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<document id="A16E65DA23DA92717673FB1173E804BA" ID-CLB-Dataset="28344" ID-DOI="10.11646/zootaxa.4567.3.2" ID-GBIF-Dataset="6e80279e-4853-4c39-8ff2-f59c18084052" ID-ISSN="1175-5326" ID-Zenodo-Dep="2599025" ID-ZooBank="C3EBBF3C-7F69-4F94-8A55-4823616C5366" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553062736640" checkinUser="plazi" docAuthor="Conde-Vela, Víctor M." docDate="2019" docId="03FE87C6FFB3C44F7BC5F89CFC7361DD" docLanguage="en" docName="zootaxa.4567.3.2.pdf" docOrigin="Zootaxa 4567 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nereididae de Blainville 1818" docType="treatment" docVersion="9" lastPageNumber="452" masterDocId="FFC7FFBEFFB2C44D7B52FFB7FF9B625D" masterDocTitle="Re-evaluation of the morphology of the monotypic genera Lycastonereis Rao, 1981 and Ganganereis Misra, 1999 (Annelida, Phyllodocida, Nereididae)" masterLastPageNumber="460" masterPageNumber="450" pageNumber="451" updateTime="1698702017085" updateUser="ExternalLinkService">
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<mods:title id="CCEA531AD41A72207846A6B2A70A06E8">Re-evaluation of the morphology of the monotypic genera Lycastonereis Rao, 1981 and Ganganereis Misra, 1999 (Annelida, Phyllodocida, Nereididae)</mods:title>
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<mods:namePart id="854618B0473C9AFEA6D2A1DCE6DE109B">Conde-Vela, Víctor M.</mods:namePart>
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<mods:title id="F5888C7EDB480DB7012C7C47378A11A3">Zootaxa</mods:title>
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<mods:date id="310FFC26EC6C2FD7A72526FE3E9630ED">2019</mods:date>
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<mods:identifier id="573BAB33B2AA6AFD82510A6FCF531828" type="DOI">10.11646/zootaxa.4567.3.2</mods:identifier>
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<treatment id="03FE87C6FFB3C44F7BC5F89CFC7361DD" ID-DOI="http://doi.org/10.5281/zenodo.5280473" ID-GBIF-Taxon="154435168" ID-Zenodo-Dep="5280473" LSID="urn:lsid:plazi:treatment:03FE87C6FFB3C44F7BC5F89CFC7361DD" httpUri="http://treatment.plazi.org/id/03FE87C6FFB3C44F7BC5F89CFC7361DD" lastPageId="2" lastPageNumber="452" pageId="1" pageNumber="451">
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<subSubSection id="C34D655BFFB3C44C7BC5F89CFD8D65D7" pageId="1" pageNumber="451" type="nomenclature">
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<paragraph id="8BE836D0FFB3C44C7BC5F89CFDF06518" blockId="1.[151,619,1835,1861]" box="[151,619,1835,1861]" pageId="1" pageNumber="451">
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<heading id="D0A081BCFFB3C44C7BC5F89CFDF06518" bold="true" box="[151,619,1835,1861]" fontSize="11" level="1" pageId="1" pageNumber="451" reason="1">
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<emphasis id="B923EAC2FFB3C44C7BC5F89CFDF06518" bold="true" box="[151,619,1835,1861]" pageId="1" pageNumber="451">
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Family
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<taxonomicName id="4C574D53FFB3C44C7BA4F89CFDF06518" ID-CoL="84KJY" authority="de Blainville, 1818" authorityName="de Blainville" authorityYear="1818" box="[246,619,1835,1861]" class="Polychaeta" family="Nereididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="451" phylum="Annelida" rank="family">
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Nereididae de
