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<document id="1AA20ABEB3EE8539C345ADE8D669E88E" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF21FF51FDEBBE48A244FDC4" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus fidenas Noyes 2023, sp.nov." docType="treatment" docUuid="4C6FB785-5062-4F1D-AF89-CC40098086E6" docUuidSource="ZooBank" docVersion="2" lastPageNumber="245" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="243" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="5D1E0F9EF8D64188F8CC6459CBD9B0FD">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="045FFBD102D6190FF28846C2684CAF3E">Noyes, John Stuart</mods:namePart>
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<mods:title id="71CA46C2D28B2F931ED8D4579D423306">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="F23D795B1BF2BF104200B0299884D8D9">2023</mods:date>
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<mods:number id="5F3EDB6870E8D2AE248877773F390C9E">2023-06-30</mods:number>
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<subSubSection id="C309653AFF21FF53FDEBBE48A529FD40" box="[521,825,620,645]" pageId="244" pageNumber="243" type="nomenclature">
<paragraph id="8BAC36B1FF21FF53FDEBBE48A529FD40" blockId="244.[521,825,620,671]" box="[521,825,620,645]" pageId="244" pageNumber="243">
<heading id="D0E481DDFF21FF53FDEBBE48A529FD40" bold="true" box="[521,825,620,645]" centered="true" fontSize="10" level="2" pageId="244" pageNumber="243" reason="2">
<emphasis id="B967EAA3FF21FF53FDEBBE48A529FD40" bold="true" box="[521,825,620,645]" pageId="244" pageNumber="243">
<taxonomicName id="4C134D32FF21FF53FDEBBE48A4F8FD41" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[521,744,620,644]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="244" pageNumber="243" phylum="Arthropoda" rank="species" species="fidenas" status="sp.nov.">
<emphasis id="B967EAA3FF21FF53FDEBBE48A4F8FD41" bold="true" box="[521,744,620,644]" italics="true" pageId="244" pageNumber="243">Cheiloneurus fidenas</emphasis>
</taxonomicName>
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</emphasis>
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</paragraph>
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<subSubSection id="C309653AFF21FF53FDFBBEA3A538FD5A" box="[537,808,647,671]" pageId="244" pageNumber="243" type="description">
<paragraph id="8BAC36B1FF21FF53FDFBBEA3A538FD5A" blockId="244.[521,825,620,671]" box="[537,808,647,671]" pageId="244" pageNumber="243">
(
<figureCitation id="13282A34FF21FF53FDC2BEA3A4BBFD5A" box="[544,683,647,671]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Figs 604-607</figureCitation>
; Hab. E 92)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF21FF51FF57BE99A29CFF5A" lastPageId="246" lastPageNumber="245" pageId="244" pageNumber="243" type="diagnosis">
<paragraph id="8BAC36B1FF21FF53FF57BE99A45AFAAA" blockId="244.[181,1165,701,1391]" pageId="244" pageNumber="243">
DIAGNOSIS.
