treatments-xml/data/03/FF/4B/03FF4B3BFF8FFFEDFF02F9AAFB8E89A6.xml

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<document id="227393BFC098478EA865D7720F28B6D9" ID-CLB-Dataset="24345" ID-DOI="10.11646/zootaxa.4722.3.4" ID-GBIF-Dataset="03d67507-fe7a-44e3-bf0f-3b842d7d2e72" ID-ISSN="1175-5326" ID-Zenodo-Dep="3607103" ID-ZooBank="B7AEB4DA-FB6B-4B7A-8539-075E39AB7050" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578986109188" checkinUser="plazi" docAuthor="Saxton, Natalie A., Powell, Gareth S., Martin, Gavin J. &amp; Bybee, Seth M." docDate="2020" docId="03FF4B3BFF8FFFEDFF02F9AAFB8E89A6" docLanguage="en" docName="zootaxa.4722.3.4.pdf" docOrigin="Zootaxa 4722 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atyphella maritimus Saxton &amp; Powell 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="272" masterDocId="FFC63343FF8EFFEFFF95FF9EFFD48E79" masterDocTitle="Two new species of coastal Atyphella Olliff (Lampyridae: Luciolinae)" masterLastPageNumber="276" masterPageNumber="270" pageNumber="271" updateTime="1698777403341" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="769B6DEA4BBAC90108C4E7E56027771D">Two new species of coastal Atyphella Olliff (Lampyridae: Luciolinae)</mods:title>
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<mods:namePart id="EE94319240CDE423B186656CC8EFFD34">Powell, Gareth S.</mods:namePart>
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<taxonomicName id="4C5681AEFF8FFFEEFF02F9AAFDBC8837" authority="Saxton &amp; Powell" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[151,616,1588,1615]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="271" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B922263FFF8FFFEEFF02F9AAFE5A8837" bold="true" box="[151,398,1588,1615]" italics="true" pageId="1" pageNumber="271">Atyphella maritimus</emphasis>
Saxton &amp; Powell
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(
<figureCitation id="136DE6A8FF8FFFEEFF0AF9C6FE90880A" box="[159,324,1624,1651]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1931]" captionTargetBox="[151,1436,410,1886]" captionTargetId="figure@3.[151,1436,410,1886]" captionTargetPageId="3" captionText="FIGURE 1. AI) Atyphella spp. holotypes. AC) dorsal pronotum: A) A. aphrogeneia, B) A. maritimus, C) A. marigenous; DF) scutellum: D) A. aphrogeneia, E) A. maritimus, F) A. marigenous; GI) left elytron, notice extent of costae G) A. aphrogeneia, H) A. maritimus, I) A. marigenous. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="1" pageNumber="271">Figs. 1B, E, H</figureCitation>
;
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)
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<emphasis id="B922263FFF8FFFEEFF02F93EFE9188C3" bold="true" box="[151,325,1695,1722]" pageId="1" pageNumber="271">Type material.</emphasis>
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. male
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(
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female
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(
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): //
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&amp;
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<collectorName id="26A39FFBFF8FFFEEFBD0F892FB62895F" box="[1093,1206,1804,1830]" pageId="1" pageNumber="271">G. Powell</collectorName>
// (5
<collectionCode id="ED4762E8FF8FFFEEFB64F892FAEC895F" box="[1265,1336,1804,1830]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="1" pageNumber="271">ANIC</collectionCode>
, 9
