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<document ID-DOI="http://doi.org/10.5281/zenodo.4763785" ID-GBIF-Dataset="2cbeea15-656e-466c-b2d2-23a455cc9428" ID-GBIF-Taxon="182074931" ID-ISSN="1854-0392" ID-Zenodo-Dep="4763785" checkinTime="1620961553388" checkinUser="felipe" docAuthor="Zwick, Peter &amp; Baumann, Richard W." docDate="2011" docId="03F98790FFCFFF9EFEB1FDD8333FF8EA" docLanguage="en" docName="Illiesia.7.15.148-155.pdf" docOrigin="Illiesia 7 (15)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Nemoura jejudoensis Zwick &amp; Baumann 2011, sp. n." docType="treatment" docVersion="5" lastPageId="2" lastPageNumber="150" masterDocId="FFC0FFE8FFCEFF9CFFFBFFE5365CFF9D" masterDocTitle="Nemoura Jejudoensis A New Species Of Stonefly And The Redescription Of Amphinemura Baei Ham And Lee (Plecoptera, Nemouridae) From Jeju Island, Korea" masterLastPageNumber="155" masterPageNumber="148" pageId="1" pageNumber="149" updateTime="1632012747112" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nemoura Jejudoensis A New Species Of Stonefly And The Redescription Of Amphinemura Baei Ham And Lee (Plecoptera, Nemouridae) From Jeju Island, Korea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zwick, Peter</mods:namePart>
<mods:affiliation>Schwarzer Stock 9, D- 36110 Schlitz, Germany E-mail: pleco-p. zwick @ t-online. de</mods:affiliation>
<mods:nameIdentifier type="email">pleco-p.zwick@t-online.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, UT, U. S. A. 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>15</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>148</mods:start>
<mods:end>155</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760189</mods:identifier>
<mods:identifier type="GBIF-Dataset">2cbeea15-656e-466c-b2d2-23a455cc9428</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4760189</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4763785" ID-GBIF-Taxon="182074931" ID-Zenodo-Dep="4763785" LSID="urn:lsid:plazi:treatment:03F98790FFCFFF9EFEB1FDD8333FF8EA" httpUri="http://treatment.plazi.org/id/03F98790FFCFFF9EFEB1FDD8333FF8EA" lastPageId="2" lastPageNumber="150" pageId="1" pageNumber="149">
<subSubSection box="[330,657,572,597]" pageId="1" pageNumber="149" type="nomenclature">
<paragraph blockId="1.[330,657,572,597]" box="[330,657,572,597]" pageId="1" pageNumber="149">
<heading bold="true" box="[330,657,572,597]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="149" reason="2">
<emphasis bold="true" box="[330,657,572,597]" pageId="1" pageNumber="149">
<taxonomicName authority="Zwick &amp; Baumann, 2011" authorityName="Zwick &amp; Baumann" authorityYear="2011" box="[330,578,572,596]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="jejudoensis" status="sp. n.">
<emphasis bold="true" box="[330,578,572,596]" italics="true" pageId="1" pageNumber="149">Nemoura jejudoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[592,657,573,597]" pageId="1" pageNumber="149" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[435,551,609,633]" pageId="1" pageNumber="149" type="description">
<paragraph blockId="1.[435,551,609,633]" box="[435,551,609,633]" pageId="1" pageNumber="149">
(
<figureCitation box="[444,543,609,633]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[189,244,1212,1236]" captionStartId-1="3.[189,241,999,1023]" captionTargetBox-0="[217,1410,252,1156]" captionTargetBox-1="[223,1370,290,946]" captionTargetId-0="figure-67@2.[211,1420,243,1163]" captionTargetId-1="figure-67@3.[211,1420,279,958]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." captionText-1="Figs. 5-6. Nemoura jejudoensis, female. 5. Ventral view of abdominal tip. 6. Dorsal view of cleared vagina." figureDoi-0="http://doi.org/10.5281/zenodo.4760191" figureDoi-1="http://doi.org/10.5281/zenodo.4760193" httpUri-0="https://zenodo.org/record/4760191/files/figure.png" httpUri-1="https://zenodo.org/record/4760193/files/figure.png" pageId="1" pageNumber="149">Figs. 1-6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="149" type="materials_examined">
