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<document id="87ED6FB8E3A47154499717C3CAA2142A" ID-DOI="10.1649/934.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5461668" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630266972764" checkinUser="valdenar" docAuthor="Liang, Hongbin &amp; Kavanaugh, David H." docDate="2007" docId="03D187BCFFCB481D291CB85C24CFFC5F" docLanguage="en" docName="934.1.pdf" docOrigin="The Coleopterists Bulletin (Oxford, England) 61 (1)" docSource="http://dx.doi.org/10.1649/934.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Dendrocellus javanus Liang and Kavanaugh 2007, new species" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="FFE8FFC4FFD348042808BC0D270CFFA4" masterDocTitle="Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species" masterLastPageNumber="39" masterPageNumber="1" pageNumber="1" updateTime="1699204912656" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="71814159CFEE589A8BDDCC951402E03C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FD6F2BE7C7DB6A3D465A230D94FE2C12">
<mods:title id="7C47C9EDE44262481BDB8117FEAC7240">Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species</mods:title>
</mods:titleInfo>
<mods:name id="BEA6113CD6875321194E7035FB32ED00" type="personal">
<mods:role id="E266F0DC2ED1BF8776F250235D5C1EE3">
<mods:roleTerm id="A293BCBE57FA45AC58DAC0ACF10D1607">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37ACC014944FCAAEB9F978B39E38CB19">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:name id="4AEDBB83DD57FBF835FF30CE5B4807DC" type="personal">
<mods:role id="5C88C8A9C8446A43D7AC00015F1F530D">
<mods:roleTerm id="4222189F77C4783F6A6609611E1D79BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FF7476C3340D5250F42A1C7E0CF709F">Kavanaugh, David H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D4688A9F72E041B3FEBC24CE2E4171BE">text</mods:typeOfResource>
<mods:relatedItem id="0AFAD57D0EC6EC5471D688BDD7026DB8" type="host">
<mods:titleInfo id="90E0297CC763C779A664612729632169">
<mods:title id="32AEADEC8BE2DB60D3A4685CBA81638B">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="9FD4D61EBAF1BACCAD82D7EE3B1487DE">
<mods:date id="ADE9D1551980F079C618D0EB022807BC">2007</mods:date>
<mods:detail id="5AFFBC5B622A713DB650D49C5327FA40" type="series">
<mods:title id="EB7290861F6EDD2877B5D6CAE7E14860">Oxford, England</mods:title>
</mods:detail>
<mods:detail id="D4F6987DF9E4A01EC2D4C2F5F281F391" type="pubDate">
<mods:number id="E5D44722C312D0A996CC65FE9E8E37EF">2007-03-31</mods:number>
</mods:detail>
<mods:detail id="3C99A94C4C9C020F47DB8E161368CCED" type="volume">
<mods:number id="31B34F548ADA1CE5940CDB0E1DF00A0E">61</mods:number>
</mods:detail>
<mods:detail id="499F779A9924B7A94657294EF2373F2C" type="issue">
<mods:number id="A3C87778F55B5719E1DDBA13E9742245">1</mods:number>
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<mods:start id="9B6882D25922251A0D9B9D1B77067595">1</mods:start>
<mods:end id="519CEE2438C1C8525C68279364FE83E0">39</mods:end>
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<mods:url id="FC1FBF6E5FB418ADF32BE67635128CD4">http://dx.doi.org/10.1649/934.1</mods:url>
</mods:location>
<mods:classification id="34D1F209C28CB6EA6F0B662D9CBE4EA8">journal article</mods:classification>
<mods:identifier id="13B9160E4DEE16A6B0B87325BFE5B0A5" type="DOI">10.1649/934.1</mods:identifier>
<mods:identifier id="AD223979E98355C8169E0906DD866092" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="641E856433A8C7045A3ABA415D1FADF3" type="Zenodo-Dep">5461668</mods:identifier>
</mods:mods>
<treatment id="03D187BCFFCB481D291CB85C24CFFC5F" LSID="urn:lsid:plazi:treatment:03D187BCFFCB481D291CB85C24CFFC5F" httpUri="http://treatment.plazi.org/id/03D187BCFFCB481D291CB85C24CFFC5F" lastPageId="25" pageId="24">