<collectionCode id="ED4762E8FF8FFFEEFACFF893FA4C895E" box="[1370,1432,1805,1831]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="1" pageNumber="271" type="University or college">BYU</collectionCode>
, 5
<collectionCode id="ED4762E8FF8FFFEEFF38F8AFFF258932" box="[173,241,1841,1867]" pageId="1" pageNumber="271">NHM</collectionCode>
, 5
<collectionCode id="ED4762E8FF8FFFEEFE87F8AEFEB18933" box="[274,357,1840,1866]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="271" type="Museum">USNM</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8FFFEEFEE6F8AFFEF48916" ID-GBIF-Occurrence="2549146466" collectingDate="2018-05-24" collectionCode="BYU" collectorName="Malekula Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-16.029" location="Tenmaru" longLatPrecision="7" longitude="167.1728" pageId="1" pageNumber="271" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8FFFEEFEE6F8AFFE078932" box="[371,467,1841,1867]" name="Vanuatu" pageId="1" pageNumber="271">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8FFFEEFE4BF8AEFDBE8932" box="[478,618,1840,1867]" pageId="1" pageNumber="271">Malekula Is.</collectorName>
/
<location id="8E89ACF6FF8FFFEEFD15F8AFFD328932" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8FFFEDFF02F9AAFB8E89A6:8E89ACF6FF8FFFEEFD15F8AFFD328932" box="[640,742,1841,1867]" country="Vanuatu" latitude="-16.029" longLatPrecision="7" longitude="167.1728" name="Tenmaru" pageId="1" pageNumber="271">Tenmaru</location>
,
<geoCoordinate id="EE629CEAFF8FFFEEFD67F8AEFC828933" box="[754,854,1840,1866]" degrees="16.0290" direction="south" orientation="latitude" pageId="1" pageNumber="271" precision="5" value="-16.029">-16.0290</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8FFFEEFCF8F8AEFC0D8933" box="[877,985,1840,1866]" degrees="167.1728" direction="east" orientation="longitude" pageId="1" pageNumber="271" precision="5" value="167.1728">167.1728</geoCoordinate>
,
<date id="FFE8DCEDFF8FFFEEFC71F8AEFB878933" box="[996,1107,1840,1866]" pageId="1" pageNumber="271" value="2018-05-24">
<collectingDate id="EFAC2505FF8FFFEEFC71F8AEFB878933" box="[996,1107,1840,1866]" pageId="1" pageNumber="271" value="2018-05-24">24.v.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8FFFEEFB37F8AEFADC8933" box="[1186,1288,1840,1866]" pageId="1" pageNumber="271">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8FFFEEFAB9F8AEFA488933" box="[1324,1436,1840,1866]" pageId="1" pageNumber="271">G. Powell</collectorName>
// (20
<collectionCode id="ED4762E8FF8FFFEEFF49F8CBFECD8916" box="[220,281,1877,1903]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="1" pageNumber="271" type="University or college">BYU</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8FFFEEFEB2F8CBFA498916" ID-GBIF-Occurrence="2549146470" box="[295,1437,1875,1903]" collectingDate="2018-06-04" collectionCode="BYU" collectorName="Santo Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-15.6296" location="Ipayato" longLatPrecision="7" longitude="166.8541" pageId="1" pageNumber="271" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8FFFEEFEB2F8CBFE538916" box="[295,391,1877,1903]" name="Vanuatu" pageId="1" pageNumber="271">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8FFFEEFE06F8CAFE228916" box="[403,502,1876,1903]" pageId="1" pageNumber="271">Santo Is.</collectorName>
/
<location id="8E89ACF6FF8FFFEEFD98F8CBFDB08916" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8FFFEDFF02F9AAFB8E89A6:8E89ACF6FF8FFFEEFD98F8CBFDB08916" box="[525,612,1877,1903]" country="Vanuatu" latitude="-15.6296" longLatPrecision="7" longitude="166.8541" name="Ipayato" pageId="1" pageNumber="271">Ipayato</location>
,