<paragraph blockId="1.[189,798,679,1913]" pageId="1" pageNumber="149">
<emphasis bold="true" box="[189,435,680,704]" pageId="1" pageNumber="149">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111697313" collectingDate="1986-12-01" collectorName="M. F. Whiting" country="South Korea" county="Cheju Island" location="Chonyeon waterfalls" municipality="Chungmoon City" pageId="1" pageNumber="149" specimenCount="4" specimenCount-male="4" stateProvince="Jeju Island" typeStatus="holotype">
<typeStatus box="[453,565,680,704]" pageId="1" pageNumber="149">Holotype</typeStatus>
<specimenCount box="[581,601,679,704]" pageId="1" pageNumber="149" type="male">♂</specimenCount>
,
<specimenCount box="[624,658,679,704]" pageId="1" pageNumber="149" type="male">3♂</specimenCount>
<typeStatus box="[674,792,680,704]" pageId="1" pageNumber="149">paratypes</typeStatus>
,
<collectingCountry box="[189,406,716,740]" name="South Korea" pageId="1" pageNumber="149">Republic of Korea</collectingCountry>
, Cheju Island,
<collectingMunicipality box="[583,793,716,740]" pageId="1" pageNumber="149">Chungmoon City</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03F98790FFCFFF9EFEB1FDD8333FF8EA:8E8F605DFFCFFF9DFF46FD0A37EEFC9A" box="[189,434,751,775]" country="South Korea" county="Cheju Island" municipality="Chungmoon City" name="Chonyeon waterfalls" pageId="1" pageNumber="149" stateProvince="Jeju Island">Chonyeon waterfalls</location>
,
<date box="[443,637,751,775]" pageId="1" pageNumber="149" value="1986-12-01">
<collectingDate box="[443,637,751,775]" pageId="1" pageNumber="149" value="1986-12-01">1 December 1986</collectingDate>
</date>
,
<collectorName box="[646,797,751,775]" pageId="1" pageNumber="149">M.F. Whiting</collectorName>
[
<collectingRegion box="[195,322,787,811]" pageId="1" pageNumber="149">Jeju Island</collectingRegion>
, Seogwipo City, Jungmun, Cheonjeyeon falls]
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111697314" collectingDate="1986-12-01" collectorName="M. F. Whiting" country="South Korea" county="Cheju Island" location="Chonyeon" municipality="Chungmoon City" pageId="1" pageNumber="149" specimenCount="16" specimenCount-female="15" specimenCount-male="1" stateProvince="Jeju">
Additional material with the same collection data:
<specimenCount box="[254,301,857,882]" pageId="1" pageNumber="149" type="female">13♀</specimenCount>
plus
<specimenCount box="[365,399,857,882]" pageId="1" pageNumber="149" type="male">1♂</specimenCount>
and
<specimenCount box="[457,490,857,882]" pageId="1" pageNumber="149" type="female">2♀</specimenCount>
used for SEM study
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="149" type="description">
<paragraph blockId="1.[189,798,679,1913]" pageId="1" pageNumber="149">
<emphasis bold="true" box="[189,404,893,917]" pageId="1" pageNumber="149">Size and habitus.</emphasis>
Front wing length 6.4-7.4 (mean 6.8) mm in
<specimenCount box="[328,348,928,953]" pageId="1" pageNumber="149" type="male">♂</specimenCount>
, 8.4-9.9 (mean 9.2) mm in
<specimenCount box="[669,689,928,953]" pageId="1" pageNumber="149" type="female">♀</specimenCount>
. Habitus typical of genus, not characteristic. Body and antennae brown, legs and palpi paler, yellowish brown. Wing veins brown, membrane of front wing with faint brownish tinge.
</paragraph>
<paragraph blockId="1.[189,798,679,1913]" pageId="1" pageNumber="149">
<emphasis bold="true" box="[189,258,1107,1131]" pageId="1" pageNumber="149">Male.</emphasis>
Tergite 9 medially short, otherwise unmodified.