<subSubSection id="C3626521FFCB481C291CB85C2441FBC3" box="[276,845,1105,1127]" pageId="24" type="nomenclature">
<paragraph id="8BC736AAFFCB481C291CB85C2441FBC3" blockId="24.[276,845,1105,1155]" box="[276,845,1105,1127]" pageId="24">
<heading id="D08F81C6FFCB481C291CB85C2441FBC3" box="[276,845,1105,1127]" centered="true" fontSize="9" level="2" pageId="24" reason="2">
<emphasis id="B90CEAB8FFCB481C291CB85C2441FBC3" bold="true" box="[276,845,1105,1127]" pageId="24">
<taxonomicName id="4C784D29FFCB481C291CB85C25C2FBC3" ID-CoL="34P5C" authority="Liang and Kavanaugh" authorityName="Liang and Kavanaugh" authorityYear="2007" box="[276,718,1105,1127]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C291CB85C26EBFBC3" bold="true" box="[276,487,1105,1127]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang and Kavanaugh
</taxonomicName>
,
<taxonomicNameLabel id="A23F57C3FFCB481C2AD3B85C2441FBC3" box="[731,845,1105,1127]" pageId="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCB481C29BCB860258AF986" pageId="24" type="description">
<paragraph id="8BC736AAFFCB481C29BCB86025A1FB27" blockId="24.[276,845,1105,1155]" box="[436,685,1133,1155]" pageId="24">
(
<figureCitation id="13432A2FFFCB481C29B4B860251CFB27" box="[444,528,1133,1155]" captionStart="Figs" captionStartId="4.[167,206,1376,1395]" captionTargetBox="[171,950,195,1353]" captionTargetId="figure-1@4.[168,953,194,1354]" captionTargetPageId="4" captionText="Figs. 1–12. Habitus of Dendrocellus species. 1) D. australis (male, Congo); 2) D. coelestinus (syntype); 3) D. coelestinus (type of D. paralellus); 4) D. confusus (male, Thailand); 5) D. crassus (male, Borneo); 6) D. elegans (male, New Guinea); 7) D. rugicollis (lectotype); 8) D. geniculatus (holotype); 9) D. geniculatus (holotype of D. gilsoni); 10) D. gestroi (lectotype); 11) D. inexpectus (holotype); 12) D. javanus (holotype). Scale line 5 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461670" httpUri="https://zenodo.org/record/5461670/files/figure.png" pageId="24">Figs. 12</figureCitation>
,
<figureCitation id="13432A2FFFCB481C2A14B860253AFB27" box="[540,566,1133,1155]" captionStart="Figs" captionStartId="6.[167,206,1502,1521]" captionTargetBox="[191,931,732,1478]" captionTargetId="figure-42@6.[188,932,731,1479]" captionTargetPageId="6" captionText="Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461676" httpUri="https://zenodo.org/record/5461676/files/figure.png" pageId="24">34</figureCitation>
,
<figureCitation id="13432A2FFFCB481C2A4BB8602551FB27" box="[579,605,1133,1155]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="24">62</figureCitation>
,
<figureCitation id="13432A2FFFCB481C2A61B86025ABFB27" box="[617,679,1133,1155]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="24">89–90</figureCitation>
)
</paragraph>
<paragraph id="8BC736AAFFCB481C28A2B898258AF986" blockId="24.[146,975,1173,1570]" pageId="24">
<emphasis id="B90CEAB8FFCB481C28A2B89827E0FB0F" bold="true" box="[170,236,1173,1195]" pageId="24">Types.</emphasis>
<materialsCitation id="3B103CF7FFCB481C28FBB898256EFB62" collectionCode="ISNB, R" collectorName="I. Sc. &amp; Coll. Le &amp; I. Sc. N. B. Java'" location="Coll. Le Moult'" pageId="24" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<typeStatus id="54C38808FFCB481C28FBB898265AFB0F" box="[243,342,1173,1195]" pageId="24" type="holotype">Holotype</typeStatus>
: a male (
<collectionCode id="ED69AE6FFFCB481C29BCB89826FEFB0F" box="[436,498,1173,1195]" pageId="24">ISNB</collectionCode>
), ‘‘
<collectingRegion id="49BCF848FFCB481C2A06B898254DFB0F" box="[526,577,1173,1195]" country="Indonesia" name="Jawa Timur" pageId="24">Java</collectingRegion>
, Seokaboemi’’/‘‘
<collectionCode id="ED69AE6FFFCB481C2AE6B898240CFB0F" box="[750,768,1173,1195]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
.
