treatments-xml/data/03/F9/87/03F987987237FF9BFF02FA79FE9AFAB8.xml

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<document id="12214169331503C6CAF127C0755E5DF3" ID-CLB-Dataset="20693" ID-DOI="10.11646/phytotaxa.533.3.3" ID-GBIF-Dataset="c8c6b23b-ad69-4b91-89d3-057fd141c4ef" ID-ISSN="1179-3163" ID-Zenodo-Dep="6048443" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644580977120" checkinUser="plazi" docAuthor="Tamayo, Maverick N., Coritico, Fulgent P., Amoroso, Victor B., Penneys, Darin S., Tandang, Danilo N. &amp; Fritsch, Peter W." docDate="2022" docId="03F987987237FF9BFF02FA79FE9AFAB8" docLanguage="en" docName="phytotaxa.533.3.3.pdf" docOrigin="Phytotaxa 533 (3)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Vaccinium carmesinum M. N. Tamayo &amp; P. W. Fritsch 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="FFC0FFE07236FF9EFF8AFFCDFF9BFFAC" masterDocTitle="Vaccinium carmesinum (Ericaceae), a new species of blueberry from Mt. Tago Range, Mindanao Island, Philippines" masterLastPageNumber="180" masterPageNumber="173" pageNumber="174" updateTime="1699309738488" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D43B371EB48FA0574B6D52FAE40536BE">Vaccinium carmesinum (Ericaceae), a new species of blueberry from Mt. Tago Range, Mindanao Island, Philippines</mods:title>
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<mods:namePart id="4FFB0B41FD0DD59196439B8CF98BC9FC">Tamayo, Maverick N.</mods:namePart>
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<mods:affiliation id="A64A087D7880EBD18F2A0E556C195129">Department of Biology, College of Science and Engineering, Texas Christian University, 2800 South University Drive, Fort Worth, Texas, 76129 USA &amp; m. n. tamayo @ tcu. edu; https: // orcid. org / 0000 - 0003 - 0157 - 5116</mods:affiliation>
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<mods:namePart id="7A94A11B23C608DE5AC8E12D8F29C536">Coritico, Fulgent P.</mods:namePart>
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<mods:affiliation id="8EB4063571F0B49A114AC9161BDDF945">Center for Biodiversity Research and Extension in Mindanao (CEBREM), Central Mindanao University, Musuan, Bukidnon 8710 Philippines &amp; Department of Biology, College of Arts and Sciences, Central Mindanao University, Musuan, Bukidnon 8710 Philippines &amp; cfulgent @ cmu. edu. ph; https: // orcid. org / 0000 - 0003 - 3876 - 6610</mods:affiliation>
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<mods:namePart id="337F09D1F38CF69EE621019374779E6D">Penneys, Darin S.</mods:namePart>
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<mods:affiliation id="4A0B7F1E4CD82CA0F8B69042CB3964FA">Department of Biology and Marine Biology, University of North Carolina Wilmington, 601 South College Road, Wilmington, North Carolina 28403 USA &amp; penneysd @ uncw. edu; https: // orcid. org / 0000 - 0003 - 0727 - 2829</mods:affiliation>
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<mods:namePart id="FC4409F20EFD1D6C280FDBE422308F54">Tandang, Danilo N.</mods:namePart>
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<mods:affiliation id="66C465B6B2E6CEED2D13FA3E0AA5E024">Herbarium (HAST), Biodiversity Research Center, Academia Sinica, Taipei, 115 Taiwan &amp; Philippine National Herbarium, Botany and National Herbarium Division, National Museum of Natural History, National Museum of the Philippines, T. M. Kalaw St., Ermita, Manila, 1000 Philippines &amp; Biodiversity Program, Taiwan International Graduate Program, Academia Sinica and National Taiwan Normal University, Taipei, 11529 Taiwan &amp; Department of Life Science, National Taiwan Normal University, Taipei, 11677 Taiwan &amp; Biodiversity Research Center, Academia Sinica, Taipei, 11529 Taiwan &amp; sue 93653 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2708 - 661 X</mods:affiliation>
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<mods:namePart id="EA4822CB76A4C1B44741E81761B1BFB6">Fritsch, Peter W.</mods:namePart>
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<mods:affiliation id="8D9C6CF4FBA19D36FBDBE62A52A16B9E">Botanical Research Institute of Texas, 1700 University Drive, Fort Worth, Texas, 76107 USA &amp; pfritsch @ brit. org; https: // orcid. org / 0000 - 0002 - 3606 - 663 X</mods:affiliation>
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<emphasis id="B924EA9C7237FF9FFF02FA79FE02FA62" bold="true" box="[136,409,1460,1486]" italics="true" pageId="1" pageNumber="174">Vaccinium carmesinum</emphasis>
M.N.Tamayo &amp; P.W.Fritsch
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, [Mt. Tago Range,] accessory trail to peak,
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Plants and Lichens of the
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611
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(
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PNH!, isotypes A!, BRIT BRIT572077!,
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!, CMUH!, NY!)
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:—
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of
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[City] of
<location id="8E8F60557237FF9FFB24F941FAA8F90A" LSID="urn:lsid:plazi:treatment:03F987987237FF9BFF02FA79FE9AFAB8:8E8F60557237FF9FFB24F941FAA8F90A" box="[1198,1331,1676,1702]" country="Philippines" county="Mindanao Island" latitude="8.2563" longLatPrecision="5" longitude="125.14799" municipality="Province" name="Malaybalay" pageId="1" pageNumber="174" stateProvince="Bukidnon">Malaybalay</location>
,
<location id="8E8F60557237FF9FFACAF940FF65F966" LSID="urn:lsid:plazi:treatment:03F987987237FF9BFF02FA79FE9AFAB8:8E8F60557237FF9FFACAF940FF65F966" country="Philippines" county="Mindanao Island" latitude="8.2563" longLatPrecision="5" longitude="125.14799" municipality="Province" name="Barangay Kibalabag" pageId="1" pageNumber="174" stateProvince="Bukidnon">Barangay Kibalabag</location>
,
<location id="8E8F60557237FF9FFE83F97CFE28F966" LSID="urn:lsid:plazi:treatment:03F987987237FF9BFF02FA79FE9AFAB8:8E8F60557237FF9FFE83F97CFE28F966" box="[265,435,1712,1739]" country="Philippines" county="Mindanao Island" latitude="8.2563" longLatPrecision="5" longitude="125.14799" municipality="Province" name="Mt. Limbawon" pageId="1" pageNumber="174" stateProvince="Bukidnon">Mt. Limbawon</location>
, [Mt. Tago Range,] open area with
<taxonomicName id="4C504D0D7237FF9FFCB6F97DFC37F966" box="[828,940,1712,1738]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="174" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7237FF9FFCB6F97DFC37F966" box="[828,940,1712,1738]" italics="true" pageId="1" pageNumber="174">Pandanus</emphasis>
</taxonomicName>
,
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,
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,
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elevation,
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<collectingDate id="EFAAE9A67237FF9FFF02F919FEBEF943" box="[136,293,1748,1775]" pageId="1" pageNumber="174" value="2015-06-30">30 June 2015</collectingDate>
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,
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<collectorName id="26A553587237FF9FFEB8F918FE75F942" box="[306,494,1748,1774]" pageId="1" pageNumber="174">Peter W. Fritsch</collectorName>
2081
</emphasis>
(BRIT BRIT554025!,
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!);
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<location id="8E8F60557237FF9FFC65F918FBF0F942" LSID="urn:lsid:plazi:treatment:03F987987237FF9BFF02FA79FE9AFAB8:8E8F60557237FF9FFC65F918FBF0F942" box="[1007,1131,1748,1775]" country="Philippines" county="Mindanao Island" latitude="8.2563" longLatPrecision="5" longitude="125.14799" municipality="Province" name="Mt. Kiamo" pageId="1" pageNumber="174" stateProvince="Bukidnon">Mt. Kiamo</location>
summit, [Mt. Tago Range,] on ridge of heathland scrub,
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,
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,
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elevation,
<date id="FFEE104E7237FF9FFC34F934FBD1F8BE" box="[958,1098,1784,1811]" pageId="1" pageNumber="174" value="2014-05-07">
<collectingDate id="EFAAE9A67237FF9FFC34F934FBD1F8BE" box="[958,1098,1784,1811]" pageId="1" pageNumber="174" value="2014-05-07">7 May 2014</collectingDate>
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,
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<collectorName id="26A553587237FF9FFBDCF934FA80F8BE" box="[1110,1307,1784,1810]" pageId="1" pageNumber="174">Darin S. Penneys</collectorName>
2377
</emphasis>
(BRIT BRIT554030!,
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!)
