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<document id="5DC029CD648EC3DD783004BD18E2DF04" ID-DOI="10.11646/zootaxa.1745.1.3" ID-ISSN="1175­5334" ID-Zenodo-Dep="5108342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626382332205" checkinUser="felipe" docAuthor="Sabu, Thomas K., Abhitha, Prabhakar &amp; Zhao, Danyang" docDate="2008" docId="03B97607FFFD647A47D62A2DFA03FCB3" docLanguage="en" docName="zt01745p046.pdf" docOrigin="Zootaxa 1745 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Physocrotaphini Chaudoir 1863" docType="key" docVersion="3" lastPageNumber="33" masterDocId="FF800E7FFFFF647947412D70FF96FF81" masterDocTitle="A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)" masterLastPageNumber="46" masterPageNumber="30" pageNumber="32" updateTime="1699011263822" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A13E96121063874AA6678FAC757C6B5D">A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)</mods:title>
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<mods:namePart id="CF6CD5E7825378A937E3F328DB17E375">Sabu, Thomas K.</mods:namePart>
<mods:affiliation id="22A6BC36ABECF18E7D6C873B0BF0E743">Litter Entomology Research Unit, P. G. &amp; Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources &amp; Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:namePart id="58F1BDFCD72BFE28807C7431AE9DBF7F">Abhitha, Prabhakar</mods:namePart>
<mods:affiliation id="CB06B6221ABBFE84E104D94F23708496">Litter Entomology Research Unit, P. G. &amp; Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources &amp; Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:namePart id="0097F9C9262000B39B22F18BB81F2631">Zhao, Danyang</mods:namePart>
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Key to the genera of the tribe 
<taxonomicName id="4C10BC92FFFD647B454D2A2DFD4FF8F6" ID-CoL="8MHW3" authorityName="Chaudoir" authorityYear="1863" box="[524,729,1885,1911]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="tribe" tribe="Physocrotaphini">Physocrotaphini</taxonomicName>
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1. Ligula prominent and distinctly longer than paraglossae (
<figureCitation id="132BDB94FFFD647B442B2AD8FC25F843" box="[874,947,1960,1986]" captionStart="FIGURES 2–5" captionStartId="3.[151,269,1179,1203]" captionTargetBox="[168,1421,876,1146]" captionTargetId="figure-497@3.[151,1436,870,1165]" captionTargetPageId="3" captionText="FIGURES 2–5. Mentum (ventral view): 2 Helluodes devagiriensis n.sp.; 3 Physocrotaphus ceylonicus; 4 Pogonoglossus giganteus; 5 Schuelea drumonti. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5108348" httpUri="https://zenodo.org/record/5108348/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
); post-orbital surface area (gena) densely setose, with more than 40 setae and without tubercles; maxillary palpomere 4 shorter than 3; TLA = 24.5– 27.5 (
<figureCitation id="132BDB94FFFC647A464B2DE8FEC9FF33" box="[266,351,152,178]" captionStart="FIGURES 23–26" captionStartId="10.[151,269,1306,1330]" captionTargetBox="[159,1424,770,1265]" captionTargetId="figure-264@10.[151,1436,764,1282]" captionTargetPageId="10" captionText="FIGURES 23–26. Habitus: 23 Helluodes devagiriensis n.sp.(Holotype); 24 Pogonoglossus giganteus (Holotype); 25 Schuelea drumonti (Holotype); 26 Physocrotaphus ceylonicus (Lectotype). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5108360" httpUri="https://zenodo.org/record/5108360/files/figure.png" pageId="3" pageNumber="33">Fig. 23</figureCitation>
) ............................................................................................................. 
<taxonomicName id="4C10BC92FFFC647A43D52DE8FA03FF33" ID-CoL="8KX6Q" authority="Westwood." authorityName="Westwood." box="[1172,1429,152,178]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B9641B03FFFC647A43D52DE8FA9DFF30" box="[1172,1291,152,177]" italics="true" pageId="3" pageNumber="33">Helluodes</emphasis>
Westwood.