<geoCoordinate id="EE629CEAFF8FFFEEFDE5F8CDFD008917" box="[624,724,1875,1902]" degrees="15.6296" direction="south" orientation="latitude" pageId="1" pageNumber="271" precision="5" value="-15.6296">-15.6296</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8FFFEEFD7EF8CDFC838917" box="[747,855,1875,1902]" degrees="166.8541" direction="east" orientation="longitude" pageId="1" pageNumber="271" precision="5" value="166.8541">166.8541</geoCoordinate>
,
<date id="FFE8DCEDFF8FFFEEFCF6F8CAFC198917" box="[867,973,1876,1902]" pageId="1" pageNumber="271" value="2018-06-04">
<collectingDate id="EFAC2505FF8FFFEEFCF6F8CAFC198917" box="[867,973,1876,1902]" pageId="1" pageNumber="271" value="2018-06-04">4.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8FFFEEFB88F8CAFB508917" box="[1053,1156,1876,1902]" pageId="1" pageNumber="271">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8FFFEEFB3CF8CAFACE8917" box="[1193,1306,1876,1902]" pageId="1" pageNumber="271">G. Powell</collectorName>
// (3
<collectionCode id="ED4762E8FF8FFFEEFACDF8CBFA418916" box="[1368,1429,1877,1903]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="1" pageNumber="271" type="University or college">BYU</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8FFFEEFF02F8E7FB3989EA" ID-GBIF-Occurrence="2549146471" box="[151,1261,1911,1939]" collectingDate="2018-06-01" collectionCode="BYU" collectorName="Santo Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-15.1598" location="Lowerie" longLatPrecision="7" longitude="166.9511" pageId="1" pageNumber="271" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8FFFEEFF02F8E7FF2389EA" box="[151,247,1913,1939]" name="Vanuatu" pageId="1" pageNumber="271">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8FFFEEFE94F8E6FEB689EA" box="[257,354,1912,1939]" pageId="1" pageNumber="271">Santo Is.</collectorName>
/
<location id="8E89ACF6FF8FFFEEFEE2F8E6FE0389EB" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8FFFEDFF02F9AAFB8E89A6:8E89ACF6FF8FFFEEFEE2F8E6FE0389EB" box="[375,471,1912,1938]" country="Vanuatu" latitude="-15.1598" longLatPrecision="7" longitude="166.9511" name="Lowerie" pageId="1" pageNumber="271">Lowerie</location>
,
<geoCoordinate id="EE629CEAFF8FFFEEFE77F8E9FD9189EB" box="[482,581,1911,1938]" degrees="15.1598" direction="south" orientation="latitude" pageId="1" pageNumber="271" precision="5" value="-15.1598">-15.1598</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8FFFEEFDC1F8E9FD6889EB" box="[596,700,1911,1938]" degrees="166.9511" direction="east" orientation="longitude" pageId="1" pageNumber="271" precision="5" value="166.9511">166.9511</geoCoordinate>
<date id="FFE8DCEDFF8FFFEEFD57F8E6FCF989EB" box="[706,813,1912,1938]" pageId="1" pageNumber="271" value="2018-06-01">
<collectingDate id="EFAC2505FF8FFFEEFD57F8E6FCF989EB" box="[706,813,1912,1938]" pageId="1" pageNumber="271" value="2018-06-01">1.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8FFFEEFCEDF8E6FC0989EB" box="[888,989,1912,1938]" pageId="1" pageNumber="271">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8FFFEEFC6AF8E6FBBB89EB" box="[1023,1135,1912,1938]" pageId="1" pageNumber="271">G. Powell</collectorName>
// (4
<collectionCode id="ED4762E8FF8FFFEEFB3DF8E7FB3189EA" box="[1192,1253,1913,1939]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="1" pageNumber="271" type="University or college">BYU</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8FFFEEFB66F8E7FC3089CF" ID-GBIF-Occurrence="2549146468" collectingDate="2018-06-05" collectorName="Santo Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-15.1598" location="Lowerie" longLatPrecision="7" longitude="166.9511" pageId="1" pageNumber="271" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8FFFEEFB66F8E7FA8789EA" box="[1267,1363,1913,1939]" name="Vanuatu" pageId="1" pageNumber="271">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8FFFEEFAC9F8E6FF6689CE" pageId="1" pageNumber="271">Santo Is.</collectorName>