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.
</paragraph>
</subSubSection>
<subSubSection id="C34A65057237FF9FFF37F88DFCADF86A" pageId="1" pageNumber="174" type="diagnosis">
<paragraph id="8BEF368E7237FF9FFF37F88DFCADF86A" blockId="1.[136,1452,1675,1991]" pageId="1" pageNumber="174">
<emphasis id="B924EA9C7237FF9FFF37F88DFDC5F8F6" box="[189,606,1856,1882]" italics="true" pageId="1" pageNumber="174">
<emphasis id="B924EA9C7237FF9FFF37F88DFECEF8F6" bold="true" box="[189,341,1856,1882]" italics="true" pageId="1" pageNumber="174">Diagnosis:—</emphasis>
<taxonomicName id="4C504D0D7237FF9FFEDCF88CFDC5F8F6" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[342,606,1857,1882]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="174" phylum="Tracheophyta" rank="species" species="carmesinum">Vaccinium carmesinum</taxonomicName>
</emphasis>
resembles
<taxonomicName id="4C504D0D7237FF9FFD68F88CFBE6F8F6" authority="Merrill (1917: 294)" authorityName="Merrill" authorityPageNumber="294" authorityYear="1917" box="[738,1149,1856,1882]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="174" phylum="Tracheophyta" rank="species" species="platyphyllum">
<emphasis id="B924EA9C7237FF9FFD68F88CFC09F8F6" box="[738,914,1856,1882]" italics="true" pageId="1" pageNumber="174">V. platyphyllum</emphasis>
<bibRefCitation id="EFC14B7F7237FF9FFC17F88DFBE6F8F6" author="Merrill, E. D." box="[925,1149,1856,1882]" pageId="1" pageNumber="174" pagination="1 - 391" refId="ref5090" refString="Merrill, E. D. (1917) New Philippine shrubs and trees. Philippine Journal of Science, section C, Botany, 12 (1): 1 - 391." type="journal article" year="1917">Merrill (1917: 294)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C504D0D7237FF9FFB33F88CFE95F8D3" authority="S. Vidal (1886: 168)" authorityName="S. Vidal" authorityPageNumber="168" authorityYear="1886" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="174" phylum="Tracheophyta" rank="species" species="luzoniense">
<emphasis id="B924EA9C7237FF9FFB33F88CFAD5F8F6" box="[1209,1358,1856,1882]" italics="true" pageId="1" pageNumber="174">V. luzoniense</emphasis>
S.
<bibRefCitation id="EFC14B7F7237FF9FFAE3F88DFE95F8D3" author="Vidal, S." pageId="1" pageNumber="174" pagination="1 - 455" refId="ref5629" refString="Vidal, S. (1886) Revision de plantas vasculares Filipinas, Fam. 67 - Vacciniaceas. Establecimiento Tipo-Litografico de M. Perez, Hijo, Binondo, Manila. pp. 1 - 455." type="book chapter" year="1886">Vidal (1886: 168)</bibRefCitation>
</taxonomicName>
, but differs from the former by longer and wider leaves, longer racemes, longer bracts, glabrous corollas, and glabrescent fruits, and from the latter by longer petioles, leaf glands distributed along the length of the blade margin, a glabrous inflorescence rachis, and lanate filaments.
</paragraph>
</subSubSection>
<caption id="DF2F66067234FF9CFF02F8F2FE4CF830" ID-DOI="http://doi.org/10.5281/zenodo.6048445" ID-Zenodo-Dep="6048445" httpUri="https://zenodo.org/record/6048445/files/figure.png" pageId="2" pageNumber="175" startId="2.[136,229,1855,1877]" targetBox="[192,1372,218,1800]" targetPageId="2">
<paragraph id="8BEF368E7234FF9CFF02F8F2FE4CF830" blockId="2.[136,1452,1855,1949]" pageId="2" pageNumber="175">
<emphasis id="B924EA9C7234FF9CFF02F8F2FE9BF8F8" bold="true" box="[136,256,1855,1877]" pageId="2" pageNumber="175">FIGURE 1.</emphasis>
<emphasis id="B924EA9C7234FF9CFE8DF88DFE6BF8F9" box="[263,496,1856,1877]" italics="true" pageId="2" pageNumber="175">
<taxonomicName id="4C504D0D7234FF9CFE8DF88DFE70F8F9" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[263,491,1856,1877]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="175" phylum="Tracheophyta" rank="species" species="carmesinum">Vaccinium carmesinum</taxonomicName>
.
</emphasis>
<emphasis id="B924EA9C7234FF9CFE7CF8F2FD95F8F8" bold="true" box="[502,526,1855,1876]" pageId="2" pageNumber="175">A.</emphasis>
Leafy branchlet.
<emphasis id="B924EA9C7234FF9CFD37F8F2FD48F8F8" bold="true" box="[701,723,1855,1876]" pageId="2" pageNumber="175">B.</emphasis>
Inflorescences showing flower buds and foliaceous bracts.
<emphasis id="B924EA9C7234FF9CFA95F8F2FAADF8F9" bold="true" box="[1311,1334,1855,1877]" pageId="2" pageNumber="175">C.</emphasis>
Corolla.
<emphasis id="B924EA9C7234FF9CFA1EF8F2FA37F8F8" bold="true" box="[1428,1452,1855,1876]" pageId="2" pageNumber="175">D.</emphasis>
Dissected flower showing stamens, style, ovary, hypanthium, and calyx.
<emphasis id="B924EA9C7234FF9CFCC4F8AEFCFFF8D4" bold="true" box="[846,868,1891,1912]" pageId="2" pageNumber="175">E.</emphasis>
Dorsal view of stamen.
<emphasis id="B924EA9C7234FF9CFBDFF8AEFBFCF8D4" bold="true" box="[1109,1127,1891,1912]" pageId="2" pageNumber="175">F.</emphasis>
Ventral view of stamen.
<emphasis id="B924EA9C7234FF9CFAD7F8AEFAEDF8D5" bold="true" box="[1373,1398,1891,1913]" pageId="2" pageNumber="175">G.</emphasis>
Fruit cross section. Illustration by MNT.
</paragraph>
</caption>
<caption id="DF2F66067235FF9DFF02F8B4FEDEF87B" ID-DOI="http://doi.org/10.5281/zenodo.6048448" ID-Zenodo-Dep="6048448" httpUri="https://zenodo.org/record/6048448/files/figure.png" pageId="3" pageNumber="176" startId="3.[136,229,1913,1935]" targetBox="[151,1436,190,1889]" targetPageId="3">
<paragraph id="8BEF368E7235FF9DFF02F8B4FEDEF87B" blockId="3.[136,1452,1913,2007]" pageId="3" pageNumber="176">
<emphasis id="B924EA9C7235FF9DFF02F8B4FF67F822" bold="true" box="[136,252,1913,1935]" pageId="3" pageNumber="176">FIGURE 2.</emphasis>
<taxonomicName id="4C504D0D7235FF9DFE8BF8B7FE44F823" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[257,479,1914,1935]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7235FF9DFE8BF8B7FE44F823" box="[257,479,1914,1935]" italics="true" pageId="3" pageNumber="176">Vaccinium carmesinum</emphasis>
</taxonomicName>
.