<emphasis id="B967EAA3FF21FF53FEADBE99A78BFD10" box="[335,411,701,725]" italics="true" pageId="244" pageNumber="243">Female</emphasis>
(length about
<quantity id="4CEB9B54FF21FF53FDD8BE99A4BEFD10" box="[570,686,701,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="244" pageNumber="243" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6-1.9mm</quantity>
): body generally dark brown to black with a weak to moderate metallic sheen; antenna (
<figureCitation id="13282A34FF21FF53FD99BEF3A4C4FD2A" box="[635,724,727,751]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 605</figureCitation>
) with scape pale orange margined brown, proximally brown; pedicel mostly pale orange, proximally dark brown dorsally; funicle pale orange, proximally slightly dusky; clava dark brown; mesoscutum with uniform pale brown setae that appear silvery in some lights; coxae white; hind tibia mostly dark brown, legs otherwise white to pale orange with brown areas; fore wing (
<figureCitation id="13282A34FF21FF53FD4FBF66A512FC9F" box="[685,770,834,858]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 604</figureCitation>
) almost completely hyaline but weakly infuscate immediately below marginal and stigmal veins and an infuscate median streak in apical half; head about 4.1X as wide as frontovertex, in facial view about as broad as long, subcircular, genae converging, moderately and evenly curved; frontovertex naked medially below anterior ocellus; area between eye and scrobe about 1.5X as wide as diameter of anterior ocellus, with shallow sculpture, scrobal margin rounded; interantennal prominence dorsally rounded with only a few long setae; mandible with two long, sharp lower teeth and a slightly shorter, sharp, upper tooth; antenna (
<figureCitation id="13282A34FF21FF53FE7BBFD9A7E1FBD0" box="[409,497,1021,1045]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 605</figureCitation>
) with scape about 4.2X as long as broad; funicle with segments quadrate or slightly longer than broad, becoming larger distad; head width about equal to length of flagellum; linear sensilla on F3-F6; clava 3-segmented, slightly shorter than F2-F6 combined, sensory area large and forming an oblique truncation, extending about 0.4X along ventral margin; mesoscutum (
<figureCitation id="13282A34FF21FF53FEA1B843A789FBBA" box="[323,409,1127,1151]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 607</figureCitation>
) with polygonally reticulate sculpture; scutellum with a distinct, but short apical tuft of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FF21FF53FD11B8A6A55CFB5F" box="[755,844,1154,1178]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 604</figureCitation>
) about 2.5X as long as broad; parastigma very weakly curved downwards; costal cell dorsally with 5 to 7 setae at apex and a single, complete line of setae ventrally; area below proximal part of parastigma with a group of about 18 setae; apices of postmarginal and stigmal veins (
<figureCitation id="13282A34FF21FF53FCF4B8F6A57FFB2F" box="[790,879,1234,1258]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 603</figureCitation>
) not connected by a naked hyaline area; apical bristle of postmarginal vein about 0.5X as long as marginal vein; mid tibial spur about as long as basitarsus; propodeum (
<figureCitation id="13282A34FF21FF53FD67B923A4CBFADA" box="[645,731,1287,1311]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 607</figureCitation>
) with about 20 setae around each spiracle, side naked; syntergum 0.9X as long as mid tibia; ovipositor about 3.3X gonostylus or 1.3X mid tibia; gonostylus about 1.2X as long as mid tibial spur; exserted part of ovipositor about 0.3X as long as mid tibial spur.
<emphasis id="B967EAA3FF21FF53FE41B973A7C7FAAA" box="[419,471,1367,1391]" italics="true" pageId="244" pageNumber="243">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF21FF53FF57B9A9A293FA60" blockId="244.[181,1165,1421,1685]" box="[181,1155,1421,1445]" pageId="244" pageNumber="243">
Female (
