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<document id="C320FDBBDFAB3843444654D1DC4A94A7" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno &amp; Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF95E8190D238E5227CBFF30" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia planispira Liew &amp; Schilthuizen &amp; Vermeulen 2009, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="541" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="541" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="FB206470F04407DBB81A02001A216BDE">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:namePart id="D3FCACF84DCBD010C57CE547B7D81A93">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="F32657AD507C84E79CACA483554E450D">Schilthuizen, Menno</mods:namePart>
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<mods:namePart id="45E15A573B4F2420690E8C61286EEE98">Vermeulen, Jaap Jan</mods:namePart>
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<mods:title id="BBC51DBD16F338122E44E6C91071BEB2">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="F16AE085123012A8D2C5980316FE2F01">2009</mods:date>
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<mods:number id="270045BA40E6EE522A92EA97C4109859">2009-10-26</mods:number>
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<mods:url id="DF688CE6FDED898407C1841331B99EEA">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
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<mods:identifier id="E8F9C718B2348A7CCAFA0434C0EB5E73" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
<mods:identifier id="06D8AC624BD73F641636AB954E620E06" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="AD82C20C725DABDBCA8A35A9033476A2" type="Zenodo-Dep">5443023</mods:identifier>
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<treatment id="03F9878AFF95E8190D238E5227CBFF30" ID-DOI="http://doi.org/10.5281/zenodo.5492203" ID-Zenodo-Dep="5492203" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8190D238E5227CBFF30" httpUri="http://treatment.plazi.org/id/03F9878AFF95E8190D238E5227CBFF30" lastPageNumber="541" pageId="26" pageNumber="541">
<subSubSection id="C34A6517FF95E8190D238E5221D5FCDC" pageId="26" pageNumber="541" type="nomenclature">
<paragraph id="8BEF369CFF95E8190D238E5221E3FD9C" blockId="26.[256,646,577,598]" box="[256,646,577,598]" pageId="26" pageNumber="541">
<heading id="D0A781F0FF95E8190D238E5221E3FD9C" box="[256,646,577,598]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="541" reason="2">
<taxonomicName id="4C504D1FFF95E8190D238E522172FD9F" ID-CoL="6HM84" authority="Liew &amp; Schilthuizen &amp; Vermeulen, 2009" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[256,535,577,598]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="541" phylum="Mollusca" rank="species" species="planispira" status="sp. nov.">EVERETTIA PLANISPIRA</taxonomicName>
<emphasis id="B924EA8EFF95E8190E028E5721E3FD9C" bold="true" box="[545,646,577,598]" pageId="26" pageNumber="541">
<taxonomicNameLabel id="A21757F5FF95E8190E028E5721E3FD9C" box="[545,646,577,598]" pageId="26" pageNumber="541" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF369CFF95E8190CB38E7B21D5FCDC" blockId="26.[144,759,615,790]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB38E7B23B0FDB7" box="[144,213,616,637]" italics="true" pageId="26" pageNumber="541">Types:</emphasis>
<materialsCitation id="3B383CC1FF95E8190CC38E7421C8FCDC" collectingDate="2004-08-14" collectionCode="T" collectorName="M. Schilthuizen &amp; S. Liew &amp; A. van Til. Date" country="Malaysia" county="State" elevation="400" latitude="5.8166666" location="Ulu Tungud Forest Reserve. East of Solonsong River" longLatPrecision="1305" longitude="117.13333" municipality="Beluran district" pageId="26" pageNumber="541" specimenCode="BMNH 20080633" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="F347760CFF95E8190CC38E74220CFDB6" box="[224,361,615,636]" name="Malaysia" pageId="26" pageNumber="541">MALAYSIA</collectingCountry>
:
<collectingCounty id="628E4E10FF95E8190D548E7422D1FDB6" box="[375,436,615,636]" pageId="26" pageNumber="541">State</collectingCounty>
of
<collectingRegion id="4994F87EFF95E8190DFE8E74214DFDB7" box="[477,552,615,637]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
:
<collectingMunicipality id="6B8BACE6FF95E8190E158E742196FDB7" box="[566,755,615,637]" pageId="26" pageNumber="541">Beluran district</collectingMunicipality>
.
