTO / 00003342 (MHNG); 2♂♂, NEPAL: distr. / Kathmandu: Phulcoki / 2500 m, 28-29.IV.[19]84 /
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<normalizedToken id="6FED3966D3F8583473996B26331FD95F" originalValue="Löbl">Loebl</normalizedToken>
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- Smetana (CMPR).
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</paragraph>
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</subSubSection>
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<subSubSection id="D5E62C6BE51894E75F2EB92F1EB07DA3" lastPageId="6" lastPageNumber="75" pageId="4" pageNumber="73" type="redescription">
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<paragraph id="C2779B3D5077C073AB2A5EE4C81E67D3" pageId="4" pageNumber="73">Redescription.</paragraph>
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<paragraph id="B5F8B9C80FD273729B9AE64F770D960B" pageId="4" pageNumber="73">
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Male. EBL: 2.9-3.0 mm. Length of different body parts: HL: AL: PL: ELL = 0.5: 0.9: 0.7: 1.6 mm; width: HW: EW: PW: ELW = 0.8: 0.1: 1.1: 1.1 mm. Proportion of antennomeres from base to tip in
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<normalizedToken id="9F48ED61BA9923CEA1E51C598B26EF2C" originalValue="μm">μm</normalizedToken>
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(length
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<normalizedToken id="66152720CCDD9B2E2BBC111812AE25CC" originalValue="×">x</normalizedToken>
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width): 132
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<normalizedToken id="2C446D72AE6155E4859B55EB0FD4912E" originalValue="×">x</normalizedToken>
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54, 89
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50, 69
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89, 29
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94.
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</paragraph>
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<paragraph id="6B93DB2722961F81F297B5189EE26225" pageId="4" pageNumber="73">Habitus (Fig. 1A) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, basal three or four antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi more or less yellowish. Dorsum continually clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
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<caption id="816C848E5D5B310C4A5EB37B5A3D0CD6" pageId="4" pageNumber="73">
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<paragraph id="74A96383660A336CD667779A32CE0E7B" pageId="4" pageNumber="73">
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Figure 1. Habitus of
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<taxonomicName id="FEDD777F52A36BB3DC531EBACA438124" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(s. str.) spp. (dorsal view).
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<normalizedToken id="DE5C4DBB0B31648D4C9BE692CF3C66E4" originalValue="A–B">A-B</normalizedToken>
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<taxonomicName id="EE0EA058327AFA86CB887001F97B5ECA" lsidName="P. nepalensis" pageId="4" pageNumber="73" rank="species" sensu="(s. str.)" species="nepalensis">P. (s. str.) nepalensis</taxonomicName>
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Perreau, 1988 A ♂ (paratype; Nepal) B ♀ (paratype; Nepal)
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<normalizedToken id="03D03B5646E0F4BD3075A5E27565473D" originalValue="C–D">C-D</normalizedToken>
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<taxonomicName id="FC0B96AD2DE3FF686CFE96AE9B8E5143" lsidName="P. masumotoi" pageId="4" pageNumber="73" rank="species" sensu="(s. str.)" species="masumotoi">P. (s. str.) masumotoi</taxonomicName>
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Nishikawa, 2011 C ♂ (paratype; Thailand) D ♀ (holotype; Thailand) E
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<taxonomicName id="9930BB85E66580D0EF8CE3AB7C6716B4" lsidName="P. piccoloi" pageId="4" pageNumber="73" rank="species" sensu="(s. str.)" species="piccoloi">P. (s. str.) piccoloi</taxonomicName>
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Wang,
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<normalizedToken id="1A5CD40DB9F7B0659BCDD244F7D75191" originalValue="Růžička">Ruzicka</normalizedToken>
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, Nishikawa, Perreau & Hayashi, 2016 ♂ (China: Zhejiang)
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<normalizedToken id="B905E68670DF6B47954D4F84071BAAB1" originalValue="F–G">F-G</normalizedToken>
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<taxonomicName id="D41E052BC7786C1E85B6F49B2403977F" lsidName="P. funiu" pageId="4" pageNumber="73" rank="species" sensu="(s. str.)" species="funiu">P. (s. str.) funiu</taxonomicName>
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sp. n. F ♂ (holotype; China: Henan) G ♀ (paratype; China: Henan)
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<normalizedToken id="CBF506B2B770583EE4D2D78FA483723F" originalValue="H–I">H-I</normalizedToken>
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<taxonomicName id="2F86B17D5878143F4EFA55DE55AC1336" lsidName="P. haba" pageId="4" pageNumber="73" rank="species" sensu="(s. str.)" species="haba">P. (s. str.) haba</taxonomicName>
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sp. n. H ♂ (holotype; China: Yunnan) I ♀ (paratype; China: Yunnan). Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="35692B2DC3E52AE3130FCFCF3F3C01DA" pageId="4" pageNumber="73">Head transverse, HW/HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin almost straight. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 2A) slender, AL/HW = 1.1; antennomere III shorter than II; VI with length/width = 0.5; XI pear-shape.</paragraph>
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<paragraph id="58AC56622399E3212F650F7E9AEDE3E6" lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73">
|
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Pronotum (Fig. 2B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, gradually narrowing from posterior to anterior; hind angles slightly pro
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<pageBreakToken id="4AD7A3BC60E2C91E49D5A4BD7042C752" pageId="5" pageNumber="74" start="start">jected</pageBreakToken>
|
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|
backwards and subacute. Posterior margin widely but shortly protruded in middle part, emarginate near hind angles.