<typeStatus id="54A88813FF21FF53FEECB9A9A77DFA60" box="[270,365,1421,1445]" pageId="244" pageNumber="243">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FF21FF53FD40B9A9A4E9FA60" box="[674,761,1421,1445]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" pageId="244" pageNumber="243" unit="mm" value="1.71">1.71mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF21FF53FC3DB9A9A223FA60" box="[991,1075,1421,1445]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="244" pageNumber="243" unit="mm" value="1.66">1.66mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF21FF53FF39B983A29CF950" blockId="244.[181,1165,1421,1685]" pageId="244" pageNumber="243">
Head black with a weak blue and purple sheen on frontovertex, more strongly purple anteriorly; scrobal area largely purple with scrobes and interantennal prominence largely metallic green with a purple band connecting toruli; mouth margin purplish; area below eye strongly metallic brassy, green and coppery; temple brassy and coppery; gena purple and brassy; frontovertex with inconspicuous dark brown setae; antenna (
<figureCitation id="13282A34FF21FF53FDB9BA36A4BFF9EF" box="[603,687,1554,1578]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="244" pageNumber="243">Fig. 605</figureCitation>
) with radicle dark brown, pale orange ventrally; antenna with scape mostly pale orange, proximally brown, dorsal and ventral margins brown; pedicel mostly pale orange, basal two-thirds dark brown dorsally; funicle pale orange, F1-F3 very slightly dusky; clava dark brown; thorax mostly dark brown to black; pronotum mainly dull, dark green mixed purple, posterior margin coppery; mesoscutum with a metallic green to blue-green
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF57BCA6A6F3FF5A" blockId="245.[181,227,130,159]" box="[181,227,130,159]" pageId="245" pageNumber="244">
<pageNumber id="BBB83D67FF20FF52FF57BCA6A6F3FF5A" ambiguity="2" box="[181,227,130,159]" fuzzyness="0" pageId="245" pageNumber="244" score="12" value="244">
<pageTitle id="CB8CEED6FF20FF52FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="245" pageNumber="244">244</pageTitle>
</pageNumber>
</paragraph>
<paragraph id="8BAC36B1FF20FF52FEB5BCA4A5FBFF5D" blockId="245.[343,1003,128,152]" box="[343,1003,128,152]" pageId="245" pageNumber="244">
<pageTitle id="CB8CEED6FF20FF52FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="245" pageNumber="244">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF20FF52FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="245" pageNumber="244" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF57BCFAA50CFD12" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">
sheen with coppery and brassy reflections, posterior margin coppery purple, setae uniform and translucent pale brown and appear silvery in some lights; tegula dark brown; axilla dull, dark purple-brown with coppery reflections; scutellum noticeably less shiny than mesoscutum, but with a distinct dark blue sheen, medially strongly coppery, clothed with dark brown setae, extreme sides and apex shining green mixed coppery, especially at apex; metanotum with slight purple sheen; mesopleuron in posterior half with a coppery-purple and brassy sheen, anterior half mixed blue, green, coppery and brassy, these narrowly separated by a coppery, dark blue and violet sheen; fore coxa white; fore femur white to pale yellow, with an elongate brown spot dorsally in distal half; fore tibia pale orange with an incomplete subbasal brown band; mid coxa white; mid femur white to pale yellow with an elongate brown spot externally in distal half, tibia pale orange with a narrow, elongate, brown, subbasal spot; hind coxa white; hind femur white in proximal half or so, otherwise dark brown, tibia mostly pale orange with a broad, diffuse brown, subbasal ring in proximal half or so; fore tarsus pale orange-brown, mid and hind tarsi pale orange, pretarsi slightly dusky; fore wing (
<figureCitation id="13282A34FF20FF52FE23BE1DA407FD94" box="[449,535,569,593]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="245" pageNumber="244">Fig. 604</figureCitation>
) almost completely hyaline but weakly infuscate immediately below marginal and stigmal veins, and a weakly infuscate median streak distad of pale brown venation; propodeum dark purple-brown; sides metallic green with a few conspicuous silvery setae in spiracular area; gaster dark purple-brown, dorsum slightly coppery, mixed purple and brassy, sides towards apex and outer plates of ovipositor mixed metallic blue, green, coppery and brassy; gonostylus dark brown, extreme apex orange-brown.