/
<location id="8E89ACF6FF8FFFEEFF5DF802FEFC89CF" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8FFFEDFF02F9AAFB8E89A6:8E89ACF6FF8FFFEEFF5DF802FEFC89CF" box="[200,296,1948,1974]" country="Vanuatu" latitude="-15.1598" longLatPrecision="7" longitude="166.9511" name="Lowerie" pageId="1" pageNumber="271">Lowerie</location>
,
<geoCoordinate id="EE629CEAFF8FFFEEFEA6F805FE4C89CF" box="[307,408,1947,1974]" degrees="15.1598" direction="south" orientation="latitude" pageId="1" pageNumber="271" precision="5" value="-15.1598">-15.1598</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8FFFEEFE3BF805FDC289CF" box="[430,534,1947,1974]" degrees="166.9511" direction="east" orientation="longitude" pageId="1" pageNumber="271" precision="5" value="166.9511">166.9511</geoCoordinate>
<date id="FFE8DCEDFF8FFFEEFD88F805FD5289CF" box="[541,646,1947,1974]" pageId="1" pageNumber="271" value="2018-06-05">
<collectingDate id="EFAC2505FF8FFFEEFD88F805FD5289CF" box="[541,646,1947,1974]" pageId="1" pageNumber="271" value="2018-06-05">5.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8FFFEEFD41F802FCEE89CF" box="[724,826,1948,1974]" pageId="1" pageNumber="271">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8FFFEEFCC8F802FC1989CF" box="[861,973,1948,1974]" pageId="1" pageNumber="271">G. Powell</collectorName>
//
</materialsCitation>
</paragraph>
<paragraph id="8BE9FA2DFF8FFFEDFF52F821FE288EAE" blockId="1.[151,1437,1695,2011]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="272" pageId="1" pageNumber="271">
<emphasis id="B922263FFF8FFFEEFF52F821FDD089A3" bold="true" box="[199,516,1983,2010]" pageId="1" pageNumber="271">Other material examined.</emphasis>
<materialsCitation id="3B3EF070FF8FFFEDFD9AF85FFE2C8EAE" ID-GBIF-Occurrence="2549146472" collectingDate="1929-06" collectingDateMax="1930-03" collectingDateMin="1929-06" collectionCode="NHM, BPBM" collectorName="L. Cheesman &amp; Malekula" country="Vanuatu" lastPageId="2" lastPageNumber="272" location="Malekula Atchin Island" municipality="Malekula Atchin Island" pageId="1" pageNumber="271" specimenCount="5" specimenCount-male="5">
New Hebrides (=
<collectingCountry id="F341BABDFF8FFFEEFD43F85EFCEE89A3" box="[726,826,1984,2010]" name="Vanuatu" pageId="1" pageNumber="271">Vanuatu</collectingCountry>
):
<location id="8E89ACF6FF8FFFEEFCC5F85EFBB689A3" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8FFFEDFF02F9AAFB8E89A6:8E89ACF6FF8FFFEEFCC5F85EFBB689A3" box="[848,1122,1984,2010]" country="Vanuatu" municipality="Malekula Atchin Island" name="Malekula Atchin Island" pageId="1" pageNumber="271">Malekula Atchin Island</location>
,
<date id="FFE8DCEDFF8FFFEEFBE7F85EFB0189A3" box="[1138,1237,1984,2010]" pageId="1" pageNumber="271" value="1929-06">
<collectingDate id="EFAC2505FF8FFFEEFBE7F85EFB0189A3" box="[1138,1237,1984,2010]" pageId="1" pageNumber="271" value="1929-06">vi. 1929</collectingDate>
</date>
,
<collectorName id="26A39FFBFF8FFFEEFB71F85FFAAB89A3" box="[1252,1407,1984,2011]" pageId="1" pageNumber="271">L. Cheesman</collectorName>
,
<specimenCount id="9D5031A4FF8FFFEDFA1BF85EFF088ECA" lastPageId="2" lastPageNumber="272" pageId="1" pageNumber="271" type="male">3 males</specimenCount>
;
<collectingMunicipality id="6B8D6057FF8CFFEDFF7FFF07FE2C8ECA" box="[234,504,153,179]" pageId="2" pageNumber="272">Malekula Atchin Island</collectingMunicipality>
,
