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<document id="A863B5CDC2A47BD0629E83643D62FC4D" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFBDF77201CA11DB4E0ECB9E" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada uwinsensis Köhler 2010, n.sp." docType="treatment" docUuid="4CCC73A5-13FC-4FD9-A525-22DEFF33ADB3" docUuidSource="ZooBank" docVersion="3" lastPageNumber="263" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="261" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="0B9ADF1629055308C0A2981D5E530AD6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="38375C9DE3E3234B8DB445608F5F3352">
<mods:title id="85DB0BFB831698892735D5E4D7D45819">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="05169E7698188E6A4D076042996C49E5" type="personal">
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</mods:role>
<mods:namePart id="06EAF98C6CE515D7A7B34D12B154E34D">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="633446B97E92215502839B30DA6AD3AD">text</mods:typeOfResource>
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<mods:title id="BB8BA48923C3D79A3F3C0E8763CC6FB9">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="AEC9C578C18D00F8ED4A5747BC74A5E5">
<mods:date id="19C383B097DD85639762C747C3395699">2010</mods:date>
<mods:detail id="7FE59C0364383D19514571DAB0B26608" type="pubDate">
<mods:number id="D09F2A128F0DB86F8E2554C6970C0BAC">2010-11-24</mods:number>
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<mods:detail id="876E37D252E05F32A27E8B3226E6F2CB" type="volume">
<mods:number id="22F4072EBE75569F800F2E664ADA527E">62</mods:number>
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<mods:number id="9488A97CAEAE10C28D127FFCC89D43A2">2</mods:number>
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<mods:location id="526A1ED170836A1F92E2D69EDCBB78B1">
<mods:url id="7E4CD283A62E2AAE5D390B7617336EB7">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
</mods:location>
<mods:classification id="9330147843EB4201E6F138B82E2549C4">journal article</mods:classification>
<mods:identifier id="FEBC1A0782181E4F55C1692D18BE2B07" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="9A7CA6573AAF438197993F0DB45057FF" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="B851DF1552F61B2B52058D0AD82BC46C" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="FDFBBFEFE4E53D770B253355B85F138C" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FFBDF77201CA11DB4E0ECB9E" ID-DOI="http://doi.org/10.5281/zenodo.5239045" ID-Zenodo-Dep="5239045" LSID="urn:lsid:zoobank.org:act:4CCC73A5-13FC-4FD9-A525-22DEFF33ADB3" httpUri="http://treatment.plazi.org/id/03F987C5FFBDF77201CA11DB4E0ECB9E" lastPageId="46" lastPageNumber="263" pageId="44" pageNumber="261">
<subSubSection id="C34A6558FFBDF77001CA11DB4856CFCC" box="[959,1347,414,440]" pageId="44" pageNumber="261" type="nomenclature">
<paragraph id="8BEF36D3FFBDF77001CA11DB4856CFCC" blockId="44.[959,1347,414,440]" box="[959,1347,414,440]" pageId="44" pageNumber="261">
<heading id="D0A781BFFFBDF77001CA11DB4856CFCC" box="[959,1347,414,440]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="261" reason="2">
<emphasis id="B924EAC1FFBDF77001CA11DB4856CFCC" bold="true" box="[959,1347,414,440]" pageId="44" pageNumber="261">
<taxonomicName id="4C504D50FFBDF77001CA11DB49EACFCC" ID-CoL="D7S8" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[959,1279,414,440]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="261" phylum="Mollusca" rank="species" species="uwinsensis" status="sp. nov.">
<emphasis id="B924EAC1FFBDF77001CA11DB49EACFCC" bold="true" box="[959,1279,414,440]" italics="true" pageId="44" pageNumber="261">Amplirhagada uwinsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFBDF770077211DB4856CFCC" box="[1287,1347,414,440]" pageId="44" pageNumber="261" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFBDF7700132119F49A1CD45" pageId="44" pageNumber="261" type="materials_examined">
<paragraph id="8BEF36D3FFBDF7700132119F4802CC13" blockId="44.[839,1467,474,615]" pageId="44" pageNumber="261">