<collectionCode box="[361,378,1143,1167]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="149">T</collectionCode>
10 forms the medially largely membranous base plate of the epiproct. Epiproct closely appressed to base plate, basally firmly connected, difficult to detach for study. The epiproct consists of a short body and a long anterior rostrumlike process.
</paragraph>
<paragraph blockId="1.[189,798,679,1913]" pageId="1" pageNumber="149">
Body of epiproct short, wider than long (
<figureCitation box="[715,780,1356,1380]" captionStart="Figs" captionStartId="2.[189,244,1212,1236]" captionTargetBox="[217,1410,252,1156]" captionTargetId="figure-67@2.[211,1420,243,1163]" captionTargetPageId="2" captionText="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." figureDoi="http://doi.org/10.5281/zenodo.4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="1" pageNumber="149">Fig. 1</figureCitation>
). The transverse anterodorsal sclerite supports two backwardly turned claw-like plates each with four marginal bare teeth. There is an additional anteromedian tooth whose mediodorsal face is beset with cuticular spicules (
<figureCitation box="[491,565,1533,1558]" captionStart="Figs" captionStartId="2.[189,244,1212,1236]" captionTargetBox="[217,1410,252,1156]" captionTargetId="figure-67@2.[211,1420,243,1163]" captionTargetPageId="2" captionText="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." figureDoi="http://doi.org/10.5281/zenodo.4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="1" pageNumber="149">Fig. 4</figureCitation>
). A short distance behind the anterodorsal sclerite lies another slender transverse sclerite which is laterally angled backward. Other sclerites are seen only in slidemounted epriprocts, by transparency (
<figureCitation box="[642,706,1675,1700]" captionStart="Figs" captionStartId="2.[189,244,1212,1236]" captionTargetBox="[217,1410,252,1156]" captionTargetId="figure-67@2.[211,1420,243,1163]" captionTargetPageId="2" captionText="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." figureDoi="http://doi.org/10.5281/zenodo.4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="1" pageNumber="149">Fig. 1</figureCitation>
). There are no ventral setae or teeth. The basolateral earshaped structures that provide the normally-shaped epiprocts of other
<taxonomicName authorityName="Zwick &amp; Baumann" authorityYear="2011" box="[447,547,1783,1806]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis box="[447,547,1783,1806]" italics="true" pageId="1" pageNumber="149">Nemoura</emphasis>
</taxonomicName>
-species with some mobility can be recognized but seem rigid, nonfunctional.
</paragraph>
<paragraph blockId="1.[189,798,679,1913]" lastBlockId="1.[835,1444,253,1912]" pageId="1" pageNumber="149">
Rostrum rising from a bulb-like base connected to the medial ends of the anterolateral sclerites. A central sclerite with median division line appears by transparency. Basal half of rostrum with several oblique folds, in side view seen to be triangularly raised near midlength (
<figureCitation box="[1110,1177,395,419]" captionStart="Figs" captionStartId="2.[189,244,1212,1236]" captionTargetBox="[217,1410,252,1156]" captionTargetId="figure-67@2.[211,1420,243,1163]" captionTargetPageId="2" captionText="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." figureDoi="http://doi.org/10.5281/zenodo.4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="1" pageNumber="149">Fig. 2</figureCitation>
).
</paragraph>
<paragraph blockId="1.[835,1444,253,1912]" pageId="1" pageNumber="149">Sternite 9 almost circular, with short pointed caudal tip. The club-shaped vesicle occupies 4/5 of the sternite length. The short blade-like median sclerites of the paraprocts lie alongside the tip of S9. A membranous section separates them from the triangular, apically quite narrow outer lobe.</paragraph>
<paragraph blockId="1.[835,1444,253,1912]" pageId="1" pageNumber="149">
Cerci curved forward and mediad, partly concealing the epiproct in dorsal view (
<figureCitation box="[1353,1428,679,703]" captionStart="Figs" captionStartId="2.[189,244,1212,1236]" captionTargetBox="[217,1410,252,1156]" captionTargetId="figure-67@2.[211,1420,243,1163]" captionTargetPageId="2" captionText="Figs.1-4. Nemoura jejudoensis sp. n., male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites. Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus." figureDoi="http://doi.org/10.5281/zenodo.4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="1" pageNumber="149">Fig. 3</figureCitation>
). Medial flat face pale and convex outer face brown, and pilose. The tiny rudiment of segment 2 is at the top of the curved cercus, on the soft medial face. There is an inconspicuous subterminal point on the front side of the cercus.
