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<document id="D25EC764E6DEE16773F435AB2819D400" ID-DOI="10.17161/jom.v0i78.7488" ID-ISSN="2325-4467" ID-Zenodo-Dep="8057375" ID-ZooBank="urn:lsid:zoobank.org:pub:6B0F82BE-DFB9-479A-B3CA-A8BDA14CE3CD" checkinTime="1687209762631" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2018" docId="C54B87F1FFDDFF81FE26FA997A9FFA6D" docLanguage="en" docName="JMelittology.2018.78.1-13.pdf" docOrigin="Journal of Melitology 2018 (78)" docSource="http://dx.doi.org/10.17161/jom.v0i78.7488" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docTitle="Varthemapistra Engel 2018, new genus" docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="3972FF89FFDCFF85FFF6FFAE7B24FFED" masterDocTitle="A new genus of anthophorine bees from Brunei (Hymenoptera: Apidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1687210098996" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="3F12F09DED7CB044BB1FB95D839320C6">
<mods:title id="0982CA95E17BDE4508E21A06FF2AA0D2">A new genus of anthophorine bees from Brunei (Hymenoptera: Apidae)</mods:title>
</mods:titleInfo>
<mods:name id="33D5EA9C6651F118B84CFB90D6E6A9F1" type="personal">
<mods:role id="643A9126951F2EDD44B1ABC54215F002">
<mods:roleTerm id="4F2DBD72BAE8D6B035F5CA7A0B07F84E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="48AD437487FD6673E5BD245315692AA3">Engel, Michael S.</mods:namePart>
<mods:affiliation id="F82648A9E194D07E7C20308893ED42CE">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu).</mods:affiliation>
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<mods:typeOfResource id="43B315EB7968F47FAF83988B2D4D2F55">text</mods:typeOfResource>
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<mods:title id="D0883523765C9B58B963AD7B62A08788">Journal of Melitology</mods:title>
</mods:titleInfo>
<mods:part id="B7A6FD247D53F1A209B629D27C62FAA9">
<mods:date id="42F7D0888BC57F7476B5D82991AC4E9A">2018</mods:date>
<mods:detail id="A4E378FDF22420407D3140E7BD90AA7A" type="pubDate">
<mods:number id="1E3B4DD74B01907AF396045AB0D90FA9">2018-05-11</mods:number>
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<mods:detail id="B680C2232CE539CD965A1B77100B56FE" type="volume">
<mods:number id="EFCAF172026523C2C00C6DCDB20D0726">2018</mods:number>
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<mods:detail id="357EB2241ADBE88EAB60C3F055F8E586" type="issue">
<mods:number id="D5191EF930680B057179FCE4F4D6342C">78</mods:number>
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<mods:start id="3A611F815B150A2A40F7B6FACD90097D">1</mods:start>
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<mods:url id="9BF73FE869BE31D398355FA013C9FCEA">http://dx.doi.org/10.17161/jom.v0i78.7488</mods:url>
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<mods:classification id="69808AB8C7BD33827271B7968BFF0B09">journal article</mods:classification>
<mods:identifier id="BD4F931C9A6D7A0DDF45FB1F44A783B1" type="DOI">10.17161/jom.v0i78.7488</mods:identifier>
<mods:identifier id="D129A72C3FF3E4D144AEA6D6B424A18A" type="ISSN">2325-4467</mods:identifier>
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<mods:identifier id="AB529AC6EEE1A5E345519E30D3F369A8" type="ZooBank">urn:lsid:zoobank.org:pub:6B0F82BE-DFB9-479A-B3CA-A8BDA14CE3CD</mods:identifier>
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<treatment id="C54B87F1FFDDFF81FE26FA997A9FFA6D" LSID="urn:lsid:plazi:treatment:C54B87F1FFDDFF81FE26FA997A9FFA6D" httpUri="http://treatment.plazi.org/id/C54B87F1FFDDFF81FE26FA997A9FFA6D" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
<subSubSection id="05F8656CFFDDFF84FE26FA997F64FA82" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="4D5D36E7FFDDFF84FE26FA997845FABD" blockId="1.[464,865,1335,1360]" box="[464,865,1335,1360]" pageId="1" pageNumber="2">