<materialsCitation id="3B383C8EFFBDF7700132119F4801CC13" collectingDate="2006-06-09" collectionCode="V" county="Kimberley" latitude="-15.258889" location="Hanover Bay" longLatPrecision="21" longitude="124.76889" municipality="Bonaparte Archipelago" pageId="44" pageNumber="261" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFBDF7700132119F4EF3CF86" bold="true" box="[839,998,474,498]" pageId="44" pageNumber="261">
<typeStatus id="54EB8871FFBDF7700132119F4E96CF86" box="[839,899,474,498]" pageId="44" pageNumber="261">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFBDF770018F119F4954CF85" box="[1018,1089,474,497]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="44" pageNumber="261">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFBDF7700628119F483BCF85" box="[1117,1326,474,497]" country="Australia" name="Western Australia" pageId="44" pageNumber="261">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFBDF7700748119F48A2CF85" box="[1341,1463,474,497]" pageId="44" pageNumber="261">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFBDF770013211BD494ECC7B" box="[839,1115,504,527]" pageId="44" pageNumber="261">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FFBDF770061F11BD4812CC7B" LSID="urn:lsid:plazi:treatment:03F987C5FFBDF77201CA11DB4E0ECB9E:8E8F6008FFBDF770061F11BD4812CC7B" box="[1130,1287,504,527]" county="Kimberley" latitude="-15.258889" longLatPrecision="21" longitude="124.76889" municipality="Bonaparte Archipelago" name="Hanover Bay" pageId="44" pageNumber="261" stateProvince="Western Australia">Hanover Bay</location>
,
<location id="8E8F6008FFBDF770076311BD48A3CC7B" LSID="urn:lsid:plazi:treatment:03F987C5FFBDF77201CA11DB4E0ECB9E:8E8F6008FFBDF770076311BD48A3CC7B" box="[1302,1462,504,527]" county="Kimberley" latitude="-15.258889" longLatPrecision="21" longitude="124.76889" municipality="Bonaparte Archipelago" name="Uwins Island" pageId="44" pageNumber="261" stateProvince="Western Australia">Uwins Island</location>
,
<geoCoordinate id="EE645014FFBDF770013212504ED5CC58" box="[839,960,533,556]" degrees="15" direction="south" minutes="15" orientation="latitude" pageId="44" pageNumber="261" precision="15" seconds="32" value="-15.258889">15°15'32&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFBDF77001B11250495BCC58" box="[964,1102,533,556]" degrees="124" direction="east" minutes="46" orientation="longitude" pageId="44" pageNumber="261" precision="15" seconds="08" value="124.76889">124°46'08&quot;E</geoCoordinate>
; KIS-3–106.
<location id="8E8F6008FFBDF77006AD1250481FCC58" LSID="urn:lsid:plazi:treatment:03F987C5FFBDF77201CA11DB4E0ECB9E:8E8F6008FFBDF77006AD1250481FCC58" box="[1240,1290,533,556]" county="Kimberley" latitude="-15.258889" longLatPrecision="21" longitude="124.76889" municipality="Bonaparte Archipelago" name="Vine" pageId="44" pageNumber="261" stateProvince="Western Australia">Vine</location>
thicket and large fig trees on upper slopes, piles of sandstone rocks, under large slabs. (leg.
<collectionCode id="ED41AE16FFBDF770018F12154918CC13" box="[1018,1037,592,615]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="44" pageNumber="261" type="Museum">V</collectionCode>
. Kessner,
<date id="FFEE1013FFBDF770060C12154819CC13" box="[1145,1292,592,615]" pageId="44" pageNumber="261" value="2006-06-09">
<collectingDate id="EFAAE9FBFFBDF770060C12154819CC13" box="[1145,1292,592,615]" pageId="44" pageNumber="261" value="2006-06-09">09 June 2006</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFBDF770013212CD4897CC90" blockId="44.[839,1467,648,740]" pageId="44" pageNumber="261">
<emphasis id="B924EAC1FFBDF770013212CD4EC7CCEF" bold="true" box="[839,978,648,667]" pageId="44" pageNumber="261">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFBDF770019712CC480BCCE8" accessionNumber="S34621" box="[994,1310,649,668]" collectionCode="WAM" pageId="44" pageNumber="261" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FFBDF770019712CC4922CCE8" box="[994,1079,649,668]" pageId="44" pageNumber="261" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFBDF770064A12CC4960CCE8" box="[1087,1141,649,668]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF770060B12CC49D6CCE8" box="[1150,1219,649,668]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34621" pageId="44" pageNumber="261" type="EnaNcbi">S34621</accessionNumber>
(Pl. 2.7)
</materialsCitation>
.