<collectorName id="268D537CFFCB481C2B03B898244CFB0F" box="[779,832,1173,1195]" pageId="24">I. Sc.</collectorName>
N. B. 19.458,
<location id="8EA76071FFCB481C289AB8BD2638FB62" LSID="urn:lsid:plazi:treatment:03D187BCFFCB481D291CB85C24CFFC5F:8EA76071FFCB481C289AB8BD2638FB62" box="[146,308,1200,1222]" name="Coll. Le Moult'" pageId="24" stateProvince="Jawa Timur">
<collectorName id="268D537CFFCB481C289AB8BD27E8FB62" box="[146,228,1200,1222]" pageId="24">Coll. Le</collectorName>
Moult’
</location>
’/‘‘Coll.
<collectionCode id="ED69AE6FFFCB481C298BB8BD2694FB62" box="[387,408,1200,1222]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
.
<collectorName id="268D537CFFCB481C29AAB8BD2541FB62" box="[418,589,1200,1222]" pageId="24">I. Sc. N. B. Java’</collectorName>
’/’’
</materialsCitation>
<typeStatus id="54C38808FFCB481C2A6BB8BD25F8FB62" box="[611,756,1200,1222]" pageId="24" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C784D29FFCB481C2AF6B8BC266FFB45" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C2AF6B8BC24C3FB62" box="[766,975,1201,1222]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCB481C2961B8C626AEFB45" box="[361,418,1227,1249]" pageId="24" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [red label].
<materialsCitation id="3B103CF7FFCB481C290BB8EB256CFAB3" collectionCode="ISNB, R" collectorName="Coll. Putzeys' &amp; I. Sc. N. B. Java'" location="Java" pageId="24" specimenCount="6" specimenCount-female="1" stateProvince="Jawa Timur" typeStatus="paratype">
<typeStatus id="54C38808FFCB481C290BB8EB266BFB58" box="[259,359,1254,1276]" pageId="24" type="paratype">Paratypes</typeStatus>
(total
<specimenCount id="9D7EFD23FFCB481C29A3B8EB2524FB58" box="[427,552,1254,1276]" pageId="24" type="generic">5 specimens</specimenCount>
):
<specimenCount id="9D7EFD23FFCB481C2A30B8EB2581FB58" box="[568,653,1254,1276]" pageId="24" type="female">1 female</specimenCount>
(
<collectionCode id="ED69AE6FFFCB481C2A93B8EB25D5FB58" box="[667,729,1254,1276]" pageId="24">ISNB</collectionCode>
), ‘‘
<collectingRegion id="49BCF848FFCB481C2AFCB8EB242BFB58" box="[756,807,1254,1276]" country="Indonesia" name="Jawa Timur" pageId="24">Java</collectingRegion>
, Soc. Ent. Belg.,
<collectorName id="268D537CFFCB481C289AB90C262AFAB3" box="[146,294,1281,1303]" pageId="24">Coll. Putzeys’</collectorName>
’/‘‘Coll.