<heading id="1615818BFFDDFF84FE26FA997845FABD" box="[464,865,1335,1360]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="8AE24D64FFDDFF84FE26FA9979F2FAA2" ID-CoL="85ZG" authority="Engel" authorityName="Engel" authorityYear="2018" box="[464,726,1335,1359]" class="Insecta" family="Apidae" genus="Varthemapistra" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="new genus">
<emphasis id="7F96EAF5FFDDFF84FE26FA9979AEFAA2" bold="true" box="[464,650,1335,1359]" italics="true" pageId="1" pageNumber="2">Varthemapistra</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="64A5578EFFDDFF84FD17FA967845FABD" box="[737,865,1336,1360]" pageId="1" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="4D5D36E7FFDDFF84FF06FAF77F64FA82" blockId="1.[240,1088,1369,1391]" box="[240,1088,1369,1391]" pageId="1" pageNumber="2">
ZooBank:
<uri id="39733AE5FFDDFF84FEABFAF77F64FA82" box="[349,1088,1369,1391]" pageId="1" pageNumber="2">
urn:lsid:zoobank.org:act:
<uuid id="39440C32FFDDFF84FDA5FAF77F64FA82" box="[595,1088,1369,1391]" pageId="1" pageNumber="2">CED25625-5914-4B08-9D73-8A3EAC58DFD6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="05F8656CFFDDFF84FF26FA36787AFA42" box="[208,862,1431,1456]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="4D5D36E7FFDDFF84FF26FA36787AFA42" blockId="1.[160,1168,1431,1776]" box="[208,862,1431,1456]" pageId="1" pageNumber="2">
<materialsCitation id="FD8A3CBAFFDDFF84FF26FA36787EFA42" box="[208,858,1431,1456]" collectionCode="SPECIES" pageId="1" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="92598845FFDDFF84FF26FA367A22FA42" box="[208,262,1432,1455]" pageId="1" pageNumber="2">TYPE</typeStatus>
<collectionCode id="2BF3AE22FFDDFF84FEFBFA367A47FA5D" box="[269,355,1432,1456]" pageId="1" pageNumber="2">SPECIES</collectionCode>
:
<taxonomicName id="8AE24D64FFDDFF84FE9AFA39799BFA42" authority="Engel" authorityName="Engel" authorityYear="2018" box="[364,703,1431,1455]" class="Insecta" family="Apidae" genus="Varthemapistra" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="edentata" status="new species">
<emphasis id="7F96EAF5FFDDFF84FE9AFA397956FA42" box="[364,626,1431,1455]" italics="true" pageId="1" pageNumber="2">Varthemapistra edentata</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="64A5578EFFDDFF84FD3FFA36787EFA42" box="[713,858,1431,1455]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="05F8656CFFDDFF84FF26FA1678B7F962" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="4D5D36E7FFDDFF84FF26FA1678B7F962" blockId="1.[160,1168,1431,1776]" pageId="1" pageNumber="2">
DIAGNOSIS: The new genus can be distinguished in the female from other anthophorines by the simple mandible, lacking any preapical teeth (
<figureCitation id="D5D92A62FFDDFF84FCAFFA7978BEFA1D" box="[857,922,1495,1520]" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
) (note that this is not the result of mandibular wear:
<emphasis id="7F96EAF5FFDDFF84FDE4FA5979A5F9E2" box="[530,641,1527,1551]" italics="true" pageId="1" pageNumber="2">vide etiam</emphasis>
Discussion,
<emphasis id="7F96EAF5FFDDFF84FCE1FA56786FF9FD" box="[791,843,1528,1552]" italics="true" pageId="1" pageNumber="2">infra</emphasis>
). In addition the combination of a weakly protuberant clypeus, linear malar space, metatibia with setae on outer surface distinctly fuscous medially, and forewing with the anterior border of the third submarginal cell only slightly shorter than the posterior border further serve to characterize the genus among other
<taxonomicName id="8AE24D64FFDDFF84FDE1F9D97998F962" box="[535,700,1655,1679]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Anthophorini">Anthophorini</taxonomicName>
, in the female sex.