<emphasis id="B924EA9C7235FF9DFE60F8B4FD87F823" bold="true" box="[490,540,1913,1935]" pageId="3" pageNumber="176">A, B.</emphasis>
Flowering branchlet.
<emphasis id="B924EA9C7235FF9DFD67F8B4FC9FF823" bold="true" box="[749,772,1913,1935]" pageId="3" pageNumber="176">C.</emphasis>
Leafy branchlet.
<emphasis id="B924EA9C7235FF9DFC20F8B7FC5AF823" bold="true" box="[938,961,1914,1935]" pageId="3" pageNumber="176">D.</emphasis>
Inflorescences showing flower buds and foliaceous bracts.
<emphasis id="B924EA9C7235FF9DFF44F853FF7FF81F" bold="true" box="[206,228,1950,1971]" pageId="3" pageNumber="176">E.</emphasis>
Longitudinal section of flower showing corolla, style and stamens.
<emphasis id="B924EA9C7235FF9DFCF2F853FC11F81F" bold="true" box="[888,906,1950,1971]" pageId="3" pageNumber="176">F.</emphasis>
Immature fruits. Photographs: A, D by DSP; B, E, F by MAKP; C by DNT.
</paragraph>
</caption>
<subSubSection id="C34A65057232FF9AFF37FF53FF6FFA94" pageId="4" pageNumber="177" type="description">
<paragraph id="8BEF368E7232FF9AFF37FF53FF6FFA94" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37FF53FEF3FF14" bold="true" box="[189,360,158,184]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37FF53FED7FF14" bold="true" box="[189,332,158,184]" italics="true" pageId="4" pageNumber="177">Description:</emphasis>
</emphasis>
Terrestrial leaning shrub or tree, evergreen,
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tall, sparsely branched.
<emphasis id="B924EA9C7232FF9AFB37FF53FAA0FF15" bold="true" box="[1213,1339,158,185]" italics="true" pageId="4" pageNumber="177">Branchlets</emphasis>
glabrous, red when young, grayish brown at maturity, terete,
<quantity id="4CA89B6B7232FF9AFD59FF0FFCA9FF71" box="[723,818,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="4" pageNumber="177" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">38 mm</quantity>
wide, lenticellate; perennating buds compressed-ovoid,
<quantity id="4CA89B6B7232FF9AFF02FF2BFE95FEAD" box="[136,270,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="4" pageNumber="177" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
long; bud scales overlapping with minutely ciliate margins.
<emphasis id="B924EA9C7232FF9AFC24FF2AFC66FEAD" bold="true" box="[942,1021,231,257]" italics="true" pageId="4" pageNumber="177">Leaves</emphasis>
persistent on older branchlets, spirally and evenly arranged, slightly overlapping, internodes
<quantity id="4CA89B6B7232FF9AFD70FEC7FCCBFE89" box="[762,848,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="4" pageNumber="177" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">15 cm</quantity>
long; petiole crimson red, in cross section rounded abaxially and slightly raised adaxially, 1018 ×
<quantity id="4CA89B6B7232FF9AFD1DFEE3FD6EFEE5" box="[663,757,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="4" pageNumber="177" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">15 mm</quantity>
, glabrous; lamina broadly elliptic, ovate, or rarely subrounded, with the larger leaves on each branchlet 715 ×
<quantity id="4CA89B6B7232FF9AFD28FE9FFC96FEC1" box="[674,781,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="9.0" metricValueMin="0.4" pageId="4" pageNumber="177" unit="cm" value="4.7" valueMax="9.0" valueMin="0.4">0.49 cm</quantity>
, coriaceous, both surfaces reddish when young turning pale green abaxially and glossy adaxially,
<emphasis id="B924EA9C7232FF9AFDBFFEBAFD15FE3C" box="[565,654,375,400]" italics="true" pageId="4" pageNumber="177">in sicco</emphasis>
both surfaces light brown to ferrugineous, without punctae, glabrous; midvein flattened or sunken adaxially, strongly raised abaxially, secondary veins 3 or 4 on each side of midvein with first pair arising from base and remainder along midvein, arc-ascending, abaxially raised, adaxially sunken, tertiary veins faintly evident or obscure, base cuneate to truncate, margin entire, weakly revolute, apex slightly acuminate, marginal glands sunken, 1018 per side, scattered along length of margin but more concentrated towards the apex,
<quantity id="4CA89B6B7232FF9AFAF0FDCBFF46FDE9" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="177" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3 0.5 mm</quantity>
wide.
<emphasis id="B924EA9C7232FF9AFEAFFDE7FE59FDE9" bold="true" box="[293,450,554,581]" italics="true" pageId="4" pageNumber="177">Inflorescence</emphasis>
pseudo-terminal or terminal, racemose, developing beyond confines of perennating bud, 1 per axil,
<quantity id="4CA89B6B7232FF9AFF61FD83FEA5FDC5" box="[235,318,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="177" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">68 cm</quantity>
long at anthesis, densely 10- to 12-flowered; peduncle and rachis crimson red, slightly ridged, terete, glabrous; flower bracts caducous, crimson red, dark brown
<emphasis id="B924EA9C7232FF9AFC90FDBEFCEBFD20" box="[794,880,627,652]" italics="true" pageId="4" pageNumber="177">in sicco</emphasis>
, foliaceous, ovate to elliptic, planar or occasionally cucullate, 615 ×
<quantity id="4CA89B6B7232FF9AFEC5FD5BFE4CFD1D" box="[335,471,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="4" pageNumber="177" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
, coriaceous, glabrous, margin entire, ciliolate, with several yellowish or reddish globose glands,
<quantity id="4CA89B6B7232FF9AFF68FD77FE1CFD79" box="[226,391,698,725]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="177" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.150.20 mm</quantity>
diameter mainly on basal half and with cilia ca.
<quantity id="4CA89B6B7232FF9AFC4BFD77FB81FD79" box="[961,1050,698,725]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="177" unit="mm" value="0.1">0.1 mm</quantity>
mainly on apical third, apex acute to obtuse with a terminal gland.
<emphasis id="B924EA9C7232FF9AFE74FD13FDC9FD55" bold="true" box="[510,594,734,761]" italics="true" pageId="4" pageNumber="177">Pedicel</emphasis>
3.515 ×
<quantity id="4CA89B6B7232FF9AFD40FD13FCC8FD55" box="[714,851,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="4" pageNumber="177" unit="mm" value="0.7" valueMax="0.9" valueMin="0.5">0.50.9 mm</quantity>
at anthesis, terete, spreading, glabrous, occasionally with 1 or 2 globose glands near base or occasionally on apical half, ebracteolate.
<emphasis id="B924EA9C7232FF9AFB93FCCFFBEDFCB1" bold="true" box="[1049,1142,770,797]" italics="true" pageId="4" pageNumber="177">Flowers</emphasis>
articulated at junction with pedicel,
<quantity id="4CA89B6B7232FF9AFF6CFCEBFEFDFCED" box="[230,358,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="12.0" metricValueMin="3.5" pageId="4" pageNumber="177" unit="mm" value="7.75" valueMax="12.0" valueMin="3.5">3.512 mm</quantity>
long.
<emphasis id="B924EA9C7232FF9AFE21FCEBFDA6FCED" bold="true" box="[427,573,806,833]" italics="true" pageId="4" pageNumber="177">Hypanthium</emphasis>
crimson red, cupuliform, 1.52 ×
<quantity id="4CA89B6B7232FF9AFC36FCEBFBB4FCED" box="[956,1071,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="177" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
, white-hirsutulous with trichomes
<quantity id="4CA89B6B7232FF9AFF02FC87FEB0FCC9" box="[136,299,842,869]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="177" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.100.15 mm</quantity>
long; calyx limb 1.0
<quantity id="4CA89B6B7232FF9AFD95FC87FDEFFCC9" box="[543,628,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="177" unit="mm" value="1.2">1.2 mm</quantity>
long, white-hirsutulous; calyx lobes broadly triangular,
<quantity id="4CA89B6B7232FF9AFB6CFC87FAF7FCC9" box="[1254,1388,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="177" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
long, white hirsutulous, margin entire, often ciliolate, apex acute, with a prominent greenish (reddish
<emphasis id="B924EA9C7232FF9AFB6CFCA2FADAFC24" box="[1254,1345,879,904]" italics="true" pageId="4" pageNumber="177">in sicco</emphasis>
) globose sessile terminal gland ca.