</paragraph>
<paragraph blockId="1.[835,1444,253,1912]" pageId="1" pageNumber="149">
<emphasis bold="true" box="[835,929,892,916]" pageId="1" pageNumber="149">Female.</emphasis>
Similar to the female of
<taxonomicName authorityName="Wu" authorityYear="1929" box="[1212,1286,893,916]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="geei">
<emphasis box="[1212,1286,893,916]" italics="true" pageId="1" pageNumber="149">N. geei</emphasis>
</taxonomicName>
as illustrated by
<bibRefCitation author="Shimizu, T." box="[875,1048,928,952]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu (1997</bibRefCitation>
, his fig. 48). The almost circular pregenital plate on S7 covers the middle of S8 almost completely, the curved edges of the unpigmented vaginal lobes may be faintly visible (
<figureCitation box="[1263,1330,1035,1059]" captionStart="Figs" captionStartId="3.[189,241,999,1023]" captionTargetBox="[223,1370,290,946]" captionTargetId="figure-67@3.[211,1420,279,958]" captionTargetPageId="3" captionText="Figs. 5-6. Nemoura jejudoensis, female. 5. Ventral view of abdominal tip. 6. Dorsal view of cleared vagina." figureDoi="http://doi.org/10.5281/zenodo.4760193" httpUri="https://zenodo.org/record/4760193/files/figure.png" pageId="1" pageNumber="149">Fig. 5</figureCitation>
). Sides of S9 with a poorly defined elongate pigmented strip. Triangular sclerites in the dorsal wall of the genital opening and the end of the oviduct, respectively, may be visible by transparency.
</paragraph>
<paragraph blockId="1.[835,1444,253,1912]" pageId="1" pageNumber="149">
In cleared genitalia (
<figureCitation box="[1108,1173,1212,1236]" captionStart="Figs" captionStartId="3.[189,241,999,1023]" captionTargetBox="[223,1370,290,946]" captionTargetId="figure-67@3.[211,1420,279,958]" captionTargetPageId="3" captionText="Figs. 5-6. Nemoura jejudoensis, female. 5. Ventral view of abdominal tip. 6. Dorsal view of cleared vagina." figureDoi="http://doi.org/10.5281/zenodo.4760193" httpUri="https://zenodo.org/record/4760193/files/figure.png" pageId="1" pageNumber="149">Fig. 6</figureCitation>
), two anterolateral caplike brown sclerites are seen at the side of a long carrot-shaped transparent tube connecting to the receptacula. Behind it the dorsal wall of the vagina has two paramedian areas with concentric cuticular rings, probably extensible pouches.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="150" pageId="1" pageNumber="149" type="diagnosis">
<paragraph blockId="1.[835,1444,253,1912]" lastBlockId="2.[189,798,1532,1911]" lastPageId="2" lastPageNumber="150" pageId="1" pageNumber="149">
<emphasis bold="true" box="[835,964,1425,1449]" pageId="1" pageNumber="149">Diagnosis.</emphasis>
A member of the East Asian
<taxonomicName authorityName="Kawai" authorityYear="1960" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="ovocercia">
<emphasis italics="true" pageId="1" pageNumber="149">Nemoura ovocercia</emphasis>
</taxonomicName>
-group (sensu
<bibRefCitation author="Shimizu, T." box="[1115,1283,1461,1486]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
).
