treatments-xml/data/03/B0/87/03B087E55B4BFFBDFF79F2BAFE2AFA0D.xml

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<document id="9AA95507F93D713D322269DB2356486A" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B4BFFBDFF79F2BAFE2AFA0D" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena marawaka Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="47" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="47" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FD32EAECB993F9628FBD47ABB3EB6E1E">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="6A683EC413163F956C5653DF81C15F02">Perkins, Philip D.</mods:namePart>
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<mods:date id="9C606BF52C665833AD5E9BF54424DB1D">2011</mods:date>
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<treatment id="03B087E55B4BFFBDFF79F2BAFE2AFA0D" ID-DOI="http://doi.org/10.5281/zenodo.5291449" ID-Zenodo-Dep="5291449" LSID="urn:lsid:plazi:treatment:03B087E55B4BFFBDFF79F2BAFE2AFA0D" httpUri="http://treatment.plazi.org/id/03B087E55B4BFFBDFF79F2BAFE2AFA0D" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection id="C30365785B4BFFBDFF79F2BAFD82FF27" box="[151,564,151,177]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA636F35B4BFFBDFF79F2BAFD82FF27" blockId="46.[151,564,151,210]" box="[151,564,151,177]" pageId="46" pageNumber="47">
<heading id="D0EE819F5B4BFFBDFF79F2BAFD82FF27" bold="true" box="[151,564,151,177]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B96DEAE15B4BFFBDFF79F2BAFD82FF27" bold="true" box="[151,564,151,177]" pageId="46" pageNumber="47">
<taxonomicName id="4C194D705B4BFFBDFF79F2BAFE23FF27" ID-CoL="3N3DD" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,405,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marawaka" status="sp. nov.">
<emphasis id="B96DEAE15B4BFFBDFF79F2BAFE23FF27" bold="true" box="[151,405,151,177]" italics="true" pageId="46" pageNumber="47">Hydraena marawaka</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B4BFFBDFE4AF2BAFD82FF27" box="[420,564,151,177]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF79F297FED0FF44" box="[151,358,186,210]" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA636F35B4BFFBDFF79F297FED0FF44" blockId="46.[151,564,151,210]" box="[151,358,186,210]" pageId="46" pageNumber="47">
(
<figureCitation id="13222A765B4BFFBDFF71F297FF4BFF44" box="[159,253,186,210]" captionStart="FIGURE 62" captionStartId="191.[151,250,1745,1768]" captionTargetBox="[199,1382,209,1713]" captionTargetId="figure-26@191.[186,1401,193,1724]" captionTargetPageId="191" captionText="FIGURE 62. Hydraena marawaka, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283522" httpUri="https://zenodo.org/record/5283522/files/figure.png" pageId="46" pageNumber="47">Figs. 62</figureCitation>
,
<figureCitation id="13222A765B4BFFBDFEE6F297FE90FF44" box="[264,294,186,210]" captionStart="FIGURES 6465" captionStartId="193.[151,264,1650,1673]" captionText="FIGURES 6465. Aedeagi of holotypes. 64. H. marawaka. 65. H. sexarcuata." pageId="46" pageNumber="47">64</figureCitation>
,
<figureCitation id="13222A765B4BFFBDFEDCF297FEE8FF44" box="[306,350,186,210]" captionStart="FIGURES 447450" captionStartId="387.[151,264,1650,1673]" captionTargetBox="[165,1421,245,1593]" captionTargetId="figure-20@387.[165,1421,193,1629]" captionTargetPageId="387" captionText="FIGURES 447450. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284079" httpUri="https://zenodo.org/record/5284079/files/figure.png" pageId="46" pageNumber="47">450</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF79F32EFB2FFEA8" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BA636F35B4BFFBDFF79F32EFB2FFEA8" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">
Type Material.