</paragraph>
</subSubSection>
<subSubSection id="05F8656CFFDDFF81FF26F9367A0BFB6D" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="description">
<paragraph id="4D5D36E7FFDDFF81FF26F93678D6FB8D" blockId="1.[160,1168,1431,1776]" lastBlockId="4.[160,1168,744,1408]" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
DESCRIPTION:
<specimenCount id="5BE4FD6EFFDDFF84FE82F9357AACF942" box="[372,392,1691,1711]" pageId="1" pageNumber="2" type="female">♀</specimenCount>
: Medium-sized anthophorines (total length approximately
<quantity id="8A1A9B02FFDDFF84FB97F9367BF6F93D" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="2" unit="mm" value="11.2">11.2 mm</quantity>
), with broad areas of yellow maculation on face (refer to species description,
<emphasis id="7F96EAF5FFDDFF84FBA5F9167FA3F93D" box="[1107,1159,1720,1744]" italics="true" pageId="1" pageNumber="2">infra</emphasis>
) (
<figureCitation id="D5D92A62FFDDFF84FF5EF9797BCFF91D" box="[168,235,1751,1776]" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
). Mandible simple, without preapical teeth (
<figureCitation id="D5D92A62FFDDFF84FD09F9797866F91D" box="[767,834,1751,1776]" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
); malar space linear (
<figureCitation id="D5D92A62FFDDFF87FBCBF9797B8BFA4A" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Figs. 3, 6</figureCitation>
); labrum broader than long, margin not thickened; clypeus weakly protuberant so that, in profile, lower anterior surface is in front of lower inner compound eye orbit by a distance of about one-half compound eye width in profile (
<figureCitation id="D5D92A62FFDEFF87FCA8FA6178BAFA0A" box="[862,926,1487,1511]" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
) [similar to
<taxonomicName id="8AE24D64FFDEFF87FBDFFA617A6DF9EA" authority="Lieftinck" authorityName="Lieftinck" authorityYear="1974" class="Insecta" family="Apidae" genus="Habrophorula" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFDEFF87FBDFFA617BFCF9EA" italics="true" pageId="2" pageNumber="3">Habrophorula</emphasis>
Lieftinck
</taxonomicName>
(
<figureCitation id="D5D92A62FFDEFF87FEACFA417A89F9EA" box="[346,429,1519,1543]" captionStart="Figures 20–23" captionStartId="9.[160,241,1028,1050]" captionTargetBox="[172,1154,248,1009]" captionTargetId="figure-235@9.[160,1168,224,1022]" captionTargetPageId="9" captionText="Figures 20–23. Representative anthophorine bees. 20. Habrophorula nubilipennis (Cockerell), lateral habitus. 21. H. nubilipennis, facial view. 22. Habropoda miserabilis (Cresson), lateral habitus. 23. H. miserabilis, facial view." figureDoi="http://doi.org/10.5281/zenodo.8057389" httpUri="https://zenodo.org/record/8057389/files/figure.png" pageId="2" pageNumber="3">Fig. 20</figureCitation>
), while
<taxonomicName id="8AE24D64FFDEFF87FDFBFA417820F9EA" authority="Lieftinck" authorityName="Lieftinck" authorityYear="1966" box="[525,772,1519,1543]" class="Insecta" family="Apidae" genus="Elaphropoda" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFDEFF87FDFBFA4179B0F9EA" box="[525,660,1519,1543]" italics="true" pageId="2" pageNumber="3">Elaphropoda</emphasis>
Lieftinck
</taxonomicName>
has clypeus greatly protuberant, with lower anterior surface in front of compound eye by compound eye width or more (