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<bibRefCitation id="EFC64B21FFB3C44C7AF8F89CFDF06518" author="de Blainville, H. - M. D." box="[426,619,1835,1861]" pageId="1" pageNumber="451" pagination="78 - 85" refId="ref6494" refString="de Blainville, H. - M. D. (1818) Memoire sur la classe des Setipodes, partie des Vers a sang rouge de M. Cuvier, et des Annelides de M. de Lamarck. Bulletin des sciences par la Societe Philomathique de Paris, 97, 78 - 85." type="journal article" year="1818">Blainville, 1818</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE836D0FFB3C44C7BC5F8C7FD8D65D7" blockId="1.[151,534,1903,1930]" box="[151,534,1903,1930]" pageId="1" pageNumber="451">
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<heading id="D0A081BCFFB3C44C7BC5F8C7FD8D65D7" bold="true" box="[151,534,1903,1930]" fontSize="11" level="1" pageId="1" pageNumber="451" reason="1">
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<emphasis id="B923EAC2FFB3C44C7BC5F8C7FD8D65D7" bold="true" box="[151,534,1903,1930]" pageId="1" pageNumber="451">
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Genus
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<emphasis id="B923EAC2FFB3C44C7BBCF8D8FE0865D4" bold="true" box="[238,403,1903,1929]" italics="true" pageId="1" pageNumber="451">Lycastonereis</emphasis>
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<bibRefCitation id="EFC64B21FFB3C44C7AC8F8C7FD8D65D7" author="Rao, C. A. N." box="[410,534,1904,1930]" pageId="1" pageNumber="451" pagination="213 - 217" refId="ref7700" refString="Rao, C. A. N. (1981) On two polychaetes (Nereidae: Annelida) from estuarine waters of India. Bulletin of the Zoological Survey of India, 3 (3), 213 - 217." type="journal article" year="1981">Rao, 1981</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D655BFFB3C44C7BC5F801FE5F6596" box="[151,452,1973,1995]" pageId="1" pageNumber="451" type="description">
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<paragraph id="8BE836D0FFB3C44C7BC5F801FE5F6596" blockId="1.[151,869,1973,2026]" box="[151,452,1973,1995]" pageId="1" pageNumber="451">
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<emphasis id="B923EAC2FFB3C44C7BC5F801FEB86596" box="[151,291,1974,1995]" italics="true" pageId="1" pageNumber="451">Lycastonereis</emphasis>
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<bibRefCitation id="EFC64B21FFB3C44C7A7AF801FE166596" author="Rao, C. A. N." box="[296,397,1973,1995]" pageId="1" pageNumber="451" pagination="213 - 217" refId="ref7700" refString="Rao, C. A. N. (1981) On two polychaetes (Nereidae: Annelida) from estuarine waters of India. Bulletin of the Zoological Survey of India, 3 (3), 213 - 217." type="journal article" year="1981">Rao 1981</bibRefCitation>
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: 213.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D655BFFB3C44C7BC5F863FCFE65B7" box="[151,869,2004,2026]" pageId="1" pageNumber="451" type="materials_examined">
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<paragraph id="8BE836D0FFB3C44C7BC5F863FCFE65B7" blockId="1.[151,869,1973,2026]" box="[151,869,2004,2026]" pageId="1" pageNumber="451">
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<emphasis id="B923EAC2FFB3C44C7BC5F863FEB965B7" bold="true" box="[151,290,2004,2026]" pageId="1" pageNumber="451">
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<typeStatus id="54EC8872FFB3C44C7BC5F863FF5765B7" box="[151,204,2004,2026]" pageId="1" pageNumber="451">Type</typeStatus>
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species.