<quantity id="4CA89B6B7232FF9AFE25FC5FFD8FFC01" box="[431,532,914,941]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="177" unit="mm" value="0.25">0.25 mm</quantity>
diameter.
<emphasis id="B924EA9C7232FF9AFD06FC5FFD78FC01" bold="true" box="[652,739,914,941]" italics="true" pageId="4" pageNumber="177">Corolla</emphasis>
broadly acute, lustrous white, conical-urceolate, 712 ×
<quantity id="4CA89B6B7232FF9AFAE6FC5FFF2DFC7D" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="6.0" metricValueMin="2.5" pageId="4" pageNumber="177" unit="mm" value="4.25" valueMax="6.0" valueMin="2.5">2.56 mm</quantity>
, outside glabrous, inside white-lanate especially on upper and lower third, trichomes
<quantity id="4CA89B6B7232FF9AFBECFC7BFB4CFC7D" box="[1126,1239,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="177" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long; corolla lobes 5 or 6, 12 ×
<quantity id="4CA89B6B7232FF9AFEAEFC17FE02FC59" box="[292,409,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="177" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
, apex acute to obtuse.
<emphasis id="B924EA9C7232FF9AFD16FC17FD65FC58" bold="true" box="[668,766,986,1012]" italics="true" pageId="4" pageNumber="177">Stamens</emphasis>
8 to 10, monomorphic, distinct,
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long; filaments white, straight, gradually dilated at base,
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long, pink towards base, densely white-lanate with trichomes
<quantity id="4CA89B6B7232FF9AFF02FBEFFF66FB91" box="[136,253,1058,1085]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="177" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long; anthers
<quantity id="4CA89B6B7232FF9AFE2FFBEFFD81FB91" box="[421,538,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="4" pageNumber="177" unit="mm" value="2.2" valueMax="2.4" valueMin="2.0">22.4 mm</quantity>
long, cells
<quantity id="4CA89B6B7232FF9AFD29FBEFFCB6FB91" box="[675,813,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="4" pageNumber="177" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
long, echinulate, tubules parallel, narrowly cylindrical, distinctly narrower than cells, opening by oblique ventrally oriented apical pores,
<quantity id="4CA89B6B7232FF9AFB84FB8BFB08FBCD" box="[1038,1171,1094,1121]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="177" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
long, pore apex rounded, spurs absent.
<emphasis id="B924EA9C7232FF9AFEAAFBA7FEFDFB28" bold="true" box="[288,358,1130,1156]" italics="true" pageId="4" pageNumber="177">Ovary</emphasis>
5- or 6-locular but appearing pseudo-10- to 12-locular with false partitions extending ca.
<quantity id="4CA89B6B7232FF9AFADCFBA7FA37FB29" box="[1366,1452,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="177" unit="mm" value="1.5">1.5 mm</quantity>
from inner wall; ovules in two columns per locule.
<emphasis id="B924EA9C7232FF9AFD37FB43FD69FB05" bold="true" box="[701,754,1166,1193]" italics="true" pageId="4" pageNumber="177">Disk</emphasis>
disciform, ca.
<quantity id="4CA89B6B7232FF9AFC1CFB43FC4DFB05" box="[918,982,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="177" unit="mm" value="2.0">2 mm</quantity>
in diameter, puberulent, margin shallowly ridged.
<emphasis id="B924EA9C7232FF9AFF55FB7FFE8CFB61" bold="true" box="[223,279,1202,1229]" italics="true" pageId="4" pageNumber="177">Style</emphasis>
reddish, not exserted from corolla,
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long, glabrous.
<emphasis id="B924EA9C7232FF9AFC6BFB7FFB85FB60" bold="true" box="[993,1054,1202,1228]" italics="true" pageId="4" pageNumber="177">Fruit</emphasis>
on pedicels
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long, deep purple, dark brown or reddish
<emphasis id="B924EA9C7232FF9AFE56FB1AFDA9FB5C" box="[476,562,1239,1264]" italics="true" pageId="4" pageNumber="177">in sicco</emphasis>
, globose, slightly ridged, 46 ×
<quantity id="4CA89B6B7232FF9AFC16FB1BFC60FB5D" box="[924,1019,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="177" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
, glabrescent except for minute cilia on calyx lobe margins; persistent calyx lobes erect; disk ca.
<quantity id="4CA89B6B7232FF9AFC8EFB37FCC1FAB9" box="[772,858,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="177" unit="mm" value="4.5">4.5 mm</quantity>
in diameter.
<emphasis id="B924EA9C7232FF9AFC66FB37FBB6FAB9" bold="true" box="[1004,1069,1274,1301]" italics="true" pageId="4" pageNumber="177">Seeds</emphasis>
numerous, minute, brown, ca.
<quantity id="4CA89B6B7232FF9AFA02FB37FF2FFA95" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="177" unit="mm" value="0.8">0.8 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C34A65057232FF9AFF37FA8FFB9BFA40" pageId="4" pageNumber="177" type="distribution">
<paragraph id="8BEF368E7232FF9AFF37FA8FFB9BFA40" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37FA8FFD64FAF0" box="[189,767,1346,1373]" italics="true" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37FA8FFE66FAF1" bold="true" box="[189,509,1346,1373]" italics="true" pageId="4" pageNumber="177">Distribution and Habitat:—</emphasis>
<taxonomicName id="4C504D0D7232FF9AFE76FA8EFD64FAF0" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[508,767,1347,1372]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="carmesinum">Vaccinium carmesinum</taxonomicName>
</emphasis>
is endemic to two mountains (Mt. Kiamo and Mt. Limbawon) in Mt. Tago Range, Mindanao, growing in tropical lower montane rainforest to upper montane rainforest. Populations of
<taxonomicName id="4C504D0D7232FF9AFF20FA46FEC8FA08" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[170,339,1419,1444]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7232FF9AFF20FA46FEC8FA08" box="[170,339,1419,1444]" italics="true" pageId="4" pageNumber="177">V. carmesinum</emphasis>
</taxonomicName>
were mostly found near summits where they thrive on volcanic-igneous or clay substrate with abundant humus. They also occur in areas of open shaded mossy forests, or on ridges covered in heathland scrub.
<typeStatus id="54EB882C7232FF9AFF02FA1EFF6EFA41" box="[136,245,1491,1517]" pageId="4" pageNumber="177" type="paratype">Paratypes</typeStatus>
of the new species were collected on ca. 1030% west-facing slopes.
</paragraph>
</subSubSection>
<subSubSection id="C34A65057232FF9AFF37FA3BFAA9F8A0" pageId="4" pageNumber="177" type="etymology">
<paragraph id="8BEF368E7232FF9AFF37FA3BFBE5F9F4" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37FA3BFEC6F9BD" bold="true" box="[189,349,1526,1553]" italics="true" pageId="4" pageNumber="177">Etymology:—</emphasis>
The epithet
<taxonomicName id="4C504D0D7232FF9AFE69FA3AFDF7F9BC" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[483,620,1527,1552]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7232FF9AFE69FA3AFDF7F9BC" box="[483,620,1527,1552]" italics="true" pageId="4" pageNumber="177">carmesinum</emphasis>
</taxonomicName>
is derived from the Greek word for crimson (blood red), as depicted by its crimson red petioles, floral bracts, peduncle, rachis, pedicels, hypanthium, and calyces. Moreover, a crimson red stain is extracted in notable quantity when the plants are soaked in a denatured alcohol solution.