<figureCitation id="D5D92A62FFDEFF87FF5EF9817BDFF9AA" box="[168,251,1583,1607]" captionStart="Figures 16–19" captionStartId="8.[160,241,1216,1238]" captionTargetBox="[180,1155,244,1197]" captionTargetId="figure-166@8.[160,1168,224,1210]" captionTargetPageId="8" captionText="Figures 16–19. Representative anthophorine bees. 16. Amegilla quadrifasciata (de Villers), lateral habitus. 17. A. quadrifasciata, facial view. 18. Elaphropoda khasiana (Schulz), lateral habitus. 19. E. khasiana, facial view." figureDoi="http://doi.org/10.5281/zenodo.8057387" httpUri="https://zenodo.org/record/8057387/files/figure.png" pageId="2" pageNumber="3">Fig. 18</figureCitation>
)]; surface of clypeus weakly and gently rounded; anterior margin of clypeus extending to base of mandible; first flagellomere slightly less than one-half length of scape; first flagellomere length about twice apical width; ocelli arranged in broad isosceles triangle, lateral ocelli closer to median ocellus than to each other; proboscis reaching beyond metacoxae in repose; galea length just under twice as long as head (
<figureCitation id="D5D92A62FFDEFF87FB95F9017B8BF90A" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); maxillary palpus short, shorter than stipes (subequal to stipes in
<taxonomicName id="8AE24D64FFDEFF87FC4BF9617F60F90A" authorityName="Lieftinck" authorityYear="1966" box="[957,1092,1743,1767]" class="Insecta" family="Apidae" genus="Elaphropoda" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFDEFF87FC4BF9617F60F90A" box="[957,1092,1743,1767]" italics="true" pageId="2" pageNumber="3">Elaphropoda</emphasis>
</taxonomicName>
), with five palpomeres; maxillary palpomere II rounded in cross-section, with scattered setae (lacking setal fringe); maxillary palpomere III about one-half length of palpomere II; glossa with flabellum present at apex (absent in
<taxonomicName id="8AE24D64FFDFFF86FD05FA117898FA3A" authority="Smith" authorityName="Smith" authorityYear="1854" box="[755,956,1471,1495]" class="Insecta" family="Apidae" genus="Habropoda" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFDFFF86FD05FA11784CFA3A" box="[755,872,1471,1495]" italics="true" pageId="3" pageNumber="4">Habropoda</emphasis>
Smith
</taxonomicName>
). Mesoscutellum slightly overhanging metanotum. Legs slender; coxae not spined; protrochanter with longitudinal row of hooked bristles ventrally along posterior length; metabasitibial plate well-developed and comparatively short, about as long as wide (similar to
<taxonomicName id="8AE24D64FFDFFF86FBACF9B17A2DF9BA" authorityName="Lieftinck" authorityYear="1974" class="Insecta" family="Apidae" genus="Habrophorula" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFDFFF86FBACF9B17A2DF9BA" italics="true" pageId="3" pageNumber="4">Habrophorula</emphasis>
</taxonomicName>
), apical margin pointed (
<figureCitation id="D5D92A62FFDFFF86FDC3F991795CF9BA" box="[565,632,1599,1623]" captionStart="Figures 3–6" captionStartId="3.[160,241,1321,1343]" captionTargetBox="[168,1162,244,1298]" captionTargetId="figure-160@3.[160,1168,224,1315]" captionTargetPageId="3" captionText="Figures 3–6. Holotype female of Varthemapistra edentata, new genus and species, from Brunei. 3. Facial view. 4. Mandible. 5. Metabasitibial plate. 6. Profile of head." figureDoi="http://doi.org/10.5281/zenodo.8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
); scopa rather short and sparse, setae simple, white except medially fuscous; pretarsal claws cleft, arolia present. Forewing with 1cu- a confluent with basal vein; pterostigma twice as long as wide (