<date id="FFE8DCEDFF8CFFEDFD92FF07FDB48ECA" box="[519,608,153,179]" pageId="2" pageNumber="272" value="1930-03">
<collectingDate id="EFAC2505FF8CFFEDFD92FF07FDB48ECA" box="[519,608,153,179]" pageId="2" pageNumber="272" value="1930-03">iii.1930</collectingDate>
</date>
,
<collectorName id="26A39FFBFF8CFFEDFDF8FF07FCD38ECA" box="[621,775,153,179]" pageId="2" pageNumber="272">L. Cheesman</collectorName>
,
<specimenCount id="9D5031A4FF8CFFEDFC81FF07FCB68ECA" box="[788,866,153,179]" pageId="2" pageNumber="272" type="male">1 male</specimenCount>
(
<collectionCode id="ED4762E8FF8CFFEDFCE1FF07FC6F8ECA" box="[884,955,153,179]" pageId="2" pageNumber="272">NHM</collectionCode>
);
<collectorName id="26A39FFBFF8CFFEDFC44FF07FBEA8ECA" box="[977,1086,153,179]" pageId="2" pageNumber="272">Malekula</collectorName>
,
<specimenCount id="9D5031A4FF8CFFEDFBD9FF07FB4D8ECA" box="[1100,1177,153,179]" pageId="2" pageNumber="272" type="male">1 male</specimenCount>
(
<collectionCode id="ED4762E8FF8CFFEDFB3EFF07FB298ECA" box="[1195,1277,153,179]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="272" type="Museum">BPBM</collectionCode>
, per image in
<bibRefCitation id="EFC787DCFF8CFFEDFF02FF23FE248EAE" author="Ballantyne, L. A. &amp; Lambkin, C." box="[151,496,189,215]" pageId="2" pageNumber="272" pagination="1 - 188" refId="ref4464" refString="Ballantyne, L. A. &amp; Lambkin, C. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997 (1), 1 - 188. https: // doi. org / 10.11646 / zootaxa. 1997.1.1" type="journal article" year="2009">Ballantyne and Lambkin 2009</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8CFFEDFF52FF7EFDD78FAA" pageId="2" pageNumber="272" type="diagnosis">
<paragraph id="8BE9FA2DFF8CFFEDFF52FF7EFDD78FAA" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52FF7EFE968E82" bold="true" box="[199,322,224,251]" pageId="2" pageNumber="272">Diagnosis.</emphasis>
<taxonomicName id="4C5681AEFF8CFFEDFEDDFF7FFDF98E82" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[328,557,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8CFFEDFEDDFF7FFDF98E82" box="[328,557,225,251]" italics="true" pageId="2" pageNumber="272">Atyphella maritimus</emphasis>
</taxonomicName>
is distinguished from all other South Pacific
<emphasis id="B922263FFF8CFFEDFBB2FF7FFB4D8E82" box="[1063,1177,225,251]" italics="true" pageId="2" pageNumber="272">
<taxonomicName id="4C5681AEFF8CFFEDFBB2FF7FFB418E82" box="[1063,1173,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="genus">Atyphella</taxonomicName>
,
</emphasis>
except
<taxonomicName id="4C5681AEFF8CFFEDFB64FF7FFA498E82" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[1265,1437,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8CFFEDFB64FF7FFA498E82" box="[1265,1437,225,251]" italics="true" pageId="2" pageNumber="272">A. aphrogeneia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8CFFEDFF5CFE9BFEBB8F66" box="[201,367,261,287]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8CFFEDFF5CFE9BFEBB8F66" box="[201,367,261,287]" italics="true" pageId="2" pageNumber="272">A. marigenous</emphasis>
</taxonomicName>
by the thickened elytral apices in conjunction with a dark median marking on the pronotum (see
<bibRefCitation id="EFC787DCFF8CFFEDFF59FEB7FDF18F3A" author="Ballantyne, L. A. &amp; Lambkin, C." box="[204,549,297,323]" pageId="2" pageNumber="272" pagination="1 - 188" refId="ref4464" refString="Ballantyne, L. A. &amp; Lambkin, C. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997 (1), 1 - 188. https: // doi. org / 10.11646 / zootaxa. 1997.1.1" type="journal article" year="2009">Ballantyne and Lambkin 2009</bibRefCitation>
).