<materialsCitation id="3B383C8EFFBDF770075F12CC486BCC90" accessionNumber="S36991, S37441" collectionCode="WAM" latitude="-15.256945" longLatPrecision="21" longitude="124.80112" pageId="44" pageNumber="261" specimenCount="3" typeStatus="paratype">
<typeStatus id="54EB8871FFBDF770075F12CC4896CCE8" box="[1322,1411,649,668]" pageId="44" pageNumber="261" type="paratype">Paratypes</typeStatus>
AMS C463715 (15 preserved specimens, as
<typeStatus id="54EB8871FFBDF77006DB12E44814CCC0" box="[1198,1281,673,692]" pageId="44" pageNumber="261" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FFBDF770076712E44852CCC0" box="[1298,1351,673,692]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF770072512E44881CCC0" box="[1360,1428,673,692]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36991" pageId="44" pageNumber="261" type="EnaNcbi">S36991</accessionNumber>
(35 preserved specimens, as
<typeStatus id="54EB8871FFBDF770066F12FC497DCCB8" box="[1050,1128,697,716]" pageId="44" pageNumber="261" type="holotype">holotype</typeStatus>
), AMS C463716 (15 shells,
<geoCoordinate id="EE645014FFBDF770072F12FC48AFCCB8" box="[1370,1466,697,716]" degrees="15" direction="south" minutes="15" orientation="latitude" pageId="44" pageNumber="261" precision="15" seconds="25" value="-15.256945">15°15'25&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFBDF770013212944EA2CC90" box="[839,951,721,740]" degrees="124" direction="east" minutes="48" orientation="longitude" pageId="44" pageNumber="261" precision="15" seconds="04" value="124.80112">124°48'04&quot;E</geoCoordinate>
),
<collectionCode id="ED41AE16FFBDF77001B012944EECCC90" box="[965,1017,721,740]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF770018B12944955CC90" box="[1022,1088,721,740]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37441" pageId="44" pageNumber="261" type="EnaNcbi">S37441</accessionNumber>
(30 shells,
<geoCoordinate id="EE645014FFBDF77006D712944816CC90" box="[1186,1283,721,740]" degrees="15" direction="south" minutes="15" orientation="latitude" pageId="44" pageNumber="261" precision="15" seconds="25" value="-15.256945">15°15'25&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFBDF770077D1294486DCC90" box="[1288,1400,721,740]" degrees="124" direction="east" minutes="48" orientation="longitude" pageId="44" pageNumber="261" precision="15" seconds="04" value="124.80112">124°48'04&quot;E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFBDF7700132134349A1CD45" blockId="44.[839,1467,774,817]" pageId="44" pageNumber="261">
<emphasis id="B924EAC1FFBDF770013213434940CD6D" bold="true" box="[839,1109,774,793]" pageId="44" pageNumber="261">Additional, non-type material</emphasis>
.
<collectionCode id="ED41AE16FFBDF770062A13434986CD6D" box="[1119,1171,774,793]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF77006ED134349CFCD6D" box="[1176,1242,774,793]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36987" pageId="44" pageNumber="261" type="EnaNcbi">S36987</accessionNumber>
–90,
<collectionCode id="ED41AE16FFBDF770077613434822CD6D" box="[1283,1335,774,793]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF77007491343486ACD6D" box="[1340,1407,774,793]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37440" pageId="44" pageNumber="261" type="EnaNcbi">S37440</accessionNumber>
,
<collectionCode id="ED41AE16FFBDF77007F2134348AECD6D" box="[1415,1467,774,793]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="44" pageNumber="261" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFBDF7700132135B4E9CCD45" box="[839,905,798,817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37442" pageId="44" pageNumber="261" type="EnaNcbi">S37442</accessionNumber>
–4, AMS C463717 (Uwins Island).
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFBDF772013213104E0ECB9E" lastPageId="46" lastPageNumber="263" pageId="44" pageNumber="261" type="etymology">
<paragraph id="8BEF36D3FFBDF770013213104E81CDFD" blockId="44.[839,1466,853,905]" pageId="44" pageNumber="261">
<emphasis id="B924EAC1FFBDF770013213104ED4CD19" bold="true" box="[839,961,853,877]" pageId="44" pageNumber="261">Etymology</emphasis>
. In reference to Uwins Island, where this species occurs.