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF39BEFDA4B2FB89" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">
Head about 4.1X as wide as frontovertex, not subtriangular in profile, occipital margin carinate, occiput without a median groove above foramen; ocelli forming an angle of about 55°; frontovertex slightly shiny, with conspicuous reticulate sculpture of mesh size generally very slightly smaller than diameter of eye facet; scrobal area and mouth margin with shallower sculpture; gena and temple with more elongate sculpture of larger mesh; scrobes shallow, meeting dorsally, dorsal and lateral margins weakly rounded; antenna as in
<figureCitation id="13282A34FF20FF52FC4EBF7BA213FCB3" box="[940,1027,862,887]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="245" pageNumber="244">Fig. 605</figureCitation>
; scape hardly broadened and flattened, about 4.2X as long as broad; F1 and F4-F6 each a little longer than broad, F2-F3 quadrate, segments broadest distad; clava hardly broadened, slightly longer than F3- F6 combined; only outer suture noticeably oblique, sensory area enlarged, extending about one-third along clava ventrally and giving it an obliquely truncate appearance; eye reaching occipital margin, clothed with inconspicuous setae, each about as long as diameter of facet; inner eye margins not converging anteriorly; malar sulcus absent; mandible with three acute teeth, upper tooth shortest. Relative measurements: HW 70,
<collectingRegion id="49D7F853FF20FF52FD75B83EA4ACFBF4" box="[663,700,1050,1073]" country="Swaziland" name="Hhohho" pageId="245" pageNumber="244">HH</collectingRegion>
69, FV 17, OD 5.5, POL 10.5, OOL 0.5, OCL 6, AOL 11, EL 45, EW 36, MS 29, SL 38, SW 9.
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF39B86BA403FA49" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">
Thorax (
<figureCitation id="13282A34FF20FF52FEDAB86BA783FBA2" box="[312,403,1103,1127]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="245" pageNumber="244">Fig. 607</figureCitation>
) with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size subequal to diameter of facet; sculpture on mesoscutum generally of similar depth to that on pronotum, but more regular and of smaller mesh; sculpture on axilla similar to that on mesoscutum, but slightly deeper; scutellum with conspicuously deeper, polygonally reticulate sculpture; apex and sides completely smooth and shiny; visible part of mesoscutum about 1.8X as broad as long; scutellum very slightly broader than long, with a short subapical tuft, setae about one-third to two-fifths as long as scutellum; hind femur about 4.4X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF20FF52FDE0B92DA482FAE4" box="[514,658,1289,1313]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="245" pageNumber="244">Figs 604, 606</figureCitation>
; costal cell with only 3 setae dorsally at apex; no naked streak present between apices of postmarginal and stigmal veins; propodeum (
<figureCitation id="13282A34FF20FF52FBCEB900A295FAF9" box="[1068,1157,1316,1340]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta – (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT – (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres – (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas – (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="245" pageNumber="244">Fig. 607</figureCitation>
) with moderately dense setae in spiracular area, side naked, medially less than 0.1X as long as scutellum and with some shallow polygonally reticulate sculpture. Relative measurements: FWL 89, FWW 36; HWL 70, HWW 18.
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF39B9ABA731FA19" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">Gaster with hypopygium reaching about half-way to apex; syntergum nearly as long as mid tibia, with apex acute; ovipositor slightly exserted, the exserted part about one-third as long as mid tibial spur.</paragraph>
<paragraph id="8BAC36B1FF20FF52FF39B9FBA5A5F9D7" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">
<typeStatus id="54A88813FF20FF52FF39B9FBA727FA32" box="[219,311,1503,1527]" pageId="245" pageNumber="244">Paratype</typeStatus>
. Funicle with linear sensilla present only on F3-F6; gaster without “gland-like” structures on Gt1 or Gt5. Relative measurements: OL 55, GL 16.5 [MT 41].
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF39BA30A4ADF982" blockId="245.[181,1165,222,1607]" pageId="245" pageNumber="244">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FF20FF52FD3BBA30A574F9E9" box="[729,868,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="1.71" metricValueMax="1.87" metricValueMin="1.55" pageId="245" pageNumber="244" unit="mm" value="1.71" valueMax="1.87" valueMin="1.55">1.55-1.87mm</quantity>
, otherwise little in material examined except that F1-F4 may be orange-brown.