<taxonomicName id="4C5681AEFF8CFFEDFDADFEB7FCC98F3A" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[568,797,297,323]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8CFFEDFDADFEB7FCC98F3A" box="[568,797,297,323]" italics="true" pageId="2" pageNumber="272">Atyphella maritimus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C5681AEFF8CFFEDFC0EFEB7FB938F3A" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[923,1095,297,323]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8CFFEDFC0EFEB7FB938F3A" box="[923,1095,297,323]" italics="true" pageId="2" pageNumber="272">A. aphrogeneia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8CFFEDFBEBFEB7FAF58F3A" box="[1150,1313,297,323]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8CFFEDFBEBFEB7FAF58F3A" box="[1150,1313,297,323]" italics="true" pageId="2" pageNumber="272">A. marigenous</emphasis>
</taxonomicName>
but differs in the following ways; excavation on frons shallow (frons deeply excavated in
<taxonomicName id="4C5681AEFF8CFFEDFB80FED3FB6D8F1E" box="[1045,1209,333,359]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8CFFEDFB80FED3FB6D8F1E" box="[1045,1209,333,359]" italics="true" pageId="2" pageNumber="272">A. marigenous</emphasis>
</taxonomicName>
); pronotum broadly rounded (pronotum with weakly obtuse, anterior angles in
<taxonomicName id="4C5681AEFF8CFFEDFCA7FEEFFC0B8FF2" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[818,991,369,395]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8CFFEDFCA7FEEFFC0B8FF2" box="[818,991,369,395]" italics="true" pageId="2" pageNumber="272">A. aphrogeneia</emphasis>
</taxonomicName>
); base of scutellum with dense punctation (punctation larger, but less dense in
<taxonomicName id="4C5681AEFF8CFFEDFDCCFE0BFD2F8FD6" box="[601,763,405,431]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8CFFEDFDCCFE0BFD2F8FD6" box="[601,763,405,431]" italics="true" pageId="2" pageNumber="272">A. marigenous</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8CFFEDFCA4FE0BFC088FD6" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[817,988,405,431]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8CFFEDFCA4FE0BFC088FD6" box="[817,988,405,431]" italics="true" pageId="2" pageNumber="272">A. aphrogeneia</emphasis>
</taxonomicName>
); weak elytral costae (more pronounced elytral costae in
<taxonomicName id="4C5681AEFF8CFFEDFEDAFE27FE278FAA" box="[335,499,441,467]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8CFFEDFEDAFE27FE278FAA" box="[335,499,441,467]" italics="true" pageId="2" pageNumber="272">A. marigenous</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8CFFEDFF52FE42FD8D8872" pageId="2" pageNumber="272" type="description">
<paragraph id="8BE9FA2DFF8CFFEDFF52FE42FD938B4A" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52FE42FE028F8E" bold="true" box="[199,470,476,503]" pageId="2" pageNumber="272">
Description.
<typeStatus id="54ED448FFF8CFFEDFEF7FE43FE058F8E" box="[354,465,477,503]" pageId="2" pageNumber="272" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Overall, body parallel-sided, length
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, width measured at elytral humeri
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.
<emphasis id="B922263FFF8CFFEDFF60FD9EFEAB8C62" bold="true" box="[245,383,512,539]" pageId="2" pageNumber="272">Coloration:</emphasis>
Elytra and head dark brown to black, pronotum notably lighter, light brown with disc darker (
<figureCitation id="136DE6A8FF8CFFEDFF35FDBBFECD8C46" box="[160,281,549,575]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1931]" captionTargetBox="[151,1436,410,1886]" captionTargetId="figure@3.[151,1436,410,1886]" captionTargetPageId="3" captionText="FIGURE 1. AI) Atyphella spp. holotypes. AC) dorsal pronotum: A) A. aphrogeneia, B) A. maritimus, C) A. marigenous; DF) scutellum: D) A. aphrogeneia, E) A. maritimus, F) A. marigenous; GI) left elytron, notice extent of costae G) A. aphrogeneia, H) A. maritimus, I) A. marigenous. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="2" pageNumber="272">Fig. 1E, H</figureCitation>
). Venter lighter than pronotal margins, becoming darker apically but before the light organ. Sculpturing on dorsal body surfaces moderately shiny beneath pubescence; pubescence fine and gold in color. Legs tan at base, darkening distally.
<emphasis id="B922263FFF8CFFEDFEE4FDF3FE6E8CFE" bold="true" box="[369,442,621,647]" pageId="2" pageNumber="272">Head:</emphasis>
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at widest point (center of eyes), eyes large, black; frons depressed, depression broad, widening and becoming shallow towards clypeus, red apunctate region at narrowest point of depression, setae long and golden on frons; antennae filiform, light brown gradually becoming lighter in color distally; scape about
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long, antennomeres 211 broadly uniform in length and about half as long as scape; mandibles sickleshaped, reddish brown, darker toward pointed apices; apical maxillary palpomere enlarged, widest almost at base, apex rounded, longer than wide; apical labial palpomere longer than wide, similar to that of apical maxillary palpomere but smaller and thinner, half as long as apical maxillary palpomere.