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<paragraph id="8BAFC711FFFC647A47D62DB0FA03FEAB" blockId="3.[151,1436,152,818]" pageId="3" pageNumber="33">
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- Ligula truncate and distinctly shorter than paraglossae (
<figureCitation id="132BDB94FFFC647A44C22DB0FC6AFF5B" box="[899,1020,192,218]" captionStart="FIGURES 2–5" captionStartId="3.[151,269,1179,1203]" captionTargetBox="[168,1421,876,1146]" captionTargetId="figure-497@3.[151,1436,870,1165]" captionTargetPageId="3" captionText="FIGURES 2–5. Mentum (ventral view): 2 Helluodes devagiriensis n.sp.; 3 Physocrotaphus ceylonicus; 4 Pogonoglossus giganteus; 5 Schuelea drumonti. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5108348" httpUri="https://zenodo.org/record/5108348/files/figure.png" pageId="3" pageNumber="33">Figs. 3–5</figureCitation>
); post-orbital surface area (gena) sparsely setose with less than 10 setae and with or without tubercles; maxillary palpomere 4 longer than 3; TLA = 19.5 ................................................................................................................................................ 2
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2. Mentum with bifid median tooth; ligula stout and of uniform width from base to apex (
<figureCitation id="132BDB94FFFC647A43FA2C48FA94FED3" box="[1211,1282,312,338]" captionStart="FIGURES 2–5" captionStartId="3.[151,269,1179,1203]" captionTargetBox="[168,1421,876,1146]" captionTargetId="figure-497@3.[151,1436,870,1165]" captionTargetPageId="3" captionText="FIGURES 2–5. Mentum (ventral view): 2 Helluodes devagiriensis n.sp.; 3 Physocrotaphus ceylonicus; 4 Pogonoglossus giganteus; 5 Schuelea drumonti. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5108348" httpUri="https://zenodo.org/record/5108348/files/figure.png" pageId="3" pageNumber="33">Fig. 3</figureCitation>
); male sternite 3 (
<figureCitation id="132BDB94FFFC647A464B2C10FEC9FEFB" box="[266,351,352,378]" captionStart="FIGURES 15–18" captionStartId="8.[151,269,1462,1486]" captionTargetBox="[201,1408,184,1426]" captionTargetId="figure-234@8.[151,1436,160,1438]" captionTargetPageId="8" captionText="FIGURES 15–18. Median depression and cluster of filamentous setae on 2nd and 3rd sternites of male: 15 Physocrotaphus ceylonicus; 16 Pogonoglossus giganteus; 17 Helluodes devagiriensis n.sp.; 18 2nd and 3rd sternites of female Helluodes devagiriensis n.sp. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108354" httpUri="https://zenodo.org/record/5108354/files/figure.png" pageId="3" pageNumber="33">Fig. 15</figureCitation>
) with single oval depression bearing filamentous elongated setae; TLA = 19–19.5 (
<figureCitation id="132BDB94FFFC647A42732C10FA11FEFB" box="[1330,1415,352,378]" captionStart="FIGURES 23–26" captionStartId="10.[151,269,1306,1330]" captionTargetBox="[159,1424,770,1265]" captionTargetId="figure-264@10.[151,1436,764,1282]" captionTargetPageId="10" captionText="FIGURES 23–26. Habitus: 23 Helluodes devagiriensis n.sp.(Holotype); 24 Pogonoglossus giganteus (Holotype); 25 Schuelea drumonti (Holotype); 26 Physocrotaphus ceylonicus (Lectotype). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5108360" httpUri="https://zenodo.org/record/5108360/files/figure.png" pageId="3" pageNumber="33">Fig. 26</figureCitation>
) 
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<taxonomicName id="4C10BC92FFFC647A43D32CF8FA03FE23" ID-CoL="8KYF5" authority="Parry." authorityName="Parry." box="[1170,1429,392,418]" class="Insecta" family="Carabidae" genus="Physocrotaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B9641B03FFFC647A43D32CF8FAD1FE20" box="[1170,1351,392,417]" italics="true" pageId="3" pageNumber="33">Physocrotaphus</emphasis>
Parry.