<collectionCode id="ED69AE6FFFCB481C297DB90C2686FAB3" box="[373,394,1281,1303]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
.
<collectorName id="268D537CFFCB481C299EB90C2547FAB3" box="[406,587,1281,1303]" pageId="24">I. Sc. N. B. Java’</collectorName>
’/’’
</materialsCitation>
<typeStatus id="54C38808FFCB481C2A68B90C25E2FAB3" box="[608,750,1281,1303]" pageId="24" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C784D29FFCB481C2AF3B90C266FFA95" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C2AF3B90C24C3FAB2" box="[763,975,1281,1302]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCB481C2961B91626AEFA95" box="[361,418,1307,1329]" pageId="24" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [yellow label];
<materialsCitation id="3B103CF7FFCB481C292CB93B25A4FAE8" box="[292,680,1334,1356]" collectionCode="ISNB" collectorName="Java'" pageId="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jawa Timur" typeStatus="paratype">
<specimenCount id="9D7EFD23FFCB481C292CB93B266BFAE8" box="[292,359,1334,1356]" pageId="24" type="male">1 male</specimenCount>
and
<specimenCount id="9D7EFD23FFCB481C2991B93B26E2FAE8" box="[409,494,1334,1356]" pageId="24" type="female">1 female</specimenCount>
(
<collectionCode id="ED69AE6FFFCB481C29F3B93B2535FAE8" box="[507,569,1334,1356]" pageId="24">ISNB</collectionCode>
), ‘‘
<collectorName id="268D537CFFCB481C2A5DB93B259CFAE8" box="[597,656,1334,1356]" pageId="24">Java’</collectorName>
’/‘‘
</materialsCitation>
<materialsCitation id="3B103CF7FFCB481C2AA0B93B2628FAC3" collectionCode="R" collectorName="Mus. Hist. Nat. Belg. I. G. &amp; Coll" pageId="24" specimenCount="1" stateProvince="Jawa Timur" typeStatus="paratype">
<collectionCode id="ED69AE6FFFCB481C2AA0B93B25B0FAE8" box="[680,700,1334,1356]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
.
<collectorName id="268D537CFFCB481C2ACDB93B24C3FAE8" box="[709,975,1334,1356]" pageId="24">Mus. Hist. Nat. Belg. I.G.</collectorName>
12.595’’/‘‘
<collectorName id="268D537CFFCB481C28FEB95C2628FAC3" box="[246,292,1361,1383]" pageId="24">Coll</collectorName>
</materialsCitation>
.
<materialsCitation id="3B103CF7FFCB481C2925B95C2507FAC3" box="[301,523,1361,1383]" collectionCode="R" collectorName="Sc. &amp; Java'" pageId="24" specimenCount="1" stateProvince="Jawa Timur" typeStatus="paratype">
<collectionCode id="ED69AE6FFFCB481C2925B95C264EFAC3" box="[301,322,1361,1383]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
. I.
<collectorName id="268D537CFFCB481C2969B95C268CFAC3" box="[353,384,1361,1383]" pageId="24">Sc.</collectorName>