</paragraph>
<caption id="DF2F665BFFBDF77002D317CE4C6AC9A1" ID-DOI="http://doi.org/10.5281/zenodo.5239225" ID-Zenodo-Dep="5239225" httpUri="https://zenodo.org/record/5239225/files/figure.png" pageId="44" pageNumber="261" startId="44.[166,229,1931,1952]" targetBox="[172,779,1526,1911]" targetPageId="44">
<paragraph id="8BEF36D3FFBDF77002D317CE4C6AC9A1" blockId="44.[166,794,1931,2005]" pageId="44" pageNumber="261">
Figure 67. Genitalia of
<taxonomicName id="4C504D50FFBDF77003F917CE4F90C9D4" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[396,645,1931,1952]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="261" phylum="Mollusca" rank="species" species="uwinsensis" status="sp. nov.">
<emphasis id="B924EAC1FFBDF77003F917CE4F90C9D4" box="[396,645,1931,1952]" italics="true" pageId="44" pageNumber="261">Amplirhagada uwinsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFBDF77000F817CE4FA9C9D4" box="[653,700,1931,1952]" pageId="44" pageNumber="261" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463715 (7 August, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFBDF770013217CE49A5C9A1" ID-DOI="http://doi.org/10.5281/zenodo.5239229" ID-Zenodo-Dep="5239229" httpUri="https://zenodo.org/record/5239229/files/figure.png" pageId="44" pageNumber="261" startId="44.[839,902,1931,1952]" targetBox="[839,1454,931,1910]" targetPageId="44">
<paragraph id="8BEF36D3FFBDF770013217CE49A5C9A1" blockId="44.[839,1467,1931,2005]" pageId="44" pageNumber="261">
Figure 68. Interior of penial chamber of
<taxonomicName id="4C504D50FFBDF77006B617CE48AFC9D4" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1219,1466,1931,1952]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="261" phylum="Mollusca" rank="species" species="uwinsensis" status="sp. nov.">
<emphasis id="B924EAC1FFBDF77006B617CE48AFC9D4" box="[1219,1466,1931,1952]" italics="true" pageId="44" pageNumber="261">Amplirhagada uwinsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFBDF770013217E34E63C9CF" box="[839,886,1958,1979]" pageId="44" pageNumber="261" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463715 (7 August, scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFBCF771020C147A4932CAD0" ID-DOI="http://doi.org/10.5281/zenodo.5239231" ID-Zenodo-Dep="5239231" httpUri="https://zenodo.org/record/5239231/files/figure.png" pageId="45" pageNumber="262" startId="45.[121,184,1087,1108]" targetBox="[121,1421,151,1075]" targetPageId="45">
<paragraph id="8BEF36D3FFBCF771020C147A4932CAD0" blockId="45.[121,1422,1087,1188]" pageId="45" pageNumber="262">
Figure 69. SEM photographs of
<taxonomicName id="4C504D50FFBCF77103CD147A4FA7CA20" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[440,690,1087,1108]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="262" phylum="Mollusca" rank="species" species="uwinsensis" status="sp. nov.">
<emphasis id="B924EAC1FFBCF77103CD147A4FA7CA20" box="[440,690,1087,1108]" italics="true" pageId="45" pageNumber="262">Amplirhagada uwinsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFBCF77100CF147A4FFDCA20" box="[698,744,1087,1108]" pageId="45" pageNumber="262" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FFBCF7710082147A4E34CA20" box="[759,801,1087,1108]" italics="true" pageId="45" pageNumber="262">A–C</emphasis>
) Shell, paratype AMS C463716: (
<emphasis id="B924EAC1FFBCF7710606147A4997CA20" box="[1139,1154,1087,1108]" italics="true" pageId="45" pageNumber="262">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFBCF771032D141F4C72CA1B" box="[344,359,1114,1135]" italics="true" pageId="45" pageNumber="262">B</emphasis>