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- Mentum with triangular median tooth; ligula slender and gradually widening towards apex (
<figureCitation id="132BDB94FFFC647A42562CC0FA1AFE4B" box="[1303,1420,432,458]" captionStart="FIGURES 2–5" captionStartId="3.[151,269,1179,1203]" captionTargetBox="[168,1421,876,1146]" captionTargetId="figure-497@3.[151,1436,870,1165]" captionTargetPageId="3" captionText="FIGURES 2–5. Mentum (ventral view): 2 Helluodes devagiriensis n.sp.; 3 Physocrotaphus ceylonicus; 4 Pogonoglossus giganteus; 5 Schuelea drumonti. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5108348" httpUri="https://zenodo.org/record/5108348/files/figure.png" pageId="3" pageNumber="33">Figs. 4, 5</figureCitation>
); male sternite 3 with two oval depressions bearing filamentous elongated setae (
<figureCitation id="132BDB94FFFC647A43272CA8FB2EFE73" box="[1126,1208,472,498]" captionStart="FIGURES 15–18" captionStartId="8.[151,269,1462,1486]" captionTargetBox="[201,1408,184,1426]" captionTargetId="figure-234@8.[151,1436,160,1438]" captionTargetPageId="8" captionText="FIGURES 15–18. Median depression and cluster of filamentous setae on 2nd and 3rd sternites of male: 15 Physocrotaphus ceylonicus; 16 Pogonoglossus giganteus; 17 Helluodes devagiriensis n.sp.; 18 2nd and 3rd sternites of female Helluodes devagiriensis n.sp. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108354" httpUri="https://zenodo.org/record/5108354/files/figure.png" pageId="3" pageNumber="33">Fig. 16</figureCitation>
)..........................3
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3. Antennae longer, extended beyond the base of pronotum by 2–3 antennomeres; dorsal and ventral surfaces of antennomeres 5–11 devoid of impressions at base; antennomeres 2–11 with apical ring of elongate setae; mandibles long, narrow and weakly arcuate; anterior transverse sulcus of pronotum weakly developed; TLA = 7–17.6 (
<figureCitation id="132BDB94FFFC647A45462F08FDCAFD13" box="[519,604,632,658]" captionStart="FIGURES 23–26" captionStartId="10.[151,269,1306,1330]" captionTargetBox="[159,1424,770,1265]" captionTargetId="figure-264@10.[151,1436,764,1282]" captionTargetPageId="10" captionText="FIGURES 23–26. Habitus: 23 Helluodes devagiriensis n.sp.(Holotype); 24 Pogonoglossus giganteus (Holotype); 25 Schuelea drumonti (Holotype); 26 Physocrotaphus ceylonicus (Lectotype). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5108360" httpUri="https://zenodo.org/record/5108360/files/figure.png" pageId="3" pageNumber="33">Fig. 24</figureCitation>
) .................................................................... 
<taxonomicName id="4C10BC92FFFC647A43282F08FA03FD13" ID-CoL="8KYH6" authority="Chaudoir." authorityName="Chaudoir." box="[1129,1429,632,658]" class="Insecta" family="Carabidae" genus="Pogonoglossus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B9641B03FFFC647A43282F08FA8CFD10" box="[1129,1306,632,657]" italics="true" pageId="3" pageNumber="33">Pogonoglossus</emphasis>
Chaudoir.
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Antennae shorter, extended beyond the base of pronotum by only the last; dorsal and ventral surfaces of antennomeres 5–11 bi-impressed at base; antennomeres 2–11 without apical ring of setae; mandibles short, wide and weakly arcuate; anterior transverse sulcus of pronotum very deep; length 
<quantity id="4CE86AF4FFFC647A43B32F80FA0EFC8B" box="[1266,1432,752,778]" metricMagnitude="-2" metricUnit="m" metricValue="1.565" metricValueMax="1.68" metricValueMin="1.45" pageId="3" pageNumber="33" unit="mm" value="15.65" valueMax="16.8" valueMin="14.5">14.5–16.8mm</quantity>
. (
<figureCitation id="132BDB94FFFC647A478E2E68FEB2FCB3" box="[207,292,792,818]" captionStart="FIGURES 23–26" captionStartId="10.[151,269,1306,1330]" captionTargetBox="[159,1424,770,1265]" captionTargetId="figure-264@10.[151,1436,764,1282]" captionTargetPageId="10" captionText="FIGURES 23–26. Habitus: 23 Helluodes devagiriensis n.sp.(Holotype); 24 Pogonoglossus giganteus (Holotype); 25 Schuelea drumonti (Holotype); 26 Physocrotaphus ceylonicus (Lectotype). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5108360" httpUri="https://zenodo.org/record/5108360/files/figure.png" pageId="3" pageNumber="33">Fig. 25</figureCitation>
) ............................................................................................................................. 
<taxonomicName id="4C10BC92FFFC647A43962E68FA03FCB3" ID-CoL="8KYTL" authority="Baehr." authorityName="Baehr." box="[1239,1429,792,818]" class="Insecta" family="Carabidae" genus="Schuelea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B9641B03FFFC647A43962E68FAD6FCB0" box="[1239,1344,792,817]" italics="true" pageId="3" pageNumber="33">Schuelea</emphasis>
Baehr.
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