<emphasis id="B922263FFF8CFFEDFC7EFCDBFBB98D26" bold="true" box="[1003,1133,837,863]" pageId="2" pageNumber="272">Pronotum:</emphasis>
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long,
<quantity id="4CAE57C8FF8CFFEDFA9CFCDBFA838D26" box="[1289,1367,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="272" unit="mm" value="3.4">3.4mm</quantity>
wide, pronotal width slightly less than humeral width; margins light brown, basal third subparallel, broadly rounded anteriorly, weakly explanate, hind angles almost at right angles; disc dark brown, dorsal surface with dense punctation, with dark median region not reaching either anterior or posterior margins, median area of dorsal surface somewhat convex.
<emphasis id="B922263FFF8CFFEDFF63FC4AFEAC8D96" bold="true" box="[246,376,980,1007]" pageId="2" pageNumber="272">Scutellum:</emphasis>
trapezoidal with dense punctation, dark brown with thin, lighter lateral and posterior margins.
<emphasis id="B922263FFF8CFFEDFF02FC67FF398A6A" bold="true" box="[151,237,1017,1043]" pageId="2" pageNumber="272">Elytra:</emphasis>
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long, dark brown almost black, parallel-sided; punctures dense, not as large as that on pronotum, unevenly spaced; epipleuron developed as ridge along margin, widest at humeri, narrowing posteriorly, weakly developed towards apex; Costae somewhat pronounced, becoming less defined apically.
<emphasis id="B922263FFF8CFFEDFB14FBDEFB2B8A22" bold="true" box="[1153,1279,1088,1115]" pageId="2" pageNumber="272">Abdomen:</emphasis>
Ventrites 14 tan, becoming darker anteriorly, fat body visible at margins of ventrite 2 to ventrite 4; light organ occupying all of ventrite 56.
<emphasis id="B922263FFF8CFFEDFEB8FB17FEBA8ADA" bold="true" box="[301,366,1161,1187]" pageId="2" pageNumber="272">Legs:</emphasis>
Dark brown except for pale coxae, trochanters, and basal half of femora; dark marking present on metacoxae.
<emphasis id="B922263FFF8CFFEDFED0FB32FE158ABE" bold="true" box="[325,449,1196,1223]" pageId="2" pageNumber="272">Aedeagus:</emphasis>
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long,
<quantity id="4CAE57C8FF8CFFEDFDF0FB33FD608ABE" box="[613,692,1197,1223]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="272" unit="mm" value="0.7">0.7mm</quantity>
wide, slender (
<figureCitation id="136DE6A8FF8CFFEDFCF2FB33FC158ABE" box="[871,961,1197,1223]" captionStart="FIGURE 2" captionStartId="4.[151,250,1230,1255]" captionTargetBox="[151,1436,409,1210]" captionTargetId="figure@4.[151,1436,409,1210]" captionTargetPageId="4" captionText="FIGURE 2. AC) Atyphella spp. dorsal view of aedeagus. A) A. aphrogeneia holotype, B) A. maritimus paratype, C) A. marigenous holotype. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607107" httpUri="https://zenodo.org/record/3607107/files/figure.png" pageId="2" pageNumber="272">Fig. 2B</figureCitation>
); median lobe slightly longer than lateral lobes; posterior end of lateral lobes curve toward each other; lateral margins of lateral lobes symmetrical, tapering to apex, and rounded, diverging anteriorly; lateral lobes more sclerotized ventrally before apical fourth; median lobe apex rounded and ventrally excavated.
</paragraph>
<paragraph id="8BE9FA2DFF8CFFEDFF52FAA2FD8D8872" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52FAA2FEEB8B2E" bold="true" box="[199,319,1340,1367]" pageId="2" pageNumber="272">Variation.</emphasis>
Males: Length
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(n=10), width
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(n=10). Pronotum of some specimens lighter in color. Depression on frons sometimes deeper with apunctate region larger. Female: Length,
<typeStatus id="54ED448FFF8CFFEDFB40FAFFFAFB8B02" box="[1237,1327,1377,1403]" pageId="2" pageNumber="272" type="allotype">allotype</typeStatus>
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, width
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. Females similar to males, pubescence more dense on elytra; head is noticeably smaller and covered entirely by pronotum. Eyes much smaller, depression on frons also less developed. Ventral surface overall lighter, without darker patch on metasternite, instead small cloudy darker region on ventrites 12. Fat tissue visible through entire ventrites 16. Abdomen enlarged.