<location id="8E8F6047FF95E8190CB38E952197FD56" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8190D238E5227CBFF30:8E8F6047FF95E8190CB38E952197FD56" box="[144,754,646,668]" country="Malaysia" county="State" latitude="5.8166666" longLatPrecision="1305" longitude="117.13333" municipality="Beluran district" name="Ulu Tungud Forest Reserve. East of Solonsong River" pageId="26" pageNumber="541" stateProvince="Sabah">Ulu Tungud Forest Reserve. East of Solonsong River</location>
,
<quantity id="4CA89B79FF95E8190CB38EB623B3FD70" box="[144,214,677,698]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="541" unit="m" value="400.0">
<elevation id="007DD1AFFF95E8190CB38EB623B3FD70" box="[144,214,677,698]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="541" unit="m" value="400.0">400 m</elevation>
</quantity>
alt. (
<geoCoordinate id="EE64505BFF95E8190D368EB72201FD70" box="[277,356,675,698]" degrees="5" direction="north" minutes="49" orientation="latitude" pageId="26" pageNumber="541" precision="925" value="5.8166666">
5°49
<emphasis id="B924EA8EFF95E8190D6B8EB0222BFD70" box="[328,334,675,698]" italics="true" pageId="26" pageNumber="541">′</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE64505BFF95E8190D518EB622AEFD70" box="[370,459,675,698]" degrees="117" direction="east" minutes="8" orientation="longitude" pageId="26" pageNumber="541" precision="925" value="117.13333">
117°8
<emphasis id="B924EA8EFF95E8190D928EB022D2FD70" box="[433,439,675,698]" italics="true" pageId="26" pageNumber="541">′</emphasis>
E
</geoCoordinate>
). Collected by
<collectorName id="26A5534AFF95E8190E558EB623B4FD13" pageId="26" pageNumber="541">M. Schilthuizen</collectorName>
, T.-
<collectorName id="26A5534AFF95E8190D238ED02233FD13" box="[256,342,707,729]" pageId="26" pageNumber="541">S. Liew</collectorName>
, and
<collectorName id="26A5534AFF95E8190DBA8ED02138FD12" box="[409,605,707,729]" pageId="26" pageNumber="541">A. van Til. Date</collectorName>
:
<date id="FFEE105CFF95E8190E488ED02197FD13" box="[619,754,707,729]" pageId="26" pageNumber="541" value="2004-08-14">
<collectingDate id="EFAAE9B4FF95E8190E488ED02197FD13" box="[619,754,707,729]" pageId="26" pageNumber="541" value="2004-08-14">14.viii.2004</collectingDate>
</date>
,
<typeStatus id="54EB883EFF95E8190CB38EF12396FD32" box="[144,243,738,760]" pageId="26" pageNumber="541" type="holotype">holotype</typeStatus>
, BOR/MOL 3213,
<figureCitation id="136B2A19FF95E8190DE28EF12126FD3D" box="[449,579,738,760]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="26" pageNumber="541">Figure 12E</figureCitation>
;
<specimenCount id="9D56FD15FF95E8190E6C8EF12196FD3D" box="[591,755,738,759]" pageId="26" pageNumber="541" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
, one in
<specimenCode id="DBF69EE7FF95E8190CC18F1222CFFCDC" box="[226,426,768,790]" collectionCode="BMNH" pageId="26" pageNumber="541">BMNH 20080633</specimenCode>
and one in SP 13065
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6517FF95E8190CB38F5F27CBFF30" pageId="26" pageNumber="541" type="etymology">
<paragraph id="8BEF369CFF95E8190CB38F5F21ADFCA8" blockId="26.[144,712,844,866]" box="[144,712,844,866]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB38F5F22EAFCAB" box="[144,399,844,865]" italics="true" pageId="26" pageNumber="541">
Etymology:
<taxonomicName id="4C504D1FFF95E8190D348F5F22EAFCAB" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[279,399,844,865]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="541" phylum="Mollusca" rank="species" species="planispira">Planispira</taxonomicName>
</emphasis>
. Latin, means flat whorls.