</paragraph>
<paragraph id="8BEF368E7232FF9AFF37F9AFFC13F9D1" blockId="4.[136,1452,158,2057]" box="[189,904,1634,1661]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37F9AFFEC5F9D0" bold="true" box="[189,350,1634,1661]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37F9AFFEDAF9D1" bold="true" box="[189,321,1634,1661]" italics="true" pageId="4" pageNumber="177">Phenology:</emphasis>
</emphasis>
Flowering in June. Fruiting from January to May.
</paragraph>
<paragraph id="8BEF368E7232FF9AFF37F94BFAA9F8A0" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37F94BFDA7F90C" bold="true" box="[189,572,1670,1697]" italics="true" pageId="4" pageNumber="177">Proposed Conservation Status:—</emphasis>
Mt. Tago Range has not been extensively explored botanically, which results in uncertainty as to the conservation status of the species. This range is a non-protected area; thus, the extent of occurrence and area of occupancy for the species cannot be assessed. There are only two populations currently known. Hence, we recommend a conservation status of data deficient (DD) (
<bibRefCitation id="EFC14B7F7232FF9AFC80F93FFABDF8A0" author="IUCN Standards &amp; Petitions Committee" box="[778,1318,1778,1804]" pageId="4" pageNumber="177" refId="ref4811" refString="IUCN Standards and Petitions Committee. (2019) Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed: 14 September 2021)" type="url" year="2019">IUCN Standards and Petitions Committee 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A65057232FF9BFF37F8DBFE9AFAB8" lastPageId="5" lastPageNumber="178" pageId="4" pageNumber="177" type="discussion">
<paragraph id="8BEF368E7232FF9AFF37F8DBFBC6F7A4" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="177">
<emphasis id="B924EA9C7232FF9AFF37F8DBFEFAF89C" bold="true" box="[189,353,1814,1840]" italics="true" pageId="4" pageNumber="177">Discussion:—</emphasis>
In its combination of morphological characters,
<taxonomicName id="4C504D0D7232FF9AFC09F8DAFB12F89C" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[899,1161,1815,1840]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7232FF9AFC09F8DAFB12F89C" box="[899,1161,1815,1840]" italics="true" pageId="4" pageNumber="177">Vaccinium carmesinum</emphasis>
</taxonomicName>
matches no other species treated in relevant taxonomic treatments. In the artificial key to the species of Philippine
<taxonomicName id="4C504D0D7232FF9AFBE6F8F6FA3CF8F8" authority="(Copeland 1930)" baseAuthorityName="Copeland" baseAuthorityYear="1930" box="[1132,1447,1850,1876]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7232FF9AFBE6F8F6FB7AF8F8" box="[1132,1249,1851,1876]" italics="true" pageId="4" pageNumber="177">Vaccinium</emphasis>
(
<bibRefCitation id="EFC14B7F7232FF9AFB65F8F7FA04F8F8" author="Copeland, H. F." box="[1263,1439,1850,1876]" pageId="4" pageNumber="177" pagination="537 - 607" refId="ref4298" refString="Copeland, H. F. (1930) Philippine Ericaceae, II: the species of Vaccinium. Philippine Journal of Science 42: 537 - 607." type="journal article" year="1930">Copeland 1930</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C504D0D7232FF9AFF02F892FEB4F8D4" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[136,303,1887,1912]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7232FF9AFF02F892FEB4F8D4" box="[136,303,1887,1912]" italics="true" pageId="4" pageNumber="177">V. carmesinum</emphasis>
</taxonomicName>
keys to
<taxonomicName id="4C504D0D7232FF9AFE1CF892FDDDF8D4" box="[406,582,1886,1912]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="platyphyllum">
<emphasis id="B924EA9C7232FF9AFE1CF892FDDDF8D4" box="[406,582,1886,1912]" italics="true" pageId="4" pageNumber="177">V. platyphyllum</emphasis>
</taxonomicName>
. The new species differs from
<taxonomicName id="4C504D0D7232FF9AFC38F892FBF9F8D4" box="[946,1122,1886,1912]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="platyphyllum">
<emphasis id="B924EA9C7232FF9AFC38F892FBF9F8D4" box="[946,1122,1886,1912]" italics="true" pageId="4" pageNumber="177">V. platyphyllum</emphasis>
</taxonomicName>
by having longer and wider leaves (715 ×
<quantity id="4CA89B6B7232FF9AFEBCF84FFE04F831" box="[310,415,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="9.0" metricValueMin="0.4" pageId="4" pageNumber="177" unit="cm" value="4.7" valueMax="9.0" valueMin="0.4">0.49 cm</quantity>
vs.
<quantity id="4CA89B6B7232FF9AFE45F84FFDA5F831" box="[463,574,1922,1949]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="4" pageNumber="177" unit="cm" value="12.5" valueMax="14.0" valueMin="11.0">1114 cm</quantity>
×
<quantity id="4CA89B6B7232FF9AFDD7F84FFD2EF831" box="[605,693,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="177" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">57 cm</quantity>
), longer racemes (
<quantity id="4CA89B6B7232FF9AFC0EF84FFC42F831" box="[900,985,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="177" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">68 cm</quantity>
vs.
<quantity id="4CA89B6B7232FF9AFB83F84FFBFAF831" box="[1033,1121,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="177" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
), longer bracts (
<quantity id="4CA89B6B7232FF9AFA9DF84FFA1FF831" box="[1303,1412,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="4" pageNumber="177" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">615 mm</quantity>
vs. ca.
<quantity id="4CA89B6B7232FF9AFF3AF86BFF6DF86D" box="[176,246,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="177" unit="mm" value="8.0">8 mm</quantity>
), longer pedicels (
<quantity id="4CA89B6B7232FF9AFE4FF86BFDDCF86D" box="[453,583,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="15.0" metricValueMin="3.5" pageId="4" pageNumber="177" unit="mm" value="9.25" valueMax="15.0" valueMin="3.5">3.515 mm</quantity>
vs. ca.
<quantity id="4CA89B6B7232FF9AFD14F86BFD7BF86D" box="[670,736,1958,1985]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="177" unit="mm" value="8.0">8 mm</quantity>
) that are glabrous (vs. slightly pubescent) and ebracteolate (vs. bracteolate), a glabrous (vs. sparsely pubescent) corolla outside, longer anthers (2.0
<quantity id="4CA89B6B7232FF9AFBD0F807FB28F849" box="[1114,1203,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="177" unit="mm" value="2.3">2.3 mm</quantity>
vs. ca.
<quantity id="4CA89B6B7232FF9AFA9AF807FAF6F849" box="[1296,1389,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="177" unit="mm" value="1.5">1.5 mm</quantity>
), and longer (4.06.0 mm vs. ca.
<quantity id="4CA89B6B7232FF9AFE31F823FE66F7A5" box="[443,509,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="177" unit="mm" value="3.0">3 mm</quantity>
) and glabrescent (vs. pubescent) fruits (
<bibRefCitation id="EFC14B7F7232FF9AFC36F823FBCBF7A4" author="Merrill, E. D." box="[956,1104,2030,2056]" pageId="4" pageNumber="177" pagination="1 - 391" refId="ref5090" refString="Merrill, E. D. (1917) New Philippine shrubs and trees. Philippine Journal of Science, section C, Botany, 12 (1): 1 - 391." type="journal article" year="1917">Merrill 1917</bibRefCitation>
).