N. B.
<collectorName id="268D537CFFCB481C29CAB95C26FAFAC3" box="[450,502,1361,1383]" pageId="24">Java’</collectorName>
’/’’
</materialsCitation>
<typeStatus id="54C38808FFCB481C2A04B95C2596FAC3" box="[524,666,1361,1383]" pageId="24" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C784D29FFCB481C2AACB95C2605FA26" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C2AACB95C2479FAC2" box="[676,885,1361,1382]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCB481C2918B9612646FA26" box="[272,330,1388,1410]" pageId="24" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [yellow label];
<materialsCitation id="3B103CF7FFCB481C28DEB98B245FFA38" box="[214,851,1414,1436]" collectionCode="ISNB" collectorName="Java' &amp; Soc. Ent. Belg. Coll. Putzeys' &amp; Coll. Putzeys' &amp; Coll" pageId="24" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur" typeStatus="paratype">
<specimenCount id="9D7EFD23FFCB481C28DEB98B2626FA38" box="[214,298,1414,1436]" pageId="24" type="female">1 female</specimenCount>
(
<collectionCode id="ED69AE6FFFCB481C2930B98B267AFA38" box="[312,374,1414,1436]" pageId="24">ISNB</collectionCode>
), ‘‘
<collectorName id="268D537CFFCB481C2999B98B26C5FA38" box="[401,457,1414,1436]" pageId="24">Java’</collectorName>
’/‘‘
<collectorName id="268D537CFFCB481C29E8B98B241CFA38" box="[480,784,1414,1436]" pageId="24">Soc. Ent. Belg. Coll. Putzeys’</collectorName>
’/‘‘
<collectorName id="268D537CFFCB481C2B2DB98B245FFA38" box="[805,851,1414,1436]" pageId="24">Coll</collectorName>
</materialsCitation>
.
<materialsCitation id="3B103CF7FFCB481C2B54B98B260EFA13" collectionCode="R" collectorName="I. Sc. N. B. Java'" pageId="24" specimenCount="1" stateProvince="Jawa Timur" typeStatus="paratype">
<collectionCode id="ED69AE6FFFCB481C2B54B98B247DFA38" box="[860,881,1414,1436]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24">R</collectionCode>
.
<collectorName id="268D537CFFCB481C2B73B98B27E1FA13" pageId="24">I. Sc. N. B. Java’</collectorName>
’/’’
</materialsCitation>
<typeStatus id="54C38808FFCB481C290AB9AC269CFA13" box="[258,400,1441,1463]" pageId="24" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C784D29FFCB481C29ADB9AF247BFA13" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" box="[421,887,1441,1463]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C29ADB9AF258DFA13" box="[421,641,1442,1463]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCB481C2B8FB9AC24C7FA13" box="[903,971,1441,1463]" pageId="24" rank="species">n. sp.</taxonomicNameLabel>
, designated by H. B. Liang &amp; D. H. Kavanaugh 2006’’ [yellow label];
<materialsCitation id="3B103CF7FFCB481C2B8FB9B12470FA49" collectionCode="ZMHU" collectorName="Occid. Dr. Standgr." location="Klug'" pageId="24" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Timur" typeStatus="paratype">
<specimenCount id="9D7EFD23FFCB481C2B8FB9B124C3FA76" box="[903,975,1468,1490]" pageId="24" type="male">1 male</specimenCount>
(
<collectionCode id="ED69AE6FFFCB481C2891B9DA27E0FA49" box="[153,236,1495,1517]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="24">ZMHU</collectionCode>
), ‘‘
<collectingRegion id="49BCF848FFCB481C2901B9DA2636FA49" box="[265,314,1495,1517]" country="Indonesia" name="Jawa Timur" pageId="24">Java</collectingRegion>
<collectorName id="268D537CFFCB481C294AB9DA251EFA49" box="[322,530,1495,1517]" pageId="24">Occid. Dr. Standgr.</collectorName>
’’/‘‘
<taxonomicName id="4C784D29FFCB481C2A26B9DA242AFA48" baseAuthorityName="Klug" baseAuthorityYear="1834" box="[558,806,1495,1516]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="B90CEAB8FFCB481C2A26B9DA242AFA48" box="[558,806,1495,1516]" italics="true" pageId="24">Dendrocellus geniculatus</emphasis>