</paragraph>
<paragraph id="8BEF369CFF95E8190CB38F8422A1FB0B" blockId="26.[144,759,919,1217]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB38F842217FC66" box="[144,370,919,940]" italics="true" pageId="26" pageNumber="541">Material examined:</emphasis>
<collectingRegion id="4994F87EFF95E8190D5D8F8B22B2FC67" box="[382,471,920,941]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">SABAH</collectingRegion>
– Danum Valley: BOR/ MOL 1041,
<date id="FFEE105CFF95E8190D028FA5221BFC06" box="[289,382,950,972]" pageId="26" pageNumber="541" value="2001-07">vii.2001</date>
; BOR/MOL 1132,
<date id="FFEE105CFF95E8190E7E8FA521CDFC01" box="[605,680,950,971]" pageId="26" pageNumber="541" value="2000-05">v.2000</date>
; BOR/ MOL 1139,
<date id="FFEE105CFF95E8190D638FC622EEFC20" box="[320,395,981,1002]" pageId="26" pageNumber="541" value="2000-05">v.2000</date>
; BOR/MOL 1142,
<date id="FFEE105CFF95E8190EB88FC62197FC21" box="[667,754,981,1003]" pageId="26" pageNumber="541" value="2001-03">iii.2001</date>
; BOR/MOL 1143,
<date id="FFEE105CFF95E8190D4A8FE022A5FBC3" box="[361,448,1011,1033]" pageId="26" pageNumber="541" value="2000-09">ix.2000</date>
. Tabin Wildlife Reserve: BOR/MOL 1146,
<date id="FFEE105CFF95E8190D47880122D5FBED" box="[356,432,1042,1063]" pageId="26" pageNumber="541" value="2001-05">v.2001</date>
. Kinabatangan: BOR/MOL 1308, Batu Keruak,
<date id="FFEE105CFF95E8190DA9882322B8FB8C" box="[394,477,1072,1094]" pageId="26" pageNumber="541" value="2002-04">iv.2002</date>
; BOR/MOL 1309, Batu Keruak,
<date id="FFEE105CFF95E8190CDA885C2229FBAF" box="[249,332,1103,1125]" pageId="26" pageNumber="541" value="2002-04">iv.2002</date>
; BMNH 1892.70.20.61, Gomantong, 1892. Mount Kinabalu: BOR/MOL 5358, eastern slope,
<quantity id="4CA89B79FF95E8190CFF889F224AFB68" box="[220,303,1164,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.952" pageId="26" pageNumber="541" unit="m" value="1952.0">1952 m</quantity>
alt.,
<date id="FFEE105CFF95E8190D50889F22A3FB68" box="[371,454,1164,1186]" pageId="26" pageNumber="541" value="2004-04">iv.2004</date>
; BOR/MOL 5358, eastern slope,
<quantity id="4CA89B79FF95E8190CF988B82249FB0A" box="[218,300,1195,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.1119999999999997" pageId="26" pageNumber="541" unit="m" value="2112.0">2112 m</quantity>
alt.,
<date id="FFEE105CFF95E8190D4E88B822A5FB0B" box="[365,448,1195,1217]" pageId="26" pageNumber="541" value="2004-04">iv.2004</date>
.
</paragraph>
<paragraph id="8BEF369CFF95E8190CB388E423A8FAA2" blockId="26.[144,758,1270,1384]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB388E422D9FAC1" box="[144,444,1270,1292]" italics="true" pageId="26" pageNumber="541">Diagnostic characteristics:</emphasis>
This species is easy to recognize by its small, shiny shell with flat apex and lack of sculpture on the upper and lower surface of the shell.