<taxonomicName authority="Wu 1929" authorityName="Wu" authorityYear="1929" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="geei">
<emphasis box="[1307,1388,1462,1485]" italics="true" pageId="1" pageNumber="149">N. geei</emphasis>
Wu 1929
</taxonomicName>
occurs on the Asian mainland (
<collectingCountry box="[1278,1352,1497,1521]" name="China" pageId="1" pageNumber="149">China</collectingCountry>
,
<collectingCountry box="[1365,1438,1497,1520]" name="South Korea" pageId="1" pageNumber="149">Korea</collectingCountry>
, Russian Far East) as well as on Hokkaido and Honshu.
<taxonomicName authority="Claassen 1929" authorityName="Claassen" authorityYear="1929" box="[948,1349,1568,1592]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="pekinensis">
<emphasis box="[948,1169,1568,1592]" italics="true" pageId="1" pageNumber="149">Nemoura pekinensis</emphasis>
Claassen 1929
</taxonomicName>
and
<taxonomicName authority="Zhiltzova 1982" authorityName="Zhiltzova" authorityYear="1982" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="brevicercia">
<emphasis italics="true" pageId="1" pageNumber="149">N. brevicercia</emphasis>
<bibRefCitation author="Zhiltzova, L. A." box="[970,1156,1603,1628]" pageId="1" pageNumber="149" pagination="37 - 43" refId="ref3738" refString="Zhiltzova, L. A. 1982. New species of Stoneflies of Family Nemouridae (Plecoptera) of the Far East. Vestnik Zoologi, 1982: 37 - 43. (In Russian, English summary)." type="journal article" year="1982">Zhiltzova 1982</bibRefCitation>
</taxonomicName>
are junior synonyms (
<bibRefCitation author="Shimizu, T." box="[843,1010,1639,1663]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
). We studied some specimens from mainland
<collectingCountry box="[954,1028,1675,1698]" name="South Korea" pageId="1" pageNumber="149">Korea</collectingCountry>
. Excellent illustrations of
<taxonomicName authorityName="Wu" authorityYear="1929" box="[1326,1401,1675,1698]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="geei">
<emphasis box="[1326,1401,1675,1698]" italics="true" pageId="1" pageNumber="149">N. geei</emphasis>
</taxonomicName>
are also available (
<bibRefCitation author="Zhiltzova, L. A." box="[1013,1192,1710,1734]" pageId="1" pageNumber="149" pagination="37 - 43" refId="ref3738" refString="Zhiltzova, L. A. 1982. New species of Stoneflies of Family Nemouridae (Plecoptera) of the Far East. Vestnik Zoologi, 1982: 37 - 43. (In Russian, English summary)." type="journal article" year="1982">Zhiltzova 1982</bibRefCitation>
,
<bibRefCitation author="Zhiltzova, L. A." box="[1206,1262,1710,1734]" pageId="1" pageNumber="149" pagination="537" refId="ref3778" refString="Zhiltzova, L. A. 2003. Plecoptera Gruppe Euholognatha. - Fauna of Russia and neighbouring countries, Insecta Plecoptera, 1 (1): 537 + 7 unnumbered pages; Russian Academy of Sciences, Zoological Institute, St. Petersburg, Nauka. (In Russian.)" type="journal article" year="2003">2003</bibRefCitation>
,
<bibRefCitation author="Shimizu, T." box="[1275,1438,1710,1734]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
, Teslenko &amp; Zhiltzova 2007). Several additional species occur in
<collectingCountry box="[1051,1120,1782,1805]" name="Japan" pageId="1" pageNumber="149">Japan</collectingCountry>
, the surrounding islands,
<collectingCountry box="[835,923,1817,1841]" name="Taiwan" pageId="1" pageNumber="149">Taiwan</collectingCountry>
(
<bibRefCitation author="Shimizu, T." box="[937,1098,1817,1841]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
), and
<collectingCountry box="[1166,1237,1817,1841]" name="China" pageId="1" pageNumber="149">China</collectingCountry>
(
<bibRefCitation author="Li, W. H. &amp; D. Yang" box="[1250,1431,1817,1841]" pageId="1" pageNumber="149" pagination="65 - 68" refId="ref3611" refString="Li, W. H. and D. Yang. 2007. Two new species of Nemoura (Plecoptera: Nemouridae) from Henan, China. Zootaxa, 1511 (21): 65 - 68." type="journal article" year="2007">Li &amp; Yang 2007</bibRefCitation>
).