</taxonomicName>
<location id="8EA76071FFCB481C2B25B9DA246BFA49" LSID="urn:lsid:plazi:treatment:03D187BCFFCB481D291CB85C24CFFC5F:8EA76071FFCB481C2B25B9DA246BFA49" box="[813,871,1495,1517]" name="Klug'" pageId="24" stateProvince="Jawa Timur">Klug’</location>
’/’’
</materialsCitation>
<materialsCitation id="3B103CF7FFCB481C2B75B9DA258FF986" collectionCode="PARA-, TYPE" collectorName="H. B. Liang &amp; D. H. Kavanaugh" pageId="24" specimenCount="1" stateProvince="Jawa Timur" typeStatus="paratype">
<collectionCode id="ED69AE6FFFCB481C2B75B9DA27D9F9AC" pageId="24">PARA- TYPE</collectionCode>
,
<taxonomicName id="4C784D29FFCB481C28ECB9FF25ACF9AC" authority="Liang &amp; Kavanaugh" authorityName="Liang &amp; Kavanaugh" authorityYear="2007" box="[228,672,1522,1544]" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="javanus" status="sp. nov.">
<emphasis id="B90CEAB8FFCB481C28ECB9FF26B6F9A3" box="[228,442,1522,1543]" italics="true" pageId="24">Dendrocellus javanus</emphasis>
Liang &amp; Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="A23F57C3FFCB481C2AA3B9FF25E5F9AC" box="[683,745,1522,1544]" pageId="24" rank="species">n. sp.</taxonomicNameLabel>
, designated by
<collectorName id="268D537CFFCB481C2B9BB9FF27C1F986" pageId="24">H. B. Liang</collectorName>
&amp;
<collectorName id="268D537CFFCB481C28F8BA0126A7F986" box="[240,427,1548,1570]" pageId="24">D. H. Kavanaugh</collectorName>
2006’’ [yellow label]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBCCF250EFF7C" box="[199,514,194,216]" pageId="25" type="materials_examined">
<paragraph id="8BC736AAFFCA481D28CFBCCF250EFF7C" blockId="25.[175,1005,194,1019]" box="[199,514,194,216]" pageId="25">
<materialsCitation id="3B103CF7FFCA481D28CFBCCF26F1FF7C" box="[199,509,194,216]" country="Indonesia" location="Java" pageId="25" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="B90CEAB8FFCA481D28CFBCCF265DFF7C" bold="true" box="[199,337,194,216]" pageId="25">
<typeStatus id="54C38808FFCA481D28CFBCCF27F5FF7C" box="[199,249,194,216]" pageId="25">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49BCF848FFCA481D2952BCCF2687FF7C" box="[346,395,194,216]" country="Indonesia" name="Jawa Timur" pageId="25">Java</collectingRegion>
,
<collectingCountry id="F36F763AFFCA481D299EBCCF26F1FF7C" box="[406,509,194,216]" name="Indonesia" pageId="25">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBCD32596FEE7" pageId="25" type="diagnosis">
<paragraph id="8BC736AAFFCA481D28CFBCD32596FEE7" blockId="25.[175,1005,194,1019]" pageId="25">
<emphasis id="B90CEAB8FFCA481D28CFBCD32623FF50" bold="true" box="[199,303,222,244]" pageId="25">Diagnosis.</emphasis>
Femora brown with apical fifth or sixth black; tarsal claws (
<figureCitation id="13432A2FFFCA481D2B93BCD324E9FF50" box="[923,997,222,244]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="25">Fig. 62</figureCitation>
) with longest pectination subequal to width of tarsal claw base; elytra with outer angles slightly dentate; aedeagus with apical lamella moderately long, slightly digitiform, not thickened at apex (
<figureCitation id="13432A2FFFCA481D2A1CBD202583FEE7" box="[532,655,301,323]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="25">Figs. 89–90</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBD442519FC87" pageId="25" type="description">
<paragraph id="8BC736AAFFCA481D28CFBD442519FC87" blockId="25.[175,1005,194,1019]" pageId="25">
<emphasis id="B90CEAB8FFCA481D28CFBD44264CFEFB" bold="true" box="[199,320,329,351]" pageId="25">Description.</emphasis>
Length