</paragraph>
<paragraph id="8BAC36B1FF20FF52FF57BA40A771F9B9" blockId="245.[181,353,1636,1660]" box="[181,353,1636,1660]" pageId="245" pageNumber="244">Male. Unknown.</paragraph>
<paragraph id="8BAC36B1FF23FF51FBBCBCA6A29CFF5A" blockId="246.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="246" pageNumber="245">245</paragraph>
</subSubSection>
<subSubSection id="C309653AFF23FF51FF57BCFBA7F9FF33" box="[181,489,222,247]" pageId="246" pageNumber="245" type="distribution">
<paragraph id="8BAC36B1FF23FF51FF57BCFBA7F9FF33" blockId="246.[181,489,222,247]" box="[181,489,222,247]" pageId="246" pageNumber="245">
DISTRIBUTION.
<collectingCountry id="F3047621FF23FF51FE91BCFBA7F5FF33" box="[371,485,222,247]" name="Costa Rica" pageId="246" pageNumber="245">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF23FF51FF57BD30A76EFEE9" box="[181,382,276,300]" pageId="246" pageNumber="245" type="biology_ecology">
<paragraph id="8BAC36B1FF23FF51FF57BD30A76EFEE9" blockId="246.[181,382,276,300]" box="[181,382,276,300]" pageId="246" pageNumber="245">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF23FF51FF57BD6FA44FFE0E" pageId="246" pageNumber="245" type="materials_examined">
<paragraph id="8BAC36B1FF23FF51FF57BD6FA7BDFEA5" blockId="246.[181,1165,331,459]" box="[181,429,331,352]" pageId="246" pageNumber="245">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF23FF51FF39BD42A44FFE0E" blockId="246.[181,1165,331,459]" pageId="246" pageNumber="245">
Type material.
<materialsCitation id="3B7B3CECFF23FF51FE86BD42A299FEBE" box="[356,1161,357,380]" collectingDate="1992-04" collectorName="de Moravia &amp; P. Hanson" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" pageId="246" pageNumber="245" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="54A88813FF23FF51FE86BD42A7A8FEB9" box="[356,440,358,380]" pageId="246" pageNumber="245">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF23FF51FE3EBD41A477FEBE" bold="true" box="[476,615,357,379]" pageId="246" pageNumber="245">
<collectingCountry id="F3047621FF23FF51FE3EBD41A477FEBE" box="[476,615,357,379]" name="Costa Rica" pageId="246" pageNumber="245">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF23FF51FD90BD41A4D7FEBE" bold="true" box="[626,711,357,379]" pageId="246" pageNumber="245">
<collectingRegion id="49D7F853FF23FF51FD90BD41A4D7FEBE" box="[626,711,357,379]" country="Costa Rica" name="San Jose" pageId="246" pageNumber="245">San José</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF23FF51FD30BD42A590FEB9" LSID="urn:lsid:plazi:treatment:03BA87A7FF21FF51FDEBBE48A244FDC4:8ECC606AFF23FF51FD30BD42A590FEB9" box="[722,896,358,380]" country="Costa Rica" name="Zurqui de Moravia" pageId="246" pageNumber="245" stateProvince="San Jose">
Zurqui
<collectorName id="26E65367FF23FF51FCF7BD42A590FEB9" box="[789,896,358,380]" pageId="246" pageNumber="245">de Moravia</collectorName>
</location>
,
<quantity id="4CEB9B54FF23FF51FC6BBD41A5DAFEBE" box="[905,970,357,379]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="246" pageNumber="245" unit="m" value="1600.0">
<elevation id="003ED182FF23FF51FC6BBD41A5DAFEBE" box="[905,970,357,379]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="246" pageNumber="245" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFAD1071FF23FF51FC31BD42A206FEB9" box="[979,1046,358,380]" pageId="246" pageNumber="245" value="1992-04">
<collectingDate id="EFE9E999FF23FF51FC31BD42A206FEB9" box="[979,1046,358,380]" pageId="246" pageNumber="245" value="1992-04">iv.1992</collectingDate>