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</emphasis>
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<emphasis id="B923EAC2FFB3C44C7A7BF862FE2F65B7" box="[297,436,2005,2026]" italics="true" pageId="1" pageNumber="451">Lycastonereis</emphasis>
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<taxonomicName id="4C574D53FFB3C44C7AE9F863FDF165B7" authority="Rao, 1981" authorityName="Rao" authorityYear="1981" box="[443,618,2004,2026]" class="Polychaeta" family="Nereididae" genus="Lycastopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="1" pageNumber="451" phylum="Annelida" rank="species" species="indica">
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<emphasis id="B923EAC2FFB3C44C7AE9F863FE6165B7" box="[443,506,2004,2026]" italics="true" pageId="1" pageNumber="451">indica</emphasis>
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<bibRefCitation id="EFC64B21FFB3C44C7952F862FDF165B7" author="Rao, C. A. N." box="[512,618,2004,2026]" pageId="1" pageNumber="451" pagination="213 - 217" refId="ref7700" refString="Rao, C. A. N. (1981) On two polychaetes (Nereidae: Annelida) from estuarine waters of India. Bulletin of the Zoological Survey of India, 3 (3), 213 - 217." type="journal article" year="1981">Rao, 1981</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D655BFFB0C44F7BC5FF20FEC563F0" pageId="2" pageNumber="452" type="diagnosis">
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<paragraph id="8BE836D0FFB0C44F7BC5FF20FEC563F0" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="452">
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<emphasis id="B923EAC2FFB0C44F7BC5FF20FE8062ED" bold="true" box="[151,283,151,176]" pageId="2" pageNumber="452">Diagnosis (</emphasis>
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amended, new features highlighted in boldface). Prostomium with anterior margin entire. One pair of antennae and two pairs of eyes. Three pairs of anterior cirri. Pharynx with rounded papillae present in both oral and maxillary ring, in all areas except area V. Chaetigers 1 and 2 with neuroaciculae only. Notopodial dorsal ligules, notopodial prechaetal lobes, and neuroacicular ligules bilobed present in anterior chaetigers; notopodial dorsal ligules, notopodial prechaetal lobes and neuropodial postchaetal lobes absent in posterior chaetigers. Neuropodial ventral ligules absent throughout body. Notochaetae homogomph spinigers; neurochaetae homogomph spinigers and falcigers in supra-acicular fascicles, homogomph and sesquigomph spinigers and homogomph falcigers in subacicular fascicles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D655BFFB0C44F7B95FE00FC7361DD" pageId="2" pageNumber="452" type="discussion">
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<paragraph id="8BE836D0FFB0C44F7B95FE00FEDB60AD" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="452">
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<emphasis id="B923EAC2FFB0C44F7B95FE00FEA0638D" bold="true" box="[199,315,439,464]" pageId="2" pageNumber="452">Remarks.</emphasis>
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In the original paper, the name of the author appears as ‘C. A. Nageswara Rao’, suggesting the last two words correspond to the surnames, hence the use of the combination ‘Nageswara-Rao’ as the authority for the taxa he proposed. This combination is used by Rao himself when citing some of his papers (e.g.
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<bibRefCitation id="EFC64B21FFB0C44F7F8AFDB7FAD26045" author="Rao, C. A. N." box="[1240,1353,512,536]" pageId="2" pageNumber="452" pagination="43 - 47" refId="ref7770" refString="Rao, C. A. N. (1999) Observations on a collection of polychaetes from Godavari Estuary, Andhra Pradesh. Records of the Zoological Survey of India, 97 (1), 43 - 47." type="journal article" year="1999">Rao 1999</bibRefCitation>
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;
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<bibRefCitation id="EFC64B21FFB0C44F7E06FDB7FA0A6045" author="Rao, C. A. N." box="[1364,1425,512,536]" pageId="2" pageNumber="452" pagination="12 - 31" refId="ref7811" refString="Rao, C. A. N. (2008) Polychaeta (Annelida). Estuarine Ecosystem Series: Krishna Estuary, 5, 12 - 31." type="journal article" year="2008">2008</bibRefCitation>
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). In other occasions, he (e.g.