<materialsCitation id="3B713CAE5B4BFFBDFEBFF32EFDD7FEA8" collectingDate="2006-11-11" collectionCode="ZSM" collectorName="Balke &amp; Kinibel" country="Papua New Guinea" elevation="1400" latitude="-7.0944" location="Marawaka" longLatPrecision="1" longitude="145.74112" municipality="Mala" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Gulf" typeStatus="holotype">
<typeStatus id="54A288515B4BFFBDFEBFF32EFE08FE8D" box="[337,446,259,283]" pageId="46" pageNumber="47" type="holotype">
<emphasis id="B96DEAE15B4BFFBDFEBFF32EFE08FE8D" bold="true" box="[337,446,259,283]" pageId="46" pageNumber="47">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B4BFFBDFDC6F32FFD6FFE8D" bold="true" box="[552,729,258,283]" pageId="46" pageNumber="47">
<collectingRegion id="49DDF8115B4BFFBDFDC6F32FFD63FE8D" box="[552,725,258,283]" country="Papua New Guinea" name="Gulf" pageId="46" pageNumber="47">Gulf Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B4BFFBDFD04F32FFCD5FE8C" LSID="urn:lsid:plazi:treatment:03B087E55B4BFFBDFF79F2BAFE2AFA0D:8EC660285B4BFFBDFD04F32FFCD5FE8C" box="[746,867,258,282]" country="Papua New Guinea" latitude="-7.0944" longLatPrecision="1" longitude="145.74112" municipality="Mala" name="Marawaka" pageId="46" pageNumber="47" stateProvince="Gulf">Marawaka</location>
,
<collectingMunicipality id="6BC2AC895B4BFFBDFC81F32FFC1DFE8C" box="[879,939,258,282]" pageId="46" pageNumber="47">Mala</collectingMunicipality>
,
<quantity id="4CE19B165B4BFFBDFC56F32FFBA7FE8C" box="[952,1041,258,282]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="47" unit="m" value="1400.0">
<elevation id="0034D1C05B4BFFBDFC56F32FFBA7FE8C" box="[952,1041,258,282]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="47" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B4BFFBDFBF0F32FFB2BFE8C" box="[1054,1181,258,282]" degrees="7" direction="south" minutes="5.664" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="-7.0944">7° 5.664' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B4BFFBDFB45F32FFAE3FE8C" box="[1195,1365,258,282]" degrees="145" direction="east" minutes="44.467" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="145.74112">145° 44.467' E</geoCoordinate>
,
<date id="FFA710335B4BFFBDFA8DF32FFF64FEA8" pageId="46" pageNumber="47" value="2006-11-11">
<collectingDate id="EFE3E9DB5B4BFFBDFA8DF32FFF64FEA8" pageId="46" pageNumber="47" value="2006-11-11">11 xi 2006</collectingDate>
</date>
,
<collectorName id="26EC53255B4BFFBDFF33F30BFEA8FEA8" box="[221,286,294,318]" pageId="46" pageNumber="47">Balke</collectorName>
&amp;
<collectorName id="26EC53255B4BFFBDFEACF30BFE20FEA8" box="[322,406,294,318]" pageId="46" pageNumber="47">Kinibel</collectorName>
(
<collectingCountry id="F30E76635B4BFFBDFE4AF30BFE68FEA8" box="[420,478,294,318]" name="Papua New Guinea" pageId="46" pageNumber="47">PNG</collectingCountry>
90) (
<collectionCode id="ED08AE365B4BFFBDFDF4F30BFDEFFEA8" box="[538,601,294,318]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="46" pageNumber="47" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B4BFFBDFD82F30AFB23FEA8" box="[620,1173,294,319]" collectingDate="2006-11-11" collectionCode="ZSM" collectorName="Balke &amp; Kinibel" country="Papua New Guinea" elevation="1400" latitude="-7.0944" location="Marawaka" longLatPrecision="1" longitude="145.74112" municipality="Mala" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Paratype" typeStatus="paratype">
<emphasis id="B96DEAE15B4BFFBDFD82F30AFD6FFEA9" bold="true" box="[620,729,295,319]" pageId="46" pageNumber="47">
<typeStatus id="54A288515B4BFFBDFD82F30AFD6FFEA9" box="[620,729,295,319]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
</emphasis>
: same data as holotype (
<specimenCount id="9D1FFD7A5B4BFFBDFC19F30BFBF5FEA8" box="[1015,1091,294,318]" pageId="46" pageNumber="47" type="male">1 male</specimenCount>
,
<collectionCode id="ED08AE365B4BFFBDFBA0F30BFB3AFEA8" box="[1102,1164,294,318]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="46" pageNumber="47" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF29F367FE83FE64" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BA636F35B4BFFBDFF29F367FE83FE64" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">
<emphasis id="B96DEAE15B4BFFBDFF29F367FE64FEF5" bold="true" box="[199,466,330,355]" pageId="46" pageNumber="47">Differential Diagnosis.</emphasis>
Similar in dorsal habitus to
<taxonomicName id="4C194D705B4BFFBDFCE3F366FC0FFEF5" authorityName="Perkins" authorityYear="2011" box="[781,953,331,355]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cheesmanae">
<emphasis id="B96DEAE15B4BFFBDFCE3F366FC97FEF4" box="[781,801,331,354]" italics="true" pageId="46" pageNumber="47">H</emphasis>
.