</date>
(
<collectorName id="26E65367FF23FF51FBC1BD42A293FEBE" box="[1059,1155,358,379]" pageId="246" pageNumber="245">P. Hanson</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF23FF51FF57BDA5A5F5FE53" box="[181,997,383,406]" collectingDate="1992-04" collectorName="de Moravia &amp; P. Hanson" country="Costa Rica" elevation="1600" location="Zurqui de Moravia" pageId="246" pageNumber="245" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
<typeStatus id="54A88813FF23FF51FF57BDA5A701FE53" box="[181,273,385,406]" pageId="246" pageNumber="245">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF23FF51FEFBBDA4A7B3FE53" bold="true" box="[281,419,384,406]" pageId="246" pageNumber="245">
<collectingCountry id="F3047621FF23FF51FEFBBDA4A7B3FE53" box="[281,419,384,406]" name="Costa Rica" pageId="246" pageNumber="245">COSTA RICA</collectingCountry>
</emphasis>
, 6E,
<emphasis id="B967EAA3FF23FF51FE34BDA4A43AFE50" bold="true" box="[470,554,383,406]" pageId="246" pageNumber="245">
<collectingRegion id="49D7F853FF23FF51FE34BDA4A43AFE50" box="[470,554,383,406]" country="Costa Rica" name="San Jose" pageId="246" pageNumber="245">San José</collectingRegion>
</emphasis>
, same data as holotype but various dates
<date id="FFAD1071FF23FF51FC7CBDA4A5F5FE53" box="[926,997,384,406]" pageId="246" pageNumber="245" value="1992-04">
<collectingDate id="EFE9E999FF23FF51FC7CBDA4A5F5FE53" box="[926,997,384,406]" pageId="246" pageNumber="245" value="1992-04">iv.1992</collectingDate>
</date>
</materialsCitation>
-
<materialsCitation id="3B7B3CECFF23FF51FC09BDA4A235FE53" box="[1003,1061,384,406]" collectingDate="1996-01" collectorName="de Moravia &amp; P. Hanson" country="Costa Rica" elevation="1600" isTailEnumeration="true" location="Zurqui de Moravia" pageId="246" pageNumber="245" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
<date id="FFAD1071FF23FF51FC09BDA4A235FE53" box="[1003,1061,384,406]" pageId="246" pageNumber="245" value="1996-01">
<collectingDate id="EFE9E999FF23FF51FC09BDA4A235FE53" box="[1003,1061,384,406]" pageId="246" pageNumber="245" value="1996-01">i.1996</collectingDate>
</date>
</materialsCitation>
; 1E,
<materialsCitation id="3B7B3CECFF23FF51FBB7BDA4A543FE74" collectingDate="2002-02" country="Costa Rica" elevation="2000" latitude="10.183333" location="6 km ENE Vara Blanca" longLatPrecision="1298" longitude="-84.01667" pageId="246" pageNumber="245" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="B967EAA3FF23FF51FBB7BDA4A6C3FE74" bold="true" pageId="246" pageNumber="245">
<collectingRegion id="49D7F853FF23FF51FBB7BDA4A6C3FE74" country="Costa Rica" name="Heredia" pageId="246" pageNumber="245">Heredia</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF23FF51FF3DBDBFA7BFFE74" LSID="urn:lsid:plazi:treatment:03BA87A7FF21FF51FDEBBE48A244FDC4:8ECC606AFF23FF51FF3DBDBFA7BFFE74" box="[223,431,411,433]" country="Costa Rica" latitude="10.183333" longLatPrecision="1298" longitude="-84.01667" name="6 km ENE Vara Blanca" pageId="246" pageNumber="245" stateProvince="Heredia">
<quantity id="4CEB9B54FF23FF51FF3DBDBFA717FE74" box="[223,263,411,433]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="246" pageNumber="245" unit="km" value="6.0">6km</quantity>
ENE Vara Blanca
</location>
,
<geoCoordinate id="EE275076FF23FF51FE5ABDBFA415FE75" box="[440,517,411,432]" degrees="10" direction="north" minutes="11" orientation="latitude" pageId="246" pageNumber="245" precision="925" value="10.183333">10°11’N</geoCoordinate>