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<bibRefCitation id="EFC64B21FFB0C44F7A9FFD92FDA06061" author="Rao, C. A. N." box="[461,571,548,572]" pageId="2" pageNumber="452" pagination="199 - 209" refId="ref7742" refString="Rao, C. A. N. (1998) Polychaeta: Annelida. Estuarine Ecosystem Series: Mahanadi Estuary, 3, 199 - 209." type="journal article" year="1998">Rao 1998</bibRefCitation>
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,
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<bibRefCitation id="EFC64B21FFB0C44F7915FD93FD1A6061" author="Rao, C. A. N." box="[583,641,548,572]" pageId="2" pageNumber="452" pagination="12 - 31" refId="ref7811" refString="Rao, C. A. N. (2008) Polychaeta (Annelida). Estuarine Ecosystem Series: Krishna Estuary, 5, 12 - 31." type="journal article" year="2008">2008</bibRefCitation>
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) and their colleagues (e.g.
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<bibRefCitation id="EFC64B21FFB0C44F78E3FD93FBE06061" author="Alfred, J. R. B. & Chakraborty, S. & Chakraborty, R. K." box="[945,1147,548,573]" pageId="2" pageNumber="452" pagination="53 - 54" refId="ref6136" refString="Alfred, J. R. B., Chakraborty, S. & Chakraborty, R. K. (2000) Annelida: Polychaeta. In: A Bibliography of the Zoological Survey of India Publications in the Twentieth Century (1916 - 1999). Director Zoological Survey of India, Calcutta, pp. 53 - 54." type="book chapter" year="2000">
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Alfred
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<emphasis id="B923EAC2FFB0C44F7F53FD92FBA16060" box="[1025,1082,548,573]" italics="true" pageId="2" pageNumber="452">et al.</emphasis>
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2000
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</bibRefCitation>
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) cited his papers as ‘Rao, C. A. N.’, suggesting that ‘Rao’ is the surname only and that ‘Nageswara’ is a middle name. When citing the species he described, the author used the combination ‘Rao’, as the case for
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<taxonomicName id="4C574D53FFB0C44F78A8FDDBFAF960D9" authority="Rao, 1981" authorityName="Rao" authorityYear="1981" box="[1018,1378,620,645]" class="Polychaeta" family="Nereididae" genus="Namalycastis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="452" phylum="Annelida" rank="species" species="fauveli">
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<emphasis id="B923EAC2FFB0C44F78A8FDDBFB7E60D8" box="[1018,1253,620,645]" italics="true" pageId="2" pageNumber="452">Namalycastis fauveli</emphasis>
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<bibRefCitation id="EFC64B21FFB0C44F7FBFFDDAFAF960D9" author="Rao, C. A. N." box="[1261,1378,620,644]" pageId="2" pageNumber="452" pagination="213 - 217" refId="ref7700" refString="Rao, C. A. N. (1981) On two polychaetes (Nereidae: Annelida) from estuarine waters of India. Bulletin of the Zoological Survey of India, 3 (3), 213 - 217." type="journal article" year="1981">Rao, 1981</bibRefCitation>
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</taxonomicName>
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(e.g.
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<bibRefCitation id="EFC64B21FFB0C44F7BC5FD27FE9360F5" author="Rao, C. A. N." box="[151,264,656,680]" pageId="2" pageNumber="452" pagination="43 - 47" refId="ref7770" refString="Rao, C. A. N. (1999) Observations on a collection of polychaetes from Godavari Estuary, Andhra Pradesh. Records of the Zoological Survey of India, 97 (1), 43 - 47." type="journal article" year="1999">Rao 1999</bibRefCitation>
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: 44;
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<bibRefCitation id="EFC64B21FFB0C44F7A6CFD27FE3460F5" author="Rao, C. A. N." box="[318,431,656,680]" pageId="2" pageNumber="452" pagination="199 - 209" refId="ref7742" refString="Rao, C. A. N. (1998) Polychaeta: Annelida. Estuarine Ecosystem Series: Mahanadi Estuary, 3, 199 - 209." type="journal article" year="1998">Rao 1998</bibRefCitation>
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: 201;
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<bibRefCitation id="EFC64B21FFB0C44F7AA1FD27FDFF60F5" author="Rao, C. A. N." box="[499,612,656,680]" pageId="2" pageNumber="452" pagination="12 - 31" refId="ref7811" refString="Rao, C. A. N. (2008) Polychaeta (Annelida). Estuarine Ecosystem Series: Krishna Estuary, 5, 12 - 31." type="journal article" year="2008">Rao 2008</bibRefCitation>
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: 16).