<quantity id="4C809B4FFFCA481D2992BD44252CFEFB" box="[410,544,329,351]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.2" metricValueMin="9.2" pageId="25" unit="mm" value="9.7" valueMax="10.2" valueMin="9.2">9.2–10.2 mm</quantity>
, width
<quantity id="4C809B4FFFCA481D2A67BD4425E7FEFB" box="[623,747,329,351]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="25" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
. Black, head, pronotum, and elytra with green luster, mixed with slight blue luster; apical third of Ant 1 and basal two thirds of Ant 3 black, basal two thirds of Ant 1, apical third of Ant 3, and Ant 4–11 brown or yellow; apical fifth or sixth of femora black, remainder brown or yellow; tibiae and tarsi brown or yellow. Head (
<figureCitation id="13432A2FFFCA481D2B28BDBE2466FE6D" box="[800,874,435,457]" captionStart="Figs" captionStartId="6.[167,206,1502,1521]" captionTargetBox="[191,931,732,1478]" captionTargetId="figure-42@6.[188,932,731,1479]" captionTargetPageId="6" captionText="Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461676" httpUri="https://zenodo.org/record/5461676/files/figure.png" pageId="25">Fig. 34</figureCitation>
) wider than pronotum, HW/PW
<emphasis id="B90CEAB8FFCA481D298CBDC32694FE40" bold="true" box="[388,408,462,484]" italics="true" pageId="25">5</emphasis>
1.18 (1.13–1.23); vertex convex, coarsely punctate; Ant 1/ Ant 3
<emphasis id="B90CEAB8FFCA481D28FABDE5260AFE5A" bold="true" box="[242,262,488,510]" italics="true" pageId="25">5</emphasis>
3.35 (3.24–3.46); labrum with medial lobe slightly protruded anteriorly; genae moderately tumid; eyes large, about 2.2 times as long as genae. Pronotum densely punctate, narrow, PL/PW
<emphasis id="B90CEAB8FFCA481D2A10BE102520FD97" bold="true" box="[536,556,541,563]" italics="true" pageId="25">5</emphasis>
1.46 (1.40–1.50); lateral margins gradually widened from apical angles to middle, strongly sinuate before basal angles; basal angles protruded laterally; basal foveae quite deep; middle line shallow. Elytra long, slightly widened from base to apical third, then narrowed to apex in both sexes (more so in female); EL/EW
<emphasis id="B90CEAB8FFCA481D2A03BE852513FD3A" bold="true" box="[523,543,648,670]" italics="true" pageId="25">5</emphasis>
1.82 (1.70–1.86), EW/PW
<emphasis id="B90CEAB8FFCA481D2B23BE852433FD3A" bold="true" box="[811,831,648,670]" italics="true" pageId="25">5</emphasis>
2.23 (2.12–2.32); outer angles slightly dentate (similar to
<figureCitation id="13432A2FFFCA481D2A59BEAE2591FD1D" box="[593,669,675,697]" captionStart="Figs" captionStartId="7.[196,235,713,732]" captionTargetBox="[191,987,195,690]" captionTargetId="figure-169@7.[191,988,194,690]" captionTargetPageId="7" captionText="Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461678" httpUri="https://zenodo.org/record/5461678/files/figure.png" pageId="25">Fig. 40</figureCitation>
); interval flat, densely punctate. Protarsomere 3 slightly asymmetrical in male; tarsal claw with 5–6 long pectinations, length of the longest pectination subequal to width of claw base (
<figureCitation id="13432A2FFFCA481D28BFBEFE2608FCAD" box="[183,260,755,777]" captionStart="Figs" captionStartId="9.[196,235,972,991]" captionTargetBox="[176,1003,194,949]" captionTargetId="figure-360@9.[176,1003,194,949]" captionTargetPageId="9" captionText="Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5461686" httpUri="https://zenodo.org/record/5461686/files/figure.png" pageId="25">Fig. 62</figureCitation>
). Aedeagus with apical lamella moderately long, slightly digitiform, unthickened at apex (