<geoCoordinate id="EE275076FF23FF51FDE9BDBFA44FFE75" box="[523,607,411,432]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="246" pageNumber="245" precision="925" value="-84.01667">84°01’W</geoCoordinate>
,
<quantity id="4CEB9B54FF23FF51FD8ABDBFA4B9FE75" box="[616,681,411,432]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="246" pageNumber="245" unit="m" value="2000.0">
<elevation id="003ED182FF23FF51FD8ABDBFA4B9FE75" box="[616,681,411,432]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="246" pageNumber="245" unit="m" value="2000.0">2000m</elevation>
</quantity>
, 20/M/TN,
<date id="FFAD1071FF23FF51FCF6BDBFA543FE74" box="[788,851,411,433]" pageId="246" pageNumber="245" value="2002-02">
<collectingDate id="EFE9E999FF23FF51FCF6BDBFA543FE74" box="[788,851,411,433]" pageId="246" pageNumber="245" value="2002-02">ii.2002</collectingDate>
</date>
</materialsCitation>
(INBio-OET-ALAS).
<materialsCitation id="3B7B3CECFF23FF51FBC2BDBFA44BFE0E" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="246" pageNumber="245" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="54A88813FF23FF51FBC2BDBFA264FE74" box="[1056,1140,411,433]" pageId="246" pageNumber="245">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF23FF51FF57BD92A718FE0E" box="[181,264,438,459]" pageId="246" pageNumber="245">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF23FF51FEF3BD92A778FE0E" box="[273,360,438,459]" pageId="246" pageNumber="245">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FF23FF51FE67BD92A7CBFE0E" box="[389,475,438,459]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="246" pageNumber="245" type="Museum">NHMUK</collectionCode>
and
<collectionCode id="ED02AE74FF23FF51FDE5BD92A44BFE0E" box="[519,603,438,459]" pageId="246" pageNumber="245">MZUCR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF23FF51FF57BDCDA244FDC4" box="[181,1108,489,513]" pageId="246" pageNumber="245" type="discussion">
<paragraph id="8BAC36B1FF23FF51FF57BDCDA244FDC4" blockId="246.[181,1108,489,513]" box="[181,1108,489,513]" pageId="246" pageNumber="245">
COMMENTS. See comments under
<taxonomicName id="4C134D32FF23FF51FDC9BDCDA483FDC4" authorityName="Noyes" authorityYear="2023" box="[555,659,489,513]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="246" pageNumber="245" phylum="Arthropoda" rank="species" species="herndonia">
<emphasis id="B967EAA3FF23FF51FDC9BDCDA483FDC4" box="[555,659,489,513]" italics="true" pageId="246" pageNumber="245">herndonia</emphasis>
</taxonomicName>
(p. 240),
<taxonomicName id="4C134D32FF23FF51FD14BDCEA526FDC4" authorityName="Noyes" authorityYear="2023" box="[758,822,490,513]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="246" pageNumber="245" phylum="Arthropoda" rank="species" species="exeres">
<emphasis id="B967EAA3FF23FF51FD14BDCEA526FDC4" box="[758,822,490,513]" italics="true" pageId="246" pageNumber="245">exeres</emphasis>
</taxonomicName>
(p. 243) and
<taxonomicName id="4C134D32FF23FF51FC5CBDCDA5E6FDC4" authorityName="Noyes" authorityYear="2023" box="[958,1014,489,513]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="246" pageNumber="245" phylum="Arthropoda" rank="species" species="miklo">
<emphasis id="B967EAA3FF23FF51FC5CBDCDA5E6FDC4" box="[958,1014,489,513]" italics="true" pageId="246" pageNumber="245">miklo</emphasis>
</taxonomicName>
(p. 247).
</paragraph>
</subSubSection>
</treatment>
</document>
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