) sculpture penultimate whorl viewed rectangularly from above (scale 100 µm); (
<emphasis id="B924EAC1FFBCF7710611141F4961CA1B" box="[1124,1140,1114,1135]" italics="true" pageId="45" pageNumber="262">C</emphasis>
) close-up of periphery of last whorl (scale 200 µm). (
<emphasis id="B924EAC1FFBCF771032C14314C7FCAFD" box="[345,362,1140,1161]" italics="true" pageId="45" pageNumber="262">D</emphasis>
) Jaw, paratype AMS C463715 (scale 100 µm). (
<emphasis id="B924EAC1FFBCF771014C14314E76CAFD" box="[825,867,1140,1161]" italics="true" pageId="45" pageNumber="262">E–F</emphasis>
) Radula, paratype AMS C463715: (
<emphasis id="B924EAC1FFBCF77106CB143149D8CAFD" box="[1214,1229,1140,1161]" italics="true" pageId="45" pageNumber="262">E</emphasis>
) close-up of central and inner lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FFBCF771039214CA4CE3CAD0" box="[487,502,1167,1188]" italics="true" pageId="45" pageNumber="262">F</emphasis>
) close-up of inner to middle marginal teeth (scale 10 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFBCF771020C14884FF8C8CE" blockId="45.[121,749,1229,1722]" pageId="45" pageNumber="262">
<emphasis id="B924EAC1FFBCF771020C14884DA4CA91" bold="true" box="[121,177,1229,1253]" pageId="45" pageNumber="262">Shell</emphasis>
(
<figureCitation id="136B2A56FFBCF77102B5148B4C56CA91" box="[192,323,1230,1253]" captionStart="Figure 69" captionStartId="45.[121,184,1087,1108]" captionTargetBox="[121,1421,151,1075]" captionTargetId="figure-422@45.[121,1421,151,1075]" captionTargetPageId="45" captionText="Figure 69. SEM photographs of Amplirhagada uwinsensis n.sp. (A–C) Shell, paratype AMS C463716: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture penultimate whorl viewed rectangularly from above (scale 100 µm); (C) close-up of periphery of last whorl (scale 200 µm). (D) Jaw, paratype AMS C463715 (scale 100 µm). (E–F) Radula, paratype AMS C463715: (E) close-up of central and inner lateral teeth (scale 10 µm); (F) close-up of inner to middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239231" httpUri="https://zenodo.org/record/5239231/files/figure.png" pageId="45" pageNumber="262">Fig. 69A–C</figureCitation>
, Pl. 2.7–9). Semi-globose with medium high spire; solid. Periphery angulate; upper and basal sectors of whorls rounded. Umbilicus completely concealed by columellar reflection. Background colour light reddish brown; sub-sutural band diffuse to well defined, light brown to chestnut brown, varies greatly in thickness and contrast; mid-whorl band diffuse, brownish to chestnut brown, thin, varies greatly in contrast and thickness, visible on last whorl(s) only; bands may blend into each other across whorl surface; ventral colour yellowish-greenish brown to ochre; outer lip colour blending into chestnut brown; inner lip blending into dark ochre to chestnut brown. Protoconch c.
<quantity id="4CA89B36FFBCF77100B916544DB7C831" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="45" pageNumber="262" unit="mm" value="2.7">2.7 mm</quantity>
in diameter, comprising 1.7 whorls, with fine, indistinct axial lirae. Teleoconch with marked growth lines, becoming oblique towards base of shell. Angle of aperture 45°; outer lip rounded, slightly expanded, not reflected, basal node of lip weak, palatal node absent. Parietal wall of inner lip absent.