<figureCitation id="D5D92A62FFDFFF86FC6DF9D178FBF97A" box="[923,991,1663,1687]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); marginal cell moderately elongate, about 4.5 times as long as wide; distance from marginal cell apex to wing tip shorter than marginal cell (
<figureCitation id="D5D92A62FFDFFF86FD85F9117993F93A" box="[627,695,1727,1751]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); free part of marginal cell shorter than bounded portion of marginal cell, about as long as anterior border of third submarginal cell (
<figureCitation id="D5D92A62FFD8FF81FF05FD467A64FCED" box="[243,320,744,768]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="4" pageNumber="5">Figs. 7</figureCitation>
,
<figureCitation id="D5D92A62FFD8FF81FEBAFD467A4DFCED" box="[332,361,744,768]" captionStart="Figures 8–15" captionStartId="7.[160,241,1291,1313]" captionTargetBox="[160,1161,224,1284]" captionTargetId="figure-128@7.[160,1168,224,1284]" captionTargetPageId="7" captionText="Figures 8–15. Wing venation from representative species of Anthophorini. 8. Anthophora (Lophanthophora) dammersi Timberlake. 9. Amegilla quadrifasciata (de Villers). 10. Habropoda depressa Fowler. 11. Elaphropoda percarinata (Cockerell). 12. Pachymelus (Pachymelus) unicolor (Saussure). 13. Deltoptila sp. 14. Habrophorula nubilipennis (Cockerell). 15. Varthemapistra edentata, new genus and species." figureDoi="http://doi.org/10.5281/zenodo.8057383" httpUri="https://zenodo.org/record/8057383/files/figure.png" pageId="4" pageNumber="5">15</figureCitation>
); first submarginal cell shorter than combined lengths of second and third submarginal cells (as measured along posterior borders); second submarginal cell shorter than either first or third submarginal cells, slightly narrowed anteriorly; 1m-cu terminating near apex of second submarginal cell (
<figureCitation id="D5D92A62FFD8FF81FCCAFCE67859FC8D" box="[828,893,840,864]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
); third submarginal cell broad anteriorly, with anterior border slightly shorter than posterior border (anterior border about three-quarters length of posterior border:
<figureCitation id="D5D92A62FFD8FF81FCC5FC2678A6FC4D" box="[819,898,904,928]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="4" pageNumber="5">Figs. 7</figureCitation>
,
<figureCitation id="D5D92A62FFD8FF81FC79FC26788EFC4D" box="[911,938,904,928]" captionStart="Figures 8–15" captionStartId="7.[160,241,1291,1313]" captionTargetBox="[160,1161,224,1284]" captionTargetId="figure-128@7.[160,1168,224,1284]" captionTargetPageId="7" captionText="Figures 8–15. Wing venation from representative species of Anthophorini. 8. Anthophora (Lophanthophora) dammersi Timberlake. 9. Amegilla quadrifasciata (de Villers). 10. Habropoda depressa Fowler. 11. Elaphropoda percarinata (Cockerell). 12. Pachymelus (Pachymelus) unicolor (Saussure). 13. Deltoptila sp. 14. Habrophorula nubilipennis (Cockerell). 15. Varthemapistra edentata, new genus and species." figureDoi="http://doi.org/10.5281/zenodo.8057383" httpUri="https://zenodo.org/record/8057383/files/figure.png" pageId="4" pageNumber="5">15</figureCitation>
) (borders subequal in