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8CFFEDFF52F98AFB3E8856" box="[199,1258,1556,1583]" pageId="2" pageNumber="272" type="distribution">
<paragraph id="8BE9FA2DFF8CFFEDFF52F98AFB3E8856" blockId="2.[151,1437,153,2015]" box="[199,1258,1556,1583]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52F98AFEB58856" bold="true" box="[199,353,1556,1583]" pageId="2" pageNumber="272">Distribution.</emphasis>
<taxonomicName id="4C5681AEFF8CFFEDFEF2F98BFD988856" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[359,588,1557,1583]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8CFFEDFEF2F98BFD988856" box="[359,588,1557,1583]" italics="true" pageId="2" pageNumber="272">Atyphella maritimus</emphasis>
</taxonomicName>
is known from the islands of Malekula and Santo,
<collectingCountry id="F341BABDFF8CFFEDFB13F98BFB328856" box="[1158,1254,1557,1583]" name="Vanuatu" pageId="2" pageNumber="272">Vanuatu</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8CFFEDFF52F9A7FE6E880E" pageId="2" pageNumber="272" type="etymology">
<paragraph id="8BE9FA2DFF8CFFEDFF52F9A7FE6E880E" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52F9A7FE9A882A" bold="true" box="[199,334,1593,1619]" pageId="2" pageNumber="272">Etymology.</emphasis>
In reference to the habitat this taxon is found in, the specific epithet for this species is derived from the Latin term for coastal.
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8CFFEDFF52F91EFB8E89A6" pageId="2" pageNumber="272" type="biology_ecology">
<paragraph id="8BE9FA2DFF8CFFEDFF52F91EFB8E89A6" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="272">
<emphasis id="B922263FFF8CFFEDFF52F91EFEF388E2" bold="true" box="[199,295,1664,1691]" pageId="2" pageNumber="272">Biology.</emphasis>
<taxonomicName id="4C5681AEFF8CFFEDFEA5F91FFDC388E2" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[304,535,1665,1691]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8CFFEDFEA5F91FFDC388E2" box="[304,535,1665,1691]" italics="true" pageId="2" pageNumber="272">Atyphella maritimus</emphasis>
</taxonomicName>
was collected on volcanic rock outcroppings subject to high tide and frequent waves from the Pacific Ocean. Larvae were observed to inhabit the crevices of these rocks with despite regular sprays of salt-water and individuals were often seen clinging to wet rocks. Adult males were caught flying out over these rocks and climbing on the same rocks as females collected. Only at one locality did we observe a second species of firefly belonging to the genus
<emphasis id="B922263FFF8CFFEDFDC9F88FFD318952" box="[604,741,1809,1835]" italics="true" pageId="2" pageNumber="272">Maganalata</emphasis>
which allowed us to infer the relationship of the males and females here. Additionally, various instars were collected at all locations suggesting that this species reproduces yearround. Nine larvae were collected from Malekula, Tenmaru and completely submerged in salt-water for fourteen hours with a 100 percent survival rate, suggesting that the larvae of this species are at least partially marine and able to withstand being submerged during high tide. The similar biology and morphology of this species to
<taxonomicName id="4C5681AEFF8CFFEDFAA4F83FFEF089A6" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8CFFEDFAA4F83FFEF089A6" italics="true" pageId="2" pageNumber="272">Atyphella aphrogeneia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8CFFEDFECFF85BFD8489A6" box="[346,592,1989,2015]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="272" phylum="Arthropoda" rank="species" species="marigneous">
<emphasis id="B922263FFF8CFFEDFECFF85BFD8489A6" box="[346,592,1989,2015]" italics="true" pageId="2" pageNumber="272">Atyphella marigneous</emphasis>
</taxonomicName>
suggests that these species are closely related.
</paragraph>
</subSubSection>
</treatment>
</document>