</paragraph>
<paragraph id="8BEF368E7233FF9BFF37FF52FAC6FEC1" blockId="5.[136,1452,158,1300]" pageId="5" pageNumber="178">
<taxonomicName id="4C504D0D7233FF9BFF37FF52FE58FF14" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[189,451,159,184]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFF37FF52FE58FF14" box="[189,451,159,184]" italics="true" pageId="5" pageNumber="178">Vaccinium carmesinum</emphasis>
</taxonomicName>
can be distinguished from all other species of Philippine
<taxonomicName id="4C504D0D7233FF9BFBC5FF52FB5FFF14" baseAuthorityName="Copeland" baseAuthorityYear="1930" box="[1103,1220,159,184]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFBC5FF52FB5FFF14" box="[1103,1220,159,184]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
</taxonomicName>
by its leaves, which are the widest of any
<taxonomicName id="4C504D0D7233FF9BFEF3FF0EFE75FF70" baseAuthorityName="Copeland" baseAuthorityYear="1930" box="[377,494,195,220]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFEF3FF0EFE75FF70" box="[377,494,195,220]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
</taxonomicName>
in the
<collectingCountry id="F347761E7233FF9BFDB0FF0FFD27FF70" box="[570,700,194,220]" name="Philippines" pageId="5" pageNumber="178">Philippines</collectingCountry>
. The pedicels are also notably ebracteolate and have 0 to 2 globose glands near the base or occasionally on the apical half. These glands might be homologous with bracteoles (typically two per pedicel in
<taxonomicName id="4C504D0D7233FF9BFED7FEC6FE49FE88" baseAuthorityName="Copeland" baseAuthorityYear="1930" box="[349,466,267,292]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFED7FEC6FE49FE88" box="[349,466,267,292]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
</taxonomicName>
) with a reduction in size and/or number.
<bibRefCitation id="EFC14B7F7233FF9BFC2DFEC7FBFEFE88" author="Copeland, H. F." box="[935,1125,266,292]" pageId="5" pageNumber="178" pagination="537 - 607" refId="ref4298" refString="Copeland, H. F. (1930) Philippine Ericaceae, II: the species of Vaccinium. Philippine Journal of Science 42: 537 - 607." type="journal article" year="1930">Copeland (1930)</bibRefCitation>
mentioned pedicel glands in
<taxonomicName id="4C504D0D7233FF9BFF02FEE2FE87FEE4" box="[136,284,302,328]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="luzoniense">
<emphasis id="B924EA9C7233FF9BFF02FEE2FE87FEE4" box="[136,284,302,328]" italics="true" pageId="5" pageNumber="178">V. luzoniense</emphasis>
</taxonomicName>
. Unfortunately, this character was not thoroughly described for the other Philippine species in former publication where the absence of bracteoles in a specimen is noted as “unobserved” (i.e.
<bibRefCitation id="EFC14B7F7233FF9BFBE0FE9FFA93FEC1" author="Sleumer, H." box="[1130,1288,338,365]" pageId="5" pageNumber="178" pagination="469 - 914" refId="ref5379" refString="Sleumer, H. (1966 - 1967) Ericaceae: In: van Steenis, C. G. G. J. (Ed.) Flora Malesiana. Ser. 1, Vol. 6, Parts 4, 5. Wolters-Noordhoff, Groningen, pp. 469 - 914." type="book chapter" year="1966">Sleumer 1966</bibRefCitation>
1967).
</paragraph>
<paragraph id="8BEF368E7233FF9BFF37FEBAFC38FD1C" blockId="5.[136,1452,158,1300]" pageId="5" pageNumber="178">
<taxonomicName id="4C504D0D7233FF9BFF37FEBAFE5DFE3C" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[189,454,375,400]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFF37FEBAFE5DFE3C" box="[189,454,375,400]" italics="true" pageId="5" pageNumber="178">Vaccinium carmesinum</emphasis>
</taxonomicName>
is a member of
<emphasis id="B924EA9C7233FF9BFD05FEBAFD3FFE3C" box="[655,676,375,400]" italics="true" pageId="5" pageNumber="178">V.</emphasis>
section
<taxonomicName id="4C504D0D7233FF9BFC83FEBAFC5EFE3C" authority="Nakai" authorityName="Nakai" box="[777,965,374,400]" class="Magnoliopsida" family="Ericaceae" genus="Bracteata" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFC83FEBAFCECFE3C" box="[777,887,375,400]" italics="true" pageId="5" pageNumber="178">Bracteata</emphasis>
Nakai
</taxonomicName>
in
<bibRefCitation id="EFC14B7F7233FF9BFC7BFEBBFAF9FE3C" author="Nakai, T. &amp; Koidzumi, G." box="[1009,1378,374,400]" pageId="5" pageNumber="178" refId="ref5187" refString="Nakai, T. &amp; Koidzumi, G. (1927) Trees and shrubs indigenous in Japan proper (revised Ed.), Vol. 1. Seibido Shoten, Tokyo, 714 pp." type="book" year="1927">Nakai &amp; Koidzumi (1927: 234)</bibRefCitation>
sensu Sleumer (
<bibRefCitation id="EFC14B7F7233FF9BFF7DFE57FE02FE19" author="Sleumer, H." box="[247,409,410,437]" pageId="5" pageNumber="178" pagination="469 - 914" refId="ref5379" refString="Sleumer, H. (1966 - 1967) Ericaceae: In: van Steenis, C. G. G. J. (Ed.) Flora Malesiana. Ser. 1, Vol. 6, Parts 4, 5. Wolters-Noordhoff, Groningen, pp. 469 - 914." type="book chapter" year="1966">Sleumer 1966</bibRefCitation>
1967) as based on the combination of many-flowered racemose inflorescences, caducous foliaceous bracts, absence of a membranaceous wing at the sinuses of the corolla, and anthers that open by short introrse slits or terminal pores (
<bibRefCitation id="EFC14B7F7233FF9BFE67FE2FFD15FE50" author="Sleumer, H." box="[493,654,482,508]" pageId="5" pageNumber="178" pagination="469 - 914" refId="ref5379" refString="Sleumer, H. (1966 - 1967) Ericaceae: In: van Steenis, C. G. G. J. (Ed.) Flora Malesiana. Ser. 1, Vol. 6, Parts 4, 5. Wolters-Noordhoff, Groningen, pp. 469 - 914." type="book chapter" year="1966">Sleumer 1966</bibRefCitation>
1967;
<bibRefCitation id="EFC14B7F7233FF9BFD69FE2FFC11FE50" author="Co, L. L. &amp; Madulid, D. &amp; Argent, G." box="[739,906,482,509]" pageId="5" pageNumber="178" pagination="373 - 376" refId="ref4243" refString="Co, L. L., Madulid, D. &amp; Argent, G. (2002) A new species of Vaccinium (Ericaceae) from the Philippines. Edinburgh Journal of Botany 59 (3): 373 - 376. https: // doi. org / 10.1017 / S 0960428602000227" type="journal article" year="2002">
Co
<emphasis id="B924EA9C7233FF9BFC86FE2EFCA4FE50" box="[780,831,482,508]" italics="true" pageId="5" pageNumber="178">et al</emphasis>
. 2002
</bibRefCitation>
;
<bibRefCitation id="EFC14B7F7233FF9BFC1CFE2FFBF5FE50" author="Salares, V. B. &amp; Obico, J. J. A. &amp; Ormerod, P. &amp; Barcelona, J. F. &amp; Pelser, P. B." box="[918,1134,482,509]" pageId="5" pageNumber="178" pagination="255 - 262" refId="ref5274" refString="Salares, V. B., Obico, J. J. A., Ormerod, P., Barcelona, J. F. &amp; Pelser, P. B. (2018) Taxonomic novelties from Cebu: a new species of Vaccinium (Ericaceae) and a new record of Phaius (Orchidaceae) for the Philippines. Phytotaxa 360 (3): 255 - 262. https: // doi. org / 10.11646 / phytotaxa. 360.3.5" type="journal article" year="2018">
Salares
<emphasis id="B924EA9C7233FF9BFC65FE2EFBB9FE50" box="[1007,1058,482,508]" italics="true" pageId="5" pageNumber="178">et al</emphasis>
. 2018
</bibRefCitation>
). In
<bibRefCitation id="EFC14B7F7233FF9BFB2BFE2FFAC7FE51" author="Sleumer, H." box="[1185,1372,482,509]" pageId="5" pageNumber="178" pagination="469 - 914" refId="ref5379" refString="Sleumer, H. (1966 - 1967) Ericaceae: In: van Steenis, C. G. G. J. (Ed.) Flora Malesiana. Ser. 1, Vol. 6, Parts 4, 5. Wolters-Noordhoff, Groningen, pp. 469 - 914." type="book chapter" year="1966">Sleumers (1966</bibRefCitation>
1967) key to the Malesian
<emphasis id="B924EA9C7233FF9BFEF9FDCAFE13FD8C" box="[371,392,519,544]" italics="true" pageId="5" pageNumber="178">V.</emphasis>
section
<taxonomicName id="4C504D0D7233FF9BFE62FDCAFDCDFD8C" box="[488,598,519,544]" class="Magnoliopsida" family="Ericaceae" genus="Bracteata" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFE62FDCAFDCDFD8C" box="[488,598,519,544]" italics="true" pageId="5" pageNumber="178">Bracteata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D0D7233FF9BFDEDFDCAFC96FD8C" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[615,781,519,544]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFDEDFDCAFC96FD8C" box="[615,781,519,544]" italics="true" pageId="5" pageNumber="178">V. carmesinum</emphasis>
</taxonomicName>
keys to
<taxonomicName id="4C504D0D7233FF9BFCFBFDCAFB9FFD8C" box="[881,1028,518,544]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="luzoniense">
<emphasis id="B924EA9C7233FF9BFCFBFDCAFB9FFD8C" box="[881,1028,518,544]" italics="true" pageId="5" pageNumber="178">V. luzoniense</emphasis>
</taxonomicName>
.