</paragraph>
<paragraph id="8BEF36D3FFBCF771020C169E4F00C93E" blockId="45.[121,749,1755,1866]" pageId="45" pageNumber="262">
<emphasis id="B924EAC1FFBCF771020C169E4C43C887" bold="true" box="[121,342,1755,1779]" pageId="45" pageNumber="262">Pallial morphology</emphasis>
. Pallial cavity short, extending ½ to
<emphasis id="B924EAC1FFBCF771020C16BD4D94C972" bold="true" box="[121,129,1784,1798]" pageId="45" pageNumber="262">
<superScript id="7C259B9BFFBCF771020C16BD4D94C972" attach="none" box="[121,129,1784,1798]" fontSize="6" pageId="45" pageNumber="262">3</superScript>
</emphasis>
⁄
<emphasis id="B924EAC1FFBCF77102F316BA4D9BC979" bold="true" box="[134,142,1791,1805]" pageId="45" pageNumber="262">4</emphasis>
whorls; mantle pigmentation consists of indistinctive, sparsely distributed brown spots. Kidney extending about half or more than half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFBCF771020C172E486FC964" blockId="45.[121,749,1899,1981]" lastBlockId="45.[794,1422,1227,1808]" pageId="45" pageNumber="262">
<emphasis id="B924EAC1FFBCF771020C172E4C74C9F7" bold="true" box="[121,353,1899,1923]" pageId="45" pageNumber="262">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFBCF7710304172E4CE2C9F6" box="[369,503,1899,1922]" captionStart-0="Figure 67" captionStart-1="Figure 68" captionStartId-0="44.[166,229,1931,1952]" captionStartId-1="44.[839,902,1931,1952]" captionTargetBox-0="[172,779,1526,1911]" captionTargetBox-1="[839,1454,931,1910]" captionTargetId-0="figure-81@44.[166,793,1518,1937]" captionTargetId-1="figure-105@44.[839,1466,927,1920]" captionTargetPageId-0="44" captionTargetPageId-1="44" captionText-0="Figure 67. Genitalia of Amplirhagada uwinsensis n.sp., paratype AMS C463715 (7 August, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 68. Interior of penial chamber of Amplirhagada uwinsensis n.sp., paratype AMS C463715 (7 August, scale 5 mm). Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239225" figureDoi-1="http://doi.org/10.5281/zenodo.5239229" httpUri-0="https://zenodo.org/record/5239225/files/figure.png" httpUri-1="https://zenodo.org/record/5239229/files/figure.png" pageId="45" pageNumber="262">Figs. 67–68</figureCitation>
). Penis rather straight, more or less of same length as anterior part of oviduct. Vas deferens coils once before entering penis. Penial retractor muscle very short to stubby. Penial verge very short (barely visible), slender to spatulate, with pointed tip. Penial wall pustules comparatively large, elongated, arranged in rows on median and basal portion of penial chamber, forming densely packed, narrow longitudinal pilasters at apical end of penial chamber. Main stimulatory pilaster welldifferentiated, large, cone-shaped, covered with flattened pustules, comprising anterior to median portion of penial chamber. Two additional pilasters are formed that are covered by flattened pustules or smooth narrow ridges. Vas deferens entering penial sheath in upper third. Vagina moderately wide, tubular to posteriorly inflated; inner vaginal wall with weakly developed longitudinal pilasters. Spermathecal duct wide, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising less than half to about half of anterior part of oviduct, more or less straight. Spermoviduct of same length as anterior part of oviduct or longer. Talon embedded in albumen gland close to anterior end of albumen gland.
</paragraph>
<paragraph id="8BEF36D3FFBCF772016F17764F82CB4E" blockId="45.[794,1422,1843,1984]" lastBlockId="46.[166,794,1227,1338]" lastPageId="46" lastPageNumber="263" pageId="45" pageNumber="262">
<emphasis id="B924EAC1FFBCF771016F17764905C93F" bold="true" box="[794,1040,1843,1867]" pageId="45" pageNumber="262">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFBCF7710656177649B3C93E" box="[1059,1190,1843,1866]" captionStart="Figure 69" captionStartId="45.[121,184,1087,1108]" captionTargetBox="[121,1421,151,1075]" captionTargetId="figure-422@45.[121,1421,151,1075]" captionTargetPageId="45" captionText="Figure 69. SEM photographs of Amplirhagada uwinsensis n.sp. (A–C) Shell, paratype AMS C463716: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture penultimate whorl viewed rectangularly from above (scale 100 µm); (C) close-up of periphery of last whorl (scale 200 µm). (D) Jaw, paratype AMS C463715 (scale 100 µm). (E–F) Radula, paratype AMS C463715: (E) close-up of central and inner lateral teeth (scale 10 µm); (F) close-up of inner to middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239231" httpUri="https://zenodo.org/record/5239231/files/figure.png" pageId="45" pageNumber="262">Fig. 69E–F</figureCitation>
). Rectangular. Tooth formula C+14–16+4+22. In average 149±16.0 rows of teeth, 29.5±1.0 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of teeth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of teeth; ectocones and endocones vestigial. Marginal teeth with elongate mesocones; ectocones smaller and narrower than mesocones; endocones shorter than ectocones.