<taxonomicName id="8AE24D64FFD8FF81FF4BFC067A76FC2D" authorityName="Lieftinck" authorityYear="1974" box="[189,338,936,960]" class="Insecta" family="Apidae" genus="Habrophorula" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="7F96EAF5FFD8FF81FF4BFC067A76FC2D" box="[189,338,936,960]" italics="true" pageId="4" pageNumber="5">Habrophorula</emphasis>
</taxonomicName>
and more distinctly narrowed anteriorly in other Asiatic genera); 2rs-m gently arched but not sinuate (sinuate form results in anteriorly narrowed third submarginal cell) (
<figureCitation id="D5D92A62FFD8FF81FEBBFC467ABDFBED" box="[333,409,1000,1024]" captionStart="Figure 7" captionStartId="4.[160,231,681,703]" captionTargetBox="[185,1158,239,660]" captionTargetId="figure-363@4.[160,1168,224,677]" captionTargetPageId="4" captionText="Figure 7. Wing venation of Varthemapistra edentata, new genus and species, from Brunei." figureDoi="http://doi.org/10.5281/zenodo.8057381" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="4" pageNumber="5">Figs. 7</figureCitation>
,
<figureCitation id="D5D92A62FFD8FF81FE52FC467AE4FBED" box="[420,448,1000,1024]" captionStart="Figures 8–15" captionStartId="7.[160,241,1291,1313]" captionTargetBox="[160,1161,224,1284]" captionTargetId="figure-128@7.[160,1168,224,1284]" captionTargetPageId="7" captionText="Figures 8–15. Wing venation from representative species of Anthophorini. 8. Anthophora (Lophanthophora) dammersi Timberlake. 9. Amegilla quadrifasciata (de Villers). 10. Habropoda depressa Fowler. 11. Elaphropoda percarinata (Cockerell). 12. Pachymelus (Pachymelus) unicolor (Saussure). 13. Deltoptila sp. 14. Habrophorula nubilipennis (Cockerell). 15. Varthemapistra edentata, new genus and species." figureDoi="http://doi.org/10.5281/zenodo.8057383" httpUri="https://zenodo.org/record/8057383/files/figure.png" pageId="4" pageNumber="5">15</figureCitation>
); hind wing cu-a about half as long as 2M+Cu, oblique relative to M+Cu; 2M+Cu slightly less than one half as long as
<specimenCount id="5BE4FD6EFFD8FF81FCEAFBA6786DFBCD" box="[796,841,1032,1056]" pageId="4" pageNumber="5" type="generic">1M.</specimenCount>
Metasomal terga II–V with lateral longitudinal parts of graduli present and strong; pygidial plate margins slightly converging apically, lateral margins weakly carinate, apex broadly blunt.
</paragraph>
<caption id="199D666FFFDEFF87FF56FA9C7908FA88" ID-DOI="http://doi.org/10.5281/zenodo.8057377" ID-Zenodo-Dep="8057377" httpUri="https://zenodo.org/record/8057377/files/figure.png" pageId="2" pageNumber="3" startId="2.[160,241,1330,1352]" targetBox="[173,1149,248,1301]" targetPageId="2" targetType="figure">
<paragraph id="4D5D36E7FFDEFF87FF56FA9C7908FA88" blockId="2.[160,1175,1330,1381]" pageId="2" pageNumber="3">
<emphasis id="7F96EAF5FFDEFF87FF56FA9C7A06FAA5" bold="true" box="[160,290,1330,1352]" pageId="2" pageNumber="3">Figures 1–2.</emphasis>
Holotype female of
<taxonomicName id="8AE24D64FFDEFF87FDF3FA9D79D2FAA5" authority="Engel, 2018" authorityName="Engel" authorityYear="2018" box="[517,758,1331,1352]" class="Insecta" family="Apidae" genus="Varthemapistra" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="edentata" status="new genus and species">
<emphasis id="7F96EAF5FFDEFF87FDF3FA9D79D2FAA5" box="[517,758,1331,1352]" italics="true" pageId="2" pageNumber="3">Varthemapistra edentata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="64A5578EFFDEFF87FCF4FA9D78D8FAA4" box="[770,1020,1331,1353]" pageId="2" pageNumber="3" rank="species">new genus and species</taxonomicNameLabel>
, from Brunei.