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<bibRefCitation id="EFC64B21FFB0C44F79F1FD38FC3360F5" author="Misra, A." box="[675,936,655,680]" pageId="2" pageNumber="452" pagination="125 - 225" refId="ref7395" refString="Misra, A. (1999) Polychaete. In: A. K. Ghosh (Ed.), State Fauna Series: Fauna of West Bengal, Part 10. Calcutta, pp. 125 - 225." type="book chapter" year="1999">Misra (1999: 147, 159)</bibRefCitation>
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and
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<bibRefCitation id="EFC64B21FFB0C44F78B2FD38FAA960F5" author="Muir, A. I. & Hossain, M. M." box="[992,1330,655,680]" pageId="2" pageNumber="452" pagination="1 - 27" refId="ref7476" refString="Muir, A. I. & Hossain, M. M. (2014) The intertidal polychaete (Annelida) fauna of the Sitakunda coast (Chittagong, Bangladesh), with notes on the Capitellidae, Glyceridae, Lumbrineridae, Nephtyidae, Nereididae and Phyllodocidae of the " Northern Bay of Bengal Ecoregion. " ZooKeys, 419, 1 - 27. https: // doi. org / 10.3897 / zookeys. 419.7557." type="journal article" year="2014">Muir & Hossain (2014: 7, 12)</bibRefCitation>
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followed this combination for
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<taxonomicName id="4C574D53FFB0C44F7ADEFD02FE686090" authorityName="Rao" authorityYear="1981" box="[396,499,692,717]" class="Polychaeta" family="Nereididae" genus="Lycastopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="452" phylum="Annelida" rank="species" species="indica">
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<emphasis id="B923EAC2FFB0C44F7ADEFD02FE686090" box="[396,499,692,717]" italics="true" pageId="2" pageNumber="452">L. indica</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C574D53FFB0C44F7960FD02FD396090" authorityName="Rao" authorityYear="1981" box="[562,674,692,717]" class="Polychaeta" family="Nereididae" genus="Namalycastis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="452" phylum="Annelida" rank="species" species="fauveli">
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<emphasis id="B923EAC2FFB0C44F7960FD02FD396090" box="[562,674,692,717]" italics="true" pageId="2" pageNumber="452">N. fauveli</emphasis>
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</taxonomicName>
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. Here the combination ‘Rao’ for the authorship, as stated by the author, is used.
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</paragraph>
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<paragraph id="8BE836D0FFB0C44F7B95FD4AFC7361DD" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="452">
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<emphasis id="B923EAC2FFB0C44F7B95FD4AFE356148" box="[199,430,764,789]" italics="true" pageId="2" pageNumber="452">
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Lycastonereis
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<taxonomicName id="4C574D53FFB0C44F7A3AFD4BFE356148" authorityName="Rao" authorityYear="1981" box="[360,430,764,789]" class="Polychaeta" family="Nereididae" genus="Lycastopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="452" phylum="Annelida" rank="species" species="indica">indica</taxonomicName>
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</emphasis>
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has a set of unique features, which are highlighted in the amended diagnosis. Additional features included in the current diagnosis are the lack of neuropodial ventral ligules along the body, the lack of several parapodial processes toward posterior end, and homogomph falcigers deprived of dentition. The presence of papillae in both oral and maxillary rings is a unique feature in this genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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