<emphasis box="[835,993,1853,1876]" italics="true" pageId="1" pageNumber="149">
<taxonomicName authorityName="Wu" authorityYear="1929" box="[835,989,1853,1876]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="geei">Nemoura geei</taxonomicName>
,
</emphasis>
the present species,
<taxonomicName authority="Shimizu 1997" authorityName="Shimizu" authorityYear="1997" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="quadrituberata">
<emphasis box="[1247,1443,1852,1876]" italics="true" pageId="1" pageNumber="149">N. quadrituberata</emphasis>
<bibRefCitation author="Shimizu, T." box="[835,1007,1888,1912]" pageId="1" pageNumber="149" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
</taxonomicName>
(Tsushima Island),
<taxonomicName authority="Shimizu 1997" authorityName="Shimizu" authorityYear="1997" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" lastPageId="2" lastPageNumber="150" order="Plecoptera" pageId="1" pageNumber="149" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis box="[1277,1443,1888,1912]" italics="true" pageId="1" pageNumber="149">N. longilobata</emphasis>
<bibRefCitation author="Shimizu, T." box="[189,352,1532,1556]" pageId="2" pageNumber="150" pagination="193 - 218" refId="ref3650" refString="Shimizu, T. 1997. The species of the Nemoura ovocercia group (Plecoptera: Nemouridae). Aquatic Insects, 19: 193 - 218." type="journal article" year="1997">Shimizu 1997</bibRefCitation>
</taxonomicName>
(Hokkaido), and
<taxonomicName authority="Li &amp; Yang 2007" authorityName="Li &amp; Yang" authorityYear="2007" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="atristrigata">
<emphasis box="[572,731,1532,1556]" italics="true" pageId="2" pageNumber="150">N. atristrigata</emphasis>
<bibRefCitation author="Li, W. H. &amp; D. Yang" pageId="2" pageNumber="150" pagination="65 - 68" refId="ref3611" refString="Li, W. H. and D. Yang. 2007. Two new species of Nemoura (Plecoptera: Nemouridae) from Henan, China. Zootaxa, 1511 (21): 65 - 68." type="journal article" year="2007">Li &amp; Yang 2007</bibRefCitation>
</taxonomicName>
(
<collectingRegion box="[325,405,1567,1591]" country="China" name="Henan" pageId="2" pageNumber="150">Henan</collectingRegion>
) share a very long rostrum of the epiproct in combination with large, dentate, claw- or hand-shaped anterolateral sclerites.
<taxonomicName authorityName="Zwick &amp; Baumann" authorityYear="2011" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="jejudoensis">
<emphasis italics="true" pageId="2" pageNumber="150">Nemoura jejudoensis</emphasis>
</taxonomicName>
possesses an anteromedial rough process above these toothed sclerites which is absent in
<taxonomicName authorityName="Wu" authorityYear="1929" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="geei">
<emphasis italics="true" pageId="2" pageNumber="150">N. geei</emphasis>
</taxonomicName>
. The shape of the transverse dorsal epiproct sclerites also differs between the two species.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760191" ID-Zenodo-Dep="4760191" httpUri="https://zenodo.org/record/4760191/files/figure.png" pageId="2" pageNumber="150" startId="2.[189,244,1212,1236]" targetBox="[217,1410,252,1156]" targetPageId="2">
<paragraph blockId="2.[189,1443,1212,1449]" pageId="2" pageNumber="150">
Figs.1-4.