<emphasis id="B96DEAE15B4BFFBDFCDEF366FC0FFEF5" box="[816,953,331,355]" italics="true" pageId="46" pageNumber="47">cheesmanae</emphasis>
</taxonomicName>
; differing therefrom in larger size (ca. 1.56 vs. 1.37), and metaventral plaque shape, which are more widely separated and have raised lateral margins (
<figureCitation id="13222A765B4BFFBDFAD4F343FA21FE10" box="[1338,1431,366,390]" captionStart="FIGURE 59" captionStartId="189.[151,250,1731,1754]" captionTargetBox="[202,1383,211,1696]" captionTargetId="figure-26@189.[186,1401,193,1710]" captionTargetPageId="189" captionText="FIGURE 59. Hydraena cheesmanae, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283514" httpUri="https://zenodo.org/record/5283514/files/figure.png" pageId="46" pageNumber="47">Figs. 59</figureCitation>
,
<figureCitation id="13222A765B4BFFBDFF79F3BFFF01FE3C" box="[151,183,402,426]" captionStart="FIGURE 62" captionStartId="191.[151,250,1745,1768]" captionTargetBox="[199,1382,209,1713]" captionTargetId="figure-26@191.[186,1401,193,1724]" captionTargetPageId="191" captionText="FIGURE 62. Hydraena marawaka, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283522" httpUri="https://zenodo.org/record/5283522/files/figure.png" pageId="46" pageNumber="47">62</figureCitation>
). The male metatibiae differ very slightly in shape in the two species, but
<taxonomicName id="4C194D705B4BFFBDFC14F3BEFB25FE3D" authorityName="Perkins" authorityYear="2011" box="[1018,1171,403,427]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marawaka">
<emphasis id="B96DEAE15B4BFFBDFC14F3BEFBB8FE3C" box="[1018,1038,403,426]" italics="true" pageId="46" pageNumber="47">H</emphasis>
.
<emphasis id="B96DEAE15B4BFFBDFBF3F3BEFB25FE3D" box="[1053,1171,403,427]" italics="true" pageId="46" pageNumber="47">marawaka</emphasis>
</taxonomicName>
has longer setae on the brush. The male genitalia of the two species also suggest a relationship, but differ distinctively in many details (
<figureCitation id="13222A765B4BFFBDFF71F3F7FF4AFE64" box="[159,252,474,498]" captionStart="FIGURES 6061" captionStartId="190.[151,264,1650,1673]" captionTargetBox="[292,1287,214,1603]" captionTargetId="figure-32@190.[250,1336,193,1629]" captionTargetPageId="190" captionText="FIGURES 6061. Aedeagi of holotypes. 60. H. remulipes. 61. H. cheesmanae." figureDoi="http://doi.org/10.5281/zenodo.5283516" httpUri="https://zenodo.org/record/5283516/files/figure.png" pageId="46" pageNumber="47">Figs. 61</figureCitation>
,
<figureCitation id="13222A765B4BFFBDFEE6F3F7FE9EFE64" box="[264,296,474,498]" captionStart="FIGURES 6465" captionStartId="193.[151,264,1650,1673]" captionText="FIGURES 6465. Aedeagi of holotypes. 64. H. marawaka. 65. H. sexarcuata." pageId="46" pageNumber="47">64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF29F3D3FDC4FAB8" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA636F35B4BFFBDFF29F3D3FD5BFD14" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">
<emphasis id="B96DEAE15B4BFFBDFF29F3D3FEEEFD81" bold="true" box="[199,344,510,535]" inLineHeading="true" pageId="46" pageNumber="47" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B4BFFBDFE42F3D3FDB8FD80" box="[428,526,510,534]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.56/0.64; head 0.23/0.36; pronotum 0.39/0.45, PA 0.39, PB 0.39; elytra 0.92/0.64. Dorsum of head piceous; pronotum testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca. 7/14/5; elytra dark brown; legs light brown; maxillary palpi testaceous, tip slightly darker.