<figureCitation id="13432A2FFFCA481D2987BF002506FC87" box="[399,522,781,803]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="25">Figs. 89–90</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBF2425FAFC9B" box="[199,758,809,831]" pageId="25" type="distribution">
<paragraph id="8BC736AAFFCA481D28CFBF2425FAFC9B" blockId="25.[175,1005,194,1019]" box="[199,758,809,831]" pageId="25">
<emphasis id="B90CEAB8FFCA481D28CFBF24264AFC9B" bold="true" box="[199,326,809,831]" pageId="25">Distribution.</emphasis>
Presently known only from
<typeStatus id="54C38808FFCA481D2A78BF242590FC9B" box="[624,668,809,831]" pageId="25">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBF492451FCFE" box="[199,861,836,858]" pageId="25" type="etymology">
<paragraph id="8BC736AAFFCA481D28CFBF492451FCFE" blockId="25.[175,1005,194,1019]" box="[199,861,836,858]" pageId="25">
<emphasis id="B90CEAB8FFCA481D28CFBF492636FCFE" bold="true" box="[199,314,836,858]" pageId="25">Etymology.</emphasis>
The new species is named for
<collectingRegion id="49BCF848FFCA481D2A72BF4925A7FCFE" box="[634,683,836,858]" country="Indonesia" name="Jawa Timur" pageId="25">Java</collectingRegion>
, its
<typeStatus id="54C38808FFCA481D2ADFBF49240FFCFE" box="[727,771,836,858]" pageId="25">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3626521FFCA481D28CFBF6D24CFFC5F" pageId="25" type="discussion">
<paragraph id="8BC736AAFFCA481D28CFBF6D24CFFC5F" blockId="25.[175,1005,194,1019]" pageId="25">
<emphasis id="B90CEAB8FFCA481D28CFBF6D262BFCD2" bold="true" box="[199,295,864,886]" pageId="25">Remarks.</emphasis>
Members of this new species are similar to those of
<taxonomicName id="4C784D29FFCA481D2B53BF6D27FEFC35" authority="(Klug)" baseAuthorityName="Klug" baseAuthorityYear="1834" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="B90CEAB8FFCA481D2B53BF6D24E0FCD1" box="[859,1004,864,885]" italics="true" pageId="25">D. geniculatus</emphasis>
(Klug)
</taxonomicName>
in having tumid genae, green body luster, and similar femoral and tibial color, but differ in having the ratio of Ant 1/Ant 3 smaller and the lamella of aedeagus not thickened (
<figureCitation id="13432A2FFFCA481D29A4BFBD26F5FC62" box="[428,505,944,966]" captionStart="Figs" captionStartId="10.[167,206,1115,1134]" captionTargetBox="[147,968,194,1092]" captionTargetId="figure-214@10.[147,974,194,1093]" captionTargetPageId="10" captionText="Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5461688" httpUri="https://zenodo.org/record/5461688/files/figure.png" pageId="25">Fig. 90</figureCitation>
). They are also similar to adults of
<taxonomicName id="4C784D29FFCA481D2B6ABFBD261DFC44" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1872" class="Insecta" family="Carabidae" genus="Dendrocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="ternatensis">
<emphasis id="B90CEAB8FFCA481D2B6ABFBD24E0FC61" box="[866,1004,944,965]" italics="true" pageId="25">D. ternatensis</emphasis>
Chaudoir
</taxonomicName>
in the form of male genitalia, but differ from the latter in femoral color and in having more tumid genae (
<figureCitation id="13432A2FFFCA481D2A18BFE82557FC5F" box="[528,603,997,1019]" captionStart="Figs" captionStartId="6.[167,206,1502,1521]" captionTargetBox="[191,931,732,1478]" captionTargetId="figure-42@6.[188,932,731,1479]" captionTargetPageId="6" captionText="Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5461676" httpUri="https://zenodo.org/record/5461676/files/figure.png" pageId="25">Fig. 34</figureCitation>
) and smaller ratio of Ant 1/Ant 3.
</paragraph>
</subSubSection>
</treatment>
</document>
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