|
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</paragraph>
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<paragraph id="55E10083A13FA7E542EF433DB9D198AA" pageId="5" pageNumber="74">Elytra oval, widest at about basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 2G) rounded. Sutural striae present. Metathoracic wings fully developed.</paragraph>
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<paragraph id="4A5C2EFF9EA6F9C23D25F402E858351B" pageId="5" pageNumber="74">Prolegs robust, with basal three protarsomeres (Fig. 2C) expanded: TW/BTW = 1.3. Protibiae (Fig. 2E) expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender, straight, but relatively short.</paragraph>
|
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<paragraph id="B2BB3F5C1DEA0A0E4B1669603AC2BEF4" pageId="5" pageNumber="74">Abdominal ventrite VIII (Fig. 2I) distinctly emarginate at posterior edge, and much deeper at median. Spiculum gastrale of genital segment (Fig. 2J) with about 2/5 of length protruding beyond anterior edge of epipleurite IX.</paragraph>
|
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<caption id="93DB47FFF39F59AB81A88CED633D5761" pageId="5" pageNumber="74">
|
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|
<paragraph id="B7F1225464A75B320AF9AC66E670F252" pageId="5" pageNumber="74">
|
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|
Figure 2.
|
|||
|
<taxonomicName id="C0C4310DEF47DBDD1726EA89E1B33997" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus nepalensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="nepalensis">Ptomaphagus (s. str.) nepalensis</taxonomicName>
|
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|
Perreau, 1988 (♂: paratype; ♀: paratype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars: 0.1 mm.
|
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</paragraph>
|
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|
</caption>
|
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|
<paragraph id="2C1702007538F6BB44F57F31344CF81F" pageId="6" pageNumber="75">
|
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|
<pageBreakToken id="416419E45D9945D950EDA6537CE47E41" pageId="6" pageNumber="75" start="start">Aedeagus</pageBreakToken>
|
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|
(Fig. 3A) rather long and slender, with median lobe gradually narrowing towards a lanceolate apex and terminated to a widely subrounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 3C) inserted with two ventrally oriented setae on the left side and three ventrally oriented setae on the right side; parameres narrow, reaching about apical 1/7 of median lobe, each apex (Fig. 3D) with two long lateral setae and one shorter apical seta. In lateral view (Fig. 3B), median lobe slender, regularly bent ventrally, and gradually tapering to a round apex. Endophallus with stylus quite slender, a transverse nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.
|
|||
|
</paragraph>
|
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|
<paragraph id="592469F0CC4A70D5639F43395AD56D7A" pageId="6" pageNumber="75">Female. Similar to male in general appearance (Fig. 1B), including elytral apices (Fig. 2H), but distinguished by the following characteristics: protarsi (Fig. 2D) simply linear; protibiae (Fig. 2F) narrower; abdominal ventrite VIII (Fig. 3E) almost rounded at posterior edge and slightly protruded at median; genital segment and ovipositor as shown in Fig. 3F; spermatheca (Fig. 3F) sinuous or coiled in distal part, not coiled in proximal part.</paragraph>
|
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|
<caption id="EA1178B64A6C1D3CC02DC9780436B730" pageId="6" pageNumber="75">
|
|||
|
<paragraph id="F41BB458301033AA7FAD6EC090B096EE" pageId="6" pageNumber="75">
|
|||
|
Figure 3.
|
|||
|
<taxonomicName id="0D99AEDAA3848C0367634049087A4F58" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus nepalensis" order="Coleoptera" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="nepalensis">Ptomaphagus (s. str.) nepalensis</taxonomicName>
|
|||
|
Perreau, 1988 (♂: paratype; ♀: paratype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (dorsal view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scale bars: 0.1 mm.
|
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</paragraph>
|
|||
|
</caption>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="A297065B110A88A0ADBA98EAF41EC2C9" pageId="6" pageNumber="75" type="distribution">
|
|||
|
<paragraph id="DE801771C2075FC3F9749397F5887C37" pageId="6" pageNumber="75">Distribution.</paragraph>
|
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|
<paragraph id="45D83076851407F9B96D2C00727475AF" pageId="6" pageNumber="75">Nepal.</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
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|
</document>
|