<taxonomicName authority="Zwick &amp; Baumann, 2011" authorityName="Zwick &amp; Baumann" authorityYear="2011" box="[293,516,1212,1236]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="jejudoensis" status="sp. n.">
<emphasis box="[293,516,1212,1236]" italics="true" pageId="2" pageNumber="150">Nemoura jejudoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[522,584,1213,1236]" pageId="2" pageNumber="150" rank="species">sp. n.</taxonomicNameLabel>
, male. 1. Epiproct in dorsal (right half of figure) and ventral view (left half of figure), respectively. 2. Slightly oblique lateral view of male epiproct, diagrammatic, (not to scale). 3. Dorsal view of abdominal tip. 4. Anterior view of the apical epiproct sclerites.
</paragraph>
<paragraph blockId="2.[189,1443,1212,1449]" pageId="2" pageNumber="150">Note that figures 3 and 4 show a strongly contracted specimen with the epiproct pulled down and the paraprocts forward. The normally barely noticed blade-like inner paraproct lobes appear as sharp spines between the cerci and the epiproct while the obtuse pilose apex of the outer lobe is concealed between the epiproct and base of the cercus.</paragraph>
</caption>
<paragraph blockId="2.[189,798,1532,1911]" lastBlockId="2.[835,1444,1532,1911]" pageId="2" pageNumber="150">
Details of epiproct structure distinguish also the other two similar species but they differ in additional characters from
<taxonomicName authorityName="Wu" authorityYear="1929" box="[392,473,1888,1911]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="geei">
<emphasis box="[392,473,1888,1911]" italics="true" pageId="2" pageNumber="150">N. geei</emphasis>
</taxonomicName>
:
<taxonomicName authorityName="Shimizu" authorityYear="1997" box="[493,692,1887,1911]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="quadrituberata">
<emphasis box="[493,692,1887,1911]" italics="true" pageId="2" pageNumber="150">N. quadrituberata</emphasis>
</taxonomicName>
by four hairy swellings on tergites 9 and 10, toothless cerci, and broad paraprocts;
<taxonomicName authorityName="Shimizu" authorityYear="1997" box="[1153,1324,1567,1591]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis box="[1153,1324,1567,1591]" italics="true" pageId="2" pageNumber="150">N. longilobata</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Li &amp; Yang" authorityYear="2007" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="atristrigata">
<emphasis italics="true" pageId="2" pageNumber="150">N. atristrigata</emphasis>
</taxonomicName>
by a large hook on the cercus and broad, tongue-shaped paraprocts.
</paragraph>
<paragraph blockId="2.[835,1444,1532,1911]" pageId="2" pageNumber="150">
<taxonomicName authorityName="Zwick &amp; Baumann" authorityYear="2011" box="[873,1096,1674,1698]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="jejudoensis">
<emphasis box="[873,1096,1674,1698]" italics="true" pageId="2" pageNumber="150">Nemoura jejudoensis</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Wu" authorityYear="1929" box="[1246,1319,1675,1698]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="geei">
<emphasis box="[1246,1319,1675,1698]" italics="true" pageId="2" pageNumber="150">N. geei</emphasis>
</taxonomicName>
only in the epiproct. We imagine that
<taxonomicName authorityName="Zwick &amp; Baumann" authorityYear="2011" box="[1156,1310,1709,1733]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="jejudoensis">
<emphasis box="[1156,1310,1709,1733]" italics="true" pageId="2" pageNumber="150">N. jejudoensis</emphasis>
</taxonomicName>
developed fairly recently from a small founder population of
<taxonomicName authorityName="Wu" authorityYear="1929" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="150" phylum="Arthropoda" rank="species" species="geei">
<emphasis italics="true" pageId="2" pageNumber="150">N. geei</emphasis>
</taxonomicName>
that ventured to the off-shore island of
<collectingRegion box="[1345,1433,1780,1804]" country="South Korea" name="Jeju" pageId="2" pageNumber="150">Jeju-do</collectingRegion>
.
<emphasis bold="true" box="[835,973,1816,1840]" pageId="2" pageNumber="150">Etymology.</emphasis>
The specific name is a noun in apposition after
<collectingRegion box="[1041,1085,1852,1876]" country="South Korea" name="Jeju" pageId="2" pageNumber="150">Jeju</collectingRegion>
Island where the specimens were collected that represent this new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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