</paragraph>
<caption id="DF2F665BFFBFF77202D314654C25CAF1" ID-DOI="http://doi.org/10.5281/zenodo.5239233" ID-Zenodo-Dep="5239233" httpUri="https://zenodo.org/record/5239233/files/figure.png" pageId="46" pageNumber="263" startId="46.[166,229,1056,1077]" targetBox="[167,1467,151,1043]" targetPageId="46">
<paragraph id="8BEF36D3FFBFF77202D314654C25CAF1" blockId="46.[166,1467,1056,1157]" pageId="46" pageNumber="263">
Figure 70. SEM photographs of
<taxonomicName id="4C504D50FFBFF772039714644FFCCA42" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[482,745,1057,1078]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="263" phylum="Mollusca" rank="species" species="sphaeroidea" status="sp. nov.">
<emphasis id="B924EAC1FFBFF772039714644FFCCA42" box="[482,745,1057,1078]" italics="true" pageId="46" pageNumber="263">Amplirhagada sphaeroidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFBFF772008514654E0BCA41" box="[752,798,1056,1077]" pageId="46" pageNumber="263" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FFBFF772015914644E40CA42" box="[812,853,1057,1078]" italics="true" pageId="46" pageNumber="263">A–B</emphasis>
) Shell, paratype AMS C463719: (
<emphasis id="B924EAC1FFBFF77206D6146449A7CA42" box="[1187,1202,1057,1078]" italics="true" pageId="46" pageNumber="263">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFBFF77203FA147E4C8BCA24" box="[399,414,1083,1104]" italics="true" pageId="46" pageNumber="263">B</emphasis>
) close-up of periphery of last whorl (scale 1 mm); (
<emphasis id="B924EAC1FFBFF77201D6147E4EA6CA24" box="[931,947,1083,1104]" italics="true" pageId="46" pageNumber="263">C</emphasis>
) jaw, paratype WAM
<accessionNumber id="9403AB30FFBFF77206F8147E49C2CA24" box="[1165,1239,1083,1104]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37027" pageId="46" pageNumber="263" type="EnaNcbi">S37027</accessionNumber>
(scale 100 µm). (
<emphasis id="B924EAC1FFBFF77207F2147E48A7CA24" box="[1415,1458,1083,1104]" italics="true" pageId="46" pageNumber="263">D–E</emphasis>
) Radula, paratype WAM
<accessionNumber id="9403AB30FFBFF77203FA14134CC9CA1F" box="[399,476,1110,1131]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37027" pageId="46" pageNumber="263" type="EnaNcbi">S37027</accessionNumber>
: (
<emphasis id="B924EAC1FFBFF772039814134CEBCA1F" box="[493,510,1110,1131]" italics="true" pageId="46" pageNumber="263">D</emphasis>
) close-up of inner lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FFBFF77201B114134EC6CA1F" box="[964,979,1110,1131]" italics="true" pageId="46" pageNumber="263">E</emphasis>
) Close-up of outer lateral and inner marginal teeth (scale 20 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFBFF77202D315184E0ECB9E" blockId="46.[166,795,1373,1514]" pageId="46" pageNumber="263">
<emphasis id="B924EAC1FFBFF77202D315184CB7CB01" bold="true" box="[166,418,1373,1397]" pageId="46" pageNumber="263">Comparative remarks</emphasis>
. One of the few species with large, dome-shaped shells. Differs from
<taxonomicName id="4C504D50FFBFF7720078153E4F80CBE6" authorityName="Köhler" authorityYear="2010" box="[525,661,1403,1426]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="263" phylum="Mollusca" rank="species" species="tricenaria">
<emphasis id="B924EAC1FFBFF7720078153E4F80CBE6" box="[525,661,1403,1426]" italics="true" pageId="46" pageNumber="263">A. tricenaria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FFBFF77200B2153E4E0CCBE6" authorityName="Köhler" authorityYear="2010" box="[711,793,1403,1426]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="263" phylum="Mollusca" rank="species" species="regia">
<emphasis id="B924EAC1FFBFF77200B2153E4E0CCBE6" box="[711,793,1403,1426]" italics="true" pageId="46" pageNumber="263">A. regia</emphasis>
</taxonomicName>
by conspicuously dark brown outer lip colour and by huge, cone-shaped main pilaster, presence of additional pilasters and characteristically elongated pustules of inner penial wall.
</paragraph>
</subSubSection>
</treatment>
</document>
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