<emphasis id="7F96EAF5FFDEFF87FF56FAE17B97FA89" bold="true" box="[160,179,1359,1380]" pageId="2" pageNumber="3">1.</emphasis>
Lateral habitus.
<emphasis id="7F96EAF5FFDEFF87FE98FAE17AA5FA89" bold="true" box="[366,385,1359,1380]" pageId="2" pageNumber="3">2.</emphasis>
Dorsal habitus.
</paragraph>
</caption>
<caption id="199D666FFFDFFF86FF56FA8778B4FAB1" ID-DOI="http://doi.org/10.5281/zenodo.8057379" ID-Zenodo-Dep="8057379" httpUri="https://zenodo.org/record/8057379/files/figure.png" pageId="3" pageNumber="4" startId="3.[160,241,1321,1343]" targetBox="[168,1162,244,1298]" targetPageId="3" targetType="figure">
<paragraph id="4D5D36E7FFDFFF86FF56FA8778B4FAB1" blockId="3.[160,1175,1321,1372]" pageId="3" pageNumber="4">
<emphasis id="7F96EAF5FFDFFF86FF56FA877A07FAD3" bold="true" box="[160,291,1321,1343]" pageId="3" pageNumber="4">Figures 3–6.</emphasis>
Holotype female of
<taxonomicName id="8AE24D64FFDFFF86FDF1FA8779D1FAD3" authority="Engel, 2018" authorityName="Engel" authorityYear="2018" box="[519,757,1321,1342]" class="Insecta" family="Apidae" genus="Varthemapistra" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="edentata" status="new genus and species">
<emphasis id="7F96EAF5FFDFFF86FDF1FA8779D1FAD3" box="[519,757,1321,1342]" italics="true" pageId="3" pageNumber="4">Varthemapistra edentata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="64A5578EFFDFFF86FCF5FA8478D8FAD2" box="[771,1020,1321,1343]" pageId="3" pageNumber="4" rank="species">new genus and species</taxonomicNameLabel>
, from Brunei.
<emphasis id="7F96EAF5FFDFFF86FF56FAE87B96FAB6" bold="true" box="[160,178,1350,1371]" pageId="3" pageNumber="4">3.</emphasis>
Facial view.
<emphasis id="7F96EAF5FFDFFF86FEB2FAE87A72FAB6" bold="true" box="[324,342,1350,1371]" pageId="3" pageNumber="4">4.</emphasis>
Mandible.
<emphasis id="7F96EAF5FFDFFF86FE20FAE87ACCFAB6" bold="true" box="[470,488,1350,1371]" pageId="3" pageNumber="4">5.</emphasis>
Metabasitibial plate.
<emphasis id="7F96EAF5FFDFFF86FD22FAE879C2FAB6" bold="true" box="[724,742,1350,1371]" pageId="3" pageNumber="4">6.</emphasis>
Profile of head.
</paragraph>
</caption>
<caption id="199D666FFFD8FF81FF56FD077F62FD52" ID-DOI="http://doi.org/10.5281/zenodo.8057381" ID-Zenodo-Dep="8057381" box="[160,1094,681,703]" httpUri="https://zenodo.org/record/8057381/files/figure.png" pageId="4" pageNumber="5" startId="4.[160,231,681,703]" targetBox="[185,1158,239,660]" targetPageId="4" targetType="figure">
<paragraph id="4D5D36E7FFD8FF81FF56FD077F62FD52" blockId="4.[160,1094,681,703]" box="[160,1094,681,703]" pageId="4" pageNumber="5">
<emphasis id="7F96EAF5FFD8FF81FF56FD077BDBFD53" bold="true" box="[160,255,681,703]" pageId="4" pageNumber="5">Figure 7.</emphasis>
Wing venation of
<taxonomicName id="8AE24D64FFD8FF81FE3EFD077997FD53" authority="Engel, 2018" authorityName="Engel" authorityYear="2018" box="[456,691,681,702]" class="Insecta" family="Apidae" genus="Varthemapistra" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="edentata" status="new genus and species">
<emphasis id="7F96EAF5FFD8FF81FE3EFD077997FD53" box="[456,691,681,702]" italics="true" pageId="4" pageNumber="5">Varthemapistra edentata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="64A5578EFFD8FF81FD36FD077891FD52" box="[704,949,681,703]" pageId="4" pageNumber="5" rank="species">new genus and species</taxonomicNameLabel>
, from Brunei.