</paragraph>
<paragraph id="8BEF369CFF95E8190CB3898C213EF90D" blockId="26.[144,759,1438,1735]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB3898C227DFA7E" box="[144,280,1439,1460]" italics="true" pageId="26" pageNumber="541">Description:</emphasis>
Shell (
<figureCitation id="136B2A19FF95E8190DA7898D2282FA7E" box="[388,487,1438,1460]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="26" pageNumber="541">Fig. 12E</figureCitation>
): small, rather thin, brownish, spire flat to slightly elevated, outer whorls flat to slightly rounded. Periphery rounded. Above the periphery, shell shiny with faded growth lines. Below the periphery, shell is smooth and shiny. Height up to
<quantity id="4CA89B79FF95E8190CB38A242382F986" box="[144,231,1591,1612]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="26" pageNumber="541" unit="mm" value="4.7">4.7 mm</quantity>
; width up to
<quantity id="4CA89B79FF95E8190DB58A242288F986" box="[406,493,1591,1612]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="26" pageNumber="541" unit="mm" value="8.3">8.3 mm</quantity>
; diameter of the first three whorls 0.8, 0.7, and
<quantity id="4CA89B79FF95E8190DDB8A45212AF9A1" box="[504,591,1622,1643]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="541" unit="mm" value="0.8">0.8 mm</quantity>
, respectively; number of whorls up to four and one-quarter; height aperture up to
<quantity id="4CA89B79FF95E8190D1D8A8022F0F962" box="[318,405,1683,1705]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="541" unit="mm" value="3.5">3.5 mm</quantity>
; width aperture up to
<quantity id="4CA89B79FF95E8190EB88A802196F962" box="[667,755,1683,1704]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="26" pageNumber="541" unit="mm" value="4.8">4.8 mm</quantity>
. Genitalia: unknown. Animal: unknown.
</paragraph>
<paragraph id="8BEF369CFF95E8190CB38AEE22CDF8A5" blockId="26.[144,759,1789,1903]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190CB38AEE22D1F8D8" box="[144,436,1789,1810]" italics="true" pageId="26" pageNumber="541">Distribution and habitat:</emphasis>
Primary forest
<quantity id="4CA89B79FF95E8190E508AEE2197F8D8" box="[627,754,1789,1811]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="2.0" metricValueMin="0.05" pageId="26" pageNumber="541" unit="m" value="1025.0" valueMax="2000.0" valueMin="50.0">50–2000 m</quantity>
, but becoming very rare above
<specimenCount id="9D56FD15FF95E8190DCC8B082159F8FB" box="[495,572,1819,1841]" pageId="26" pageNumber="541" type="generic">
<quantity id="4CA89B79FF95E8190DCC8B08215DF8FB" box="[495,568,1819,1841]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="541" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
<collectingRegion id="4994F87EFF95E8190E678B0F21EAF8F8" box="[580,655,1820,1842]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
: Kinabatangan, Tabin, Danum valley, and Tawau (east coast of
<collectingRegion id="4994F87EFF95E8190C8D8B4A2393F8A5" box="[174,246,1881,1903]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
) (
<figureCitation id="136B2A19FF95E8190D2C8B4A220EF8A5" box="[271,363,1881,1903]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="26" pageNumber="541">Figs 5D</figureCitation>
,
<figureCitation id="136B2A19FF95E8190D5A8B4A22FEF8A5" box="[377,411,1881,1903]" captionStart="Figure 9" captionStartId="15.[164,243,1622,1641]" captionTargetBox="[325,1281,196,1590]" captionTargetId="figure-58@15.[323,1283,194,1592]" captionTargetPageId="15" captionText="Figure 9. Distribution maps (Mount Kinabalu and Mount Tambuyukon). A, Everettia monticola sp. nov. and Everettia safriei sp. nov.; B, Everettia planispira sp. nov., Everettia jasilini sp. nov., and Everettia corrugata corrugata; C, Everettia dominiki sp. nov.; D, Everettia themis, Everettia subconsul, Everettia corrugata williamsi ssp. nov. and Everettia layanglayang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443041" httpUri="https://zenodo.org/record/5443041/files/figure.png" pageId="26" pageNumber="541">9B</figureCitation>
).
</paragraph>
<paragraph id="8BEF369CFF95E8190F058CD527CBFF30" blockId="26.[806,1421,198,250]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190F058CD520F7FF11" box="[806,914,198,219]" italics="true" pageId="26" pageNumber="541">Remarks:</emphasis>
This species has a shiny shell and is mainly found on the east coast of
<collectingRegion id="4994F87EFF95E81908438CF727CEFF30" box="[1120,1195,228,250]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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