</paragraph>
<paragraph id="8BA636F35B4BFFBDFF29F0A2FA20FCA0" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">Frons punctures ca. 12xef, larger near eyes than medially; interstices shining, 15xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum very sparsely very finely punctulate, shining; postmentum effacedly micropunctulate. Genae raised, shining, without posterior ridge. Pronotum cordiform, median 2/ 3 of anterior margin emarginate; punctures on disc ca. 1xpd those of frons, interstices shining, 14xpd, punctures slightly larger and denser at anterior and posterior; PF1 absent; PF2 very shallow, obsolete; PF3 deep; PF4 absent.</paragraph>
<paragraph id="8BA636F35B4BFFBDFF29F16FFF47FC7C" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 13xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B4BFFBDFF29F1DBFDC4FAB8" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/3/8/4. P1 laminate; median carina sinuate in profile. P2 raised, l/w ca. 2/1, sides slightly converging toward blunt apex. Plaques large, converging slightly anteriorly, transversely raised, more so laterally than medially, located at sides of deep median depression, each plaque tapering slightly from posterior to anterior. Metaventrite without midlongitudinal ridge. AIS width at straight posterior margin ca. 1.7x P2. All legs long and slender. Profemur (male) with small, sharply pointed tubercle next to trochanter; protibia straight, widest subapically, cluster of small spines on anteromedial margin at this point, moderately long spine on medial surface at ca. distal 1/3. Mesotibia straight. Metatibia straight, medial margin in distal 1/2 emarginate and bearing sparse brush of long hair-like setae. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B4BFFBDFDFFF73BFDD3FAB8" box="[529,613,1302,1326]" captionStart="FIGURES 6465" captionStartId="193.[151,264,1650,1673]" captionText="FIGURES 6465. Aedeagi of holotypes. 64. H. marawaka. 65. H. sexarcuata." pageId="46" pageNumber="47">Fig. 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF29F716FC47FAC4" box="[199,1009,1338,1363]" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8BA636F35B4BFFBDFF29F716FC47FAC4" blockId="46.[151,1437,258,1435]" box="[199,1009,1338,1363]" pageId="46" pageNumber="47">
<emphasis id="B96DEAE15B4BFFBDFF29F716FEFBFAC5" bold="true" box="[199,333,1339,1363]" inLineHeading="true" pageId="46" pageNumber="47" reason="1">Etymology.</emphasis>
Named in reference to the known geographical distribution.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B4BFFBDFF29F773FE2AFA0D" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8BA636F35B4BFFBDFF29F773FE2AFA0D" blockId="46.[151,1437,258,1435]" pageId="46" pageNumber="47">
<emphasis id="B96DEAE15B4BFFBDFF29F773FED6FAE1" bold="true" box="[199,352,1374,1399]" inLineHeading="true" pageId="46" pageNumber="47" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B4BFFBDFD27F772FD4FFAE1" box="[713,761,1375,1399]" pageId="46" pageNumber="47">type</typeStatus>
locality near the eastern and southern limit of Area 1; elevation
<quantity id="4CE19B165B4BFFBDFF27F7AFFEA8FA0C" box="[201,286,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="47" unit="m" value="1400.0">1400 m</quantity>
(
<figureCitation id="13222A765B4BFFBDFEC3F7AFFE38FA0C" box="[301,398,1410,1434]" captionStart="FIGURES 447450" captionStartId="387.[151,264,1650,1673]" captionTargetBox="[165,1421,245,1593]" captionTargetId="figure-20@387.[165,1421,193,1629]" captionTargetPageId="387" captionText="FIGURES 447450. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284079" httpUri="https://zenodo.org/record/5284079/files/figure.png" pageId="46" pageNumber="47">Fig. 450</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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