</paragraph>
</caption>
<paragraph id="4D5D36E7FFD8FF81FF26FBC57A0BFB6D" blockId="4.[160,1168,744,1408]" box="[208,303,1129,1152]" pageId="4" pageNumber="5">
<specimenCount id="5BE4FD6EFFD8FF81FF26FBC57BC0FB92" box="[208,228,1131,1151]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
:
<emphasis id="7F96EAF5FFD8FF81FF07FBC77A0CFB6D" box="[241,296,1129,1152]" italics="true" pageId="4" pageNumber="5">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="05F8656CFFD8FF81FF26FB267A9FFA6D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="4D5D36E7FFD8FF81FF26FB267A9FFA6D" blockId="4.[160,1168,744,1408]" pageId="4" pageNumber="5">
ETYMOLOGY: The new generic name is a combination of Varthema; after Ludovico di Varthema (1470–1517), the Italian explorer who, among other things, was apparently the earliest to record accounts of
<collectingCountry id="35F57677FFD8FF81FDB0FB6679B1FB0D" box="[582,661,1224,1248]" name="Brunei" pageId="4" pageNumber="5">Brunei</collectingCountry>
based on his possible visit to the then Empire in 1505 (
<bibRefCitation id="29734B16FFD8FF81FEC3FB467AF0FAED" author="Temple, R. C." box="[309,468,1256,1280]" pageId="4" pageNumber="5" refId="ref5354" refString="Temple, R. C. 1928. The Itinerary of Ludovico di Varthema of Bologna from 1501 to 1508 as Translated from the Original Italian Edition of 1510, by John Winter Jones, F. S. A. in 1863 for the Hakluyt Society with a Discourse on Varthema and his Travels in Southern Asia. Argonaut Press; London, UK; lxxxv + 121 pp." type="book" year="1928">Temple, 1928</bibRefCitation>
: it should be noted, however, that while his account accords with a visit to
<collectingCountry id="35F57677FFD8FF81FEA2FAA67A81FACD" box="[340,421,1288,1312]" name="Brunei" pageId="4" pageNumber="5">Brunei</collectingCountry>
, he may have landed elsewhere on Borneo and learned trade details from other ports on the island,
<emphasis id="7F96EAF5FFD8FF81FD92FA8779A7FAAD" box="[612,643,1321,1344]" italics="true" pageId="4" pageNumber="5">e.g</emphasis>
.,
<bibRefCitation id="29734B16FFD8FF81FD61FA867814FAAD" author="Vienne, M. - S." box="[663,816,1320,1344]" pageId="4" pageNumber="5" refId="ref5425" refString="Vienne, M. - S., de. 2015. Brunei: From the Age of Commerce to the 21 st Century. NUS [National University of Singapore] Press; Singapore; xviii + 345 pp." type="book" year="2015">Vienne, 2015</bibRefCitation>
); and the Latin
<emphasis id="7F96EAF5FFD8FF81FC11FA877F36FAAD" box="[999,1042,1321,1344]" italics="true" pageId="4" pageNumber="5">apis</emphasis>
(meaning, “bee”), and –
<emphasis id="7F96EAF5FFD8FF81FECAFAE77A49FA8D" box="[316,365,1353,1376]" italics="true" pageId="4" pageNumber="5">istra</emphasis>
, the diminutive suffix referencing “wild resemblance”. The gender of the name is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>
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