<taxonomicName id="4C504D0D7233FF9BFB9EFDCAFA80FD8C" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[1044,1307,519,544]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFB9EFDCAFA80FD8C" box="[1044,1307,519,544]" italics="true" pageId="5" pageNumber="178">Vaccinium carmesinum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C504D0D7233FF9BFF02FDE6FE81FDE8" box="[136,282,554,580]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="luzoniense">
<emphasis id="B924EA9C7233FF9BFF02FDE6FE81FDE8" box="[136,282,554,580]" italics="true" pageId="5" pageNumber="178">V. luzoniense</emphasis>
</taxonomicName>
, however, by having longer petioles (
<quantity id="4CA89B6B7233FF9BFD49FDE7FCA6FDE9" box="[707,829,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="5" pageNumber="178" unit="mm" value="14.0" valueMax="18.0" valueMin="10.0">1018 mm</quantity>
vs. ca.
<quantity id="4CA89B6B7233FF9BFC19FDE7FC7DFDE9" box="[915,998,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="178" unit="mm" value="10.0">10 mm</quantity>
), longer and wider leaves (715 ×
<quantity id="4CA89B6B7233FF9BFAE6FDE7FF31FDC5" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="9.0" metricValueMin="0.4" pageId="5" pageNumber="178" unit="cm" value="4.7" valueMax="9.0" valueMin="0.4">0.49 cm</quantity>
vs.
<quantity id="4CA89B6B7233FF9BFF51FD83FEABFDC5" box="[219,304,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="5" pageNumber="178" unit="cm" value="8.0" valueMax="9.0" valueMin="7.0">79 cm</quantity>
×
<quantity id="4CA89B6B7233FF9BFEDBFD83FE24FDC5" box="[337,447,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="5" pageNumber="178" unit="cm" value="3.75" valueMax="4.5" valueMin="3.0">34.5 cm</quantity>
), with leaf glands distributed along the length of the leaf margin (vs. with merely a pair of glands near the base), glabrous rachis (vs. with capitate-glandular trichomes), white (vs. red) corollas, and densely lanate (vs. sparsely pubescent) filaments (
<bibRefCitation id="EFC14B7F7233FF9BFDD6FD5BFD47FD1C" author="Vidal, S." box="[604,732,662,688]" pageId="5" pageNumber="178" pagination="1 - 455" refId="ref5629" refString="Vidal, S. (1886) Revision de plantas vasculares Filipinas, Fam. 67 - Vacciniaceas. Establecimiento Tipo-Litografico de M. Perez, Hijo, Binondo, Manila. pp. 1 - 455." type="book chapter" year="1886">Vidal 1886</bibRefCitation>
;
<bibRefCitation id="EFC14B7F7233FF9BFD62FD5BFC0DFD1C" author="Copeland, H. F." box="[744,918,662,688]" pageId="5" pageNumber="178" pagination="537 - 607" refId="ref4298" refString="Copeland, H. F. (1930) Philippine Ericaceae, II: the species of Vaccinium. Philippine Journal of Science 42: 537 - 607." type="journal article" year="1930">Copeland 1930</bibRefCitation>
).
</paragraph>
<paragraph id="8BEF368E7233FF9BFF37FD76FA1AFCED" blockId="5.[136,1452,158,1300]" pageId="5" pageNumber="178">
In the key to the Bornean species of
<taxonomicName id="4C504D0D7233FF9BFDE7FD76FC0BFD78" authority="(Argent 2018)" baseAuthorityName="Argent" baseAuthorityYear="2018" box="[621,912,698,724]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFDE7FD76FD79FD78" box="[621,738,699,724]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
(
<bibRefCitation id="EFC14B7F7233FF9BFD79FD77FC13FD78" author="Argent, G." box="[755,904,698,724]" pageId="5" pageNumber="178" pagination="55 - 172" refId="ref4153" refString="Argent, G. (2018) Rigiolepis and Vaccinium (Ericaceae) in Borneo. Edinburgh Journal of Botany 76: 55 - 172. https: // doi. org / 10.1017 / S 0960428618000276" type="journal article" year="2018">Argent 2018</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C504D0D7233FF9BFC14FD76FBDEFD78" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[926,1093,699,724]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFC14FD76FBDEFD78" box="[926,1093,699,724]" italics="true" pageId="5" pageNumber="178">V. carmesinum</emphasis>
</taxonomicName>
keys to
<taxonomicName id="4C504D0D7233FF9BFB26FD76FE42FD54" authority="Argent (2018: 108)" authorityName="Argent" authorityPageNumber="108" authorityYear="2018" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="subSpecies" species="sarawakense" subSpecies="montanum">
<emphasis id="B924EA9C7233FF9BFB26FD76FAC0FD78" box="[1196,1371,698,724]" italics="true" pageId="5" pageNumber="178">V. sarawakense</emphasis>
subsp.
<emphasis id="B924EA9C7233FF9BFF02FD12FF65FD54" box="[136,254,735,760]" italics="true" pageId="5" pageNumber="178">montanum</emphasis>
<bibRefCitation id="EFC14B7F7233FF9BFE8EFD13FE42FD54" author="Argent, G." box="[260,473,734,760]" pageId="5" pageNumber="178" pagination="55 - 172" refId="ref4153" refString="Argent, G. (2018) Rigiolepis and Vaccinium (Ericaceae) in Borneo. Edinburgh Journal of Botany 76: 55 - 172. https: // doi. org / 10.1017 / S 0960428618000276" type="journal article" year="2018">Argent (2018: 108)</bibRefCitation>
</taxonomicName>
but differs from it by having an inflorescence with fewer flowers (10- to 12-flowered vs. 7- to 20-flowered), glabrous rachis (vs. densely covered by short brown curved glandular trichomes), calyx lobes with a sessile terminal gland (vs. absent), white (vs. pale pink) corollas, and the absence of anther spurs (vs. presence).
</paragraph>
<paragraph id="8BEF368E7233FF9BFF37FC86FD2CFB90" blockId="5.[136,1452,158,1300]" pageId="5" pageNumber="178">
In the sectional treatment of
<taxonomicName id="4C504D0D7233FF9BFD8CFC86FB87FCC8" authority="(Vander Kloet and Dickinson 2009)" baseAuthorityName="Vander Kloet and Dickinson" baseAuthorityYear="2009" box="[518,1052,842,868]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFD8CFC86FDE0FCC8" box="[518,635,843,868]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
(
<bibRefCitation id="EFC14B7F7233FF9BFD01FC87FB8FFCC8" author="Vander Kloet, S. P. &amp; Dickinson, T. A." box="[651,1044,842,868]" pageId="5" pageNumber="178" pagination="253 - 268" refId="ref5549" refString="Vander Kloet, S. P. &amp; Dickinson, T. A. (2009) A subgeneric classification of the genus Vaccinium and the metamorphosis of V. section Bracteata Nakai: more terrestrial and less epiphytic in habit, more continental and less insular in distribution. Journal of Plant Research 122 (3): 253 - 268. https: // doi. org / 10.1007 / s 10265 - 008 - 0211 - 7" type="journal article" year="2009">Vander Kloet and Dickinson 2009</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C504D0D7233FF9BFBA3FC86FB54FCC8" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[1065,1231,843,868]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFBA3FC86FB54FCC8" box="[1065,1231,843,868]" italics="true" pageId="5" pageNumber="178">V. carmesinum</emphasis>
</taxonomicName>
can be treated as a member of
<emphasis id="B924EA9C7233FF9BFE8CFCA2FE8CFC24" box="[262,279,879,904]" italics="true" pageId="5" pageNumber="178">V</emphasis>
. section
<taxonomicName id="4C504D0D7233FF9BFEF2FCA2FC96FC24" authority="Schlechter (1919: 174)" authorityName="Schlechter" authorityPageNumber="174" authorityYear="1919" box="[376,781,878,904]" class="Magnoliopsida" family="Ericaceae" genus="Euepigynium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFEF2FCA2FD97FC24" box="[376,524,879,904]" italics="true" pageId="5" pageNumber="178">Euepigynium</emphasis>
<bibRefCitation id="EFC14B7F7233FF9BFD9BFCA3FC96FC24" author="Schlechter, R." box="[529,781,878,904]" pageId="5" pageNumber="178" pagination="137 - 194" refId="ref5359" refString="Schlechter, R. (1919) Die Ericaceen von Deutsch-Neu-Guinea. Botanische Jahrbucher 55: 137 - 194." type="journal article" year="1919">Schlechter (1919: 174)</bibRefCitation>
</taxonomicName>
by its evergreen habit, monomorphic perennating buds, each with more than two scales, one perennating bud per leaf axil, plinerved leaf blade venation, entire leaf blade margin, peduncle longer than pedicels, calyx tube completely fused to the ovary, and pseudo-10-locular ovary. However, the boundaries of
<emphasis id="B924EA9C7233FF9BFEDDFC16FEF7FC58" box="[343,364,987,1012]" italics="true" pageId="5" pageNumber="178">V.</emphasis>
section
<taxonomicName id="4C504D0D7233FF9BFE46FC16FDFBFC58" authorityName="Schlechter" authorityYear="1919" box="[460,608,987,1012]" class="Magnoliopsida" family="Ericaceae" genus="Euepigynium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFE46FC16FDFBFC58" box="[460,608,987,1012]" italics="true" pageId="5" pageNumber="178">Euepigynium</emphasis>
</taxonomicName>
and other sections of Malesian
<taxonomicName id="4C504D0D7233FF9BFC45FC16FBDFFC58" baseAuthorityName="Vander Kloet and Dickinson" baseAuthorityYear="2009" box="[975,1092,987,1012]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFC45FC16FBDFFC58" box="[975,1092,987,1012]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
</taxonomicName>
delimited by
<bibRefCitation id="EFC14B7F7233FF9BFB6EFC17FED0FBB4" author="Vander Kloet, S. P. &amp; Dickinson, T. A." pageId="5" pageNumber="178" pagination="253 - 268" refId="ref5549" refString="Vander Kloet, S. P. &amp; Dickinson, T. A. (2009) A subgeneric classification of the genus Vaccinium and the metamorphosis of V. section Bracteata Nakai: more terrestrial and less epiphytic in habit, more continental and less insular in distribution. Journal of Plant Research 122 (3): 253 - 268. https: // doi. org / 10.1007 / s 10265 - 008 - 0211 - 7" type="journal article" year="2009">Vander Kloet and Dickinson (2009)</bibRefCitation>
were vaguely defined (i.e. the species included in each section are not provided). Hence, the sectional limits of
<taxonomicName id="4C504D0D7233FF9BFF67FBEEFEF9FB90" baseAuthorityName="Vander Kloet and Dickinson" baseAuthorityYear="2009" box="[237,354,1059,1084]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9C7233FF9BFF67FBEEFEF9FB90" box="[237,354,1059,1084]" italics="true" pageId="5" pageNumber="178">Vaccinium</emphasis>
</taxonomicName>
in
<collectingCountry id="F347761E7233FF9BFE0CFBEFFE44FB90" box="[390,479,1058,1084]" name="Malaysia" pageId="5" pageNumber="178">Malesia</collectingCountry>
need further study.
</paragraph>
<paragraph id="8BEF368E7233FF9BFF37FB8BFE9AFAB8" blockId="5.[136,1452,158,1300]" pageId="5" pageNumber="178">
During the process of diagnosing
<taxonomicName id="4C504D0D7233FF9BFDC4FB8AFCCCFBCC" authorityName="M. N. Tamayo &amp; P. W. Fritsch" authorityYear="2022" box="[590,855,1095,1120]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="carmesinum">
<emphasis id="B924EA9C7233FF9BFDC4FB8AFCCCFBCC" box="[590,855,1095,1120]" italics="true" pageId="5" pageNumber="178">Vaccinium carmesinum</emphasis>
</taxonomicName>
as distinct from other Philippine species, we have become cognizant of problems in the taxonomy of the Philippine species. For example,
<taxonomicName id="4C504D0D7233FF9BFBF1FBA6FF20FB04" authority="Merrill (1919: 441)" authorityName="Merrill" authorityPageNumber="441" authorityYear="1919" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="ilocanum">
<emphasis id="B924EA9C7233FF9BFBF1FBA6FA9AFB28" box="[1147,1281,1130,1156]" italics="true" pageId="5" pageNumber="178">V. ilocanum</emphasis>
<bibRefCitation id="EFC14B7F7233FF9BFA80FBA7FF20FB04" author="Merrill, E. D." pageId="5" pageNumber="178" pagination="1 - 694" refId="ref5124" refString="Merrill, E. D. (1919) New or noteworthy Philippine plants, XV. Philippine Journal of Science 14 (4): 1 - 694." type="journal article" year="1919">Merrill (1919: 441)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C504D0D7233FF9BFF79FB42FDD3FB04" authority="Merrill (1925: 43)" authorityName="Merrill" authorityPageNumber="43" authorityYear="1925" box="[243,584,1166,1193]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="rizalense">
<emphasis id="B924EA9C7233FF9BFF79FB42FEEFFB04" box="[243,372,1166,1192]" italics="true" pageId="5" pageNumber="178">V. rizalense</emphasis>
<bibRefCitation id="EFC14B7F7233FF9BFEF1FB43FDD3FB04" author="Merrill, E. D." box="[379,584,1166,1193]" pageId="5" pageNumber="178" pagination="1 - 59" refId="ref5154" refString="Merrill, E. D. (1925) New species of Philippine plants collected by A. Loher. Philippine Journal of Science 27 (1): 1 - 59." type="journal article" year="1925">Merrill (1925: 43)</bibRefCitation>
</taxonomicName>
were synonymized under
<taxonomicName id="4C504D0D7233FF9BFCFBFB42FB84FB04" box="[881,1055,1166,1192]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="platyphyllum">
<emphasis id="B924EA9C7233FF9BFCFBFB42FB84FB04" box="[881,1055,1166,1192]" italics="true" pageId="5" pageNumber="178">V. platyphyllum</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC14B7F7233FF9BFBC0FB43FA9CFB04" author="Copeland, H. F." box="[1098,1287,1166,1192]" pageId="5" pageNumber="178" pagination="537 - 607" refId="ref4298" refString="Copeland, H. F. (1930) Philippine Ericaceae, II: the species of Vaccinium. Philippine Journal of Science 42: 537 - 607." type="journal article" year="1930">Copeland (1930)</bibRefCitation>
but characters seem divergent among these species and the justification relied mainly on macroscopic characters. A detailed study of this complex is currently in progress with emphasis on, e.g., ovary indumentum, corolla surfaces, and stamen characters.
</paragraph>
</subSubSection>
</treatment>
</document>