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172 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.840.33326" ID-GBIF-Dataset="363f2c7c-0635-46c8-b598-9e9aa014b6d2" ID-PMC="PMC6482118" ID-Pensoft-Pub="1313-2970-840-35" ID-PubMed="31065227" ID-ZBK="C607C2FDC17644FABBC582D561C6434C" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-840-35" ModsDocOrigin="ZooKeys 840" ModsDocTitle="Conchological and molecular analysis of the “non-scaly” Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1555526082661" checkinUser="pensoft" docAuthor="Zacaery Khalik, Mohd, P. Hendriks, Kasper, J. Vermeulen, Jaap & Menno Schilthuizen," docDate="2019" docId="DCA977919AF9B3BDEF9D709DC50F7733" docLanguage="en" docName="ZooKeys 840: 35-86" docOrigin="ZooKeys 840" docSource="http://dx.doi.org/10.3897/zookeys.840.33326" docTitle="Georissa corrugata Khalik, Hendriks, Vermeulen & Schilthuizen, 2019, sp. n." docType="treatment" docUuid="81610B3D-0DB9-48ED-89D3-DAA69BDD031B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="47" masterDocId="647C1D35F570C105FFFCFFCDB169212D" masterDocTitle="Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="86" masterPageNumber="35" pageNumber="44" updateTime="1668167174253" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zacaery Khalik, Mohd</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>P. Hendriks, Kasper</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>J. Vermeulen, Jaap</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menno Schilthuizen,</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>840</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>35</mods:start>
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<mods:end>86</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.840.33326</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.840.33326</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-840-35</mods:identifier>
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<mods:identifier type="ZBK">C607C2FDC17644FABBC582D561C6434C</mods:identifier>
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<mods:identifier type="ZooBank">C607C2FDC17644FABBC582D561C6434C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156787936" LSID="urn:lsid:zoobank.org:act:81610B3D-0DB9-48ED-89D3-DAA69BDD031B" httpUri="http://treatment.plazi.org/id/DCA977919AF9B3BDEF9D709DC50F7733" lastPageId="12" lastPageNumber="47" pageId="9" pageNumber="44">
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<subSubSection pageId="9" pageNumber="44" type="nomenclature">
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<paragraph pageId="9" pageNumber="44">
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<taxonomicName LSID="http://zoobank.org/81610B3D-0DB9-48ED-89D3-DAA69BDD031B" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa corrugata" order="Cycloneritida" pageId="9" pageNumber="44" phylum="Mollusca" rank="species" species="corrugata">Georissa corrugata</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="44">sp. n.</taxonomicNameLabel>
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Figure 6
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<normalizedToken originalValue="A–I">A-I</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="45" type="type locality">
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<paragraph pageId="10" pageNumber="45">
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<pageBreakToken pageId="10" pageNumber="45" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph pageId="10" pageNumber="45">
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Batu Tomanggong, Kinabatangan valley, Sandakan, Sabah, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.531">05°31.86'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.304">118°18.24'E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="45" type="type material">
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<paragraph pageId="10" pageNumber="45">Type material.</paragraph>
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<paragraph pageId="10" pageNumber="45">
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Holotype. Batu Tomanggong, Kinabatangan valley, Sandakan, Sabah, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.531">05°31.86'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.304">118°18.24'E</geoCoordinate>
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): MZU/MOL 16.15 (Fig. 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) Paratypes. Batu Tomanggong, Kinabatangan valley, Sandakan, Sabah, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.531">05°31.86'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.304">118°18.24'E</geoCoordinate>
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): MZU/MOL 16.16 (Fig. 6
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<normalizedToken originalValue="D–I">D-I</normalizedToken>
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). Batu Punggul, Sepulut valley, Sabah: JJV 1903 (1). N Borneo: RMNH/MOL 152848. Batu Keruak, Kinabatangan valley, Sabah (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.523">05°31.38'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.285">118°17.10'E</geoCoordinate>
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): BOR/MOL 1467, BOR/MOL 1844, BOR/MOL 11661 (1). Unnamed hill, Kinabatangan valley, Sabah (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.5185">05°31.11'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.28717">118°17.23'E</geoCoordinate>
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): BOR/MOL 2218 (1, juvenile).
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</paragraph>
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<caption pageId="10" pageNumber="45">
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<paragraph pageId="10" pageNumber="45">
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Figure 6.
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa corrugata" order="Cycloneritida" pageId="10" pageNumber="45" phylum="Mollusca" rank="species" species="corrugata">Georissa corrugata</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Holotype: MZU/MOL 16.15
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<normalizedToken originalValue="D–I">D-I</normalizedToken>
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paratype: MZU/MOL 16.16 A, D shell apertural view B shell side view C shell rear view E, F shell cross-section from 3D model G shell top view H protoconch side view I close up of protoconch from top at 1000
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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); 200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(H); 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(I).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="45" type="etymology">
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<paragraph pageId="10" pageNumber="45">Etymology.</paragraph>
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<paragraph pageId="10" pageNumber="45">
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The name is derived from a Latin word corrugatus, meaning
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<normalizedToken originalValue="“wrinkled”">"wrinkled"</normalizedToken>
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, referring to the coarse and irregular radial sculpture.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="46" pageId="10" pageNumber="45" type="description">
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<paragraph pageId="10" pageNumber="45">Description.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="46" pageId="10" pageNumber="45">
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Protoconch. Colour: white. Sculpture pattern: parallel lines of varying width, widening before splitting in two. Mesh width: 3.1-6.2
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, with the distance between each of the sculptural units as wide as the mesh width. Teleoconch. Colour: white. First whorl: rounded. Subsequent whorls: rounded. Suture: clearly impressed. Shoulder: narrow. Number of whorls: 2
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<normalizedToken originalValue="½–">1/2-</normalizedToken>
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3. SH: 1.14-1.43 mm. SW: 1.01-1.11 mm. SI: 1.10-1.29. Shell sculpture. Radial sculpture: present, more prominent after the first whorl, wavy, irregular and widely spaced, often distinctly higher and strongly projected on and above the periphery. Spiral sculpture: present, regularly spaced, thin, discontinuous due the interruption by radial sculpture. Aperture. Shape: rounded to slightly ovoid, parietal side straight, palatal edge contiguous with the parietal side, palatal and
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<pageBreakToken pageId="11" pageNumber="46" start="start">basal</pageBreakToken>
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sides convex. AH: 0.82-1.07 mm. AW: 1.00-1.09 mm. AI: 0.75-1.02. Holotype dimensions. SH: 1.71 mm, SW: 0.99 mm, AH: 0.52 mm, AW: 0.58 mm.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="47" pageId="11" pageNumber="46" type="diagnosis">
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<paragraph pageId="11" pageNumber="46">Diagnosis.</paragraph>
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<paragraph lastPageId="12" lastPageNumber="47" pageId="11" pageNumber="46">
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The wavy and irregular, widely spaced and strong radial sculpture, with thin regularly arranged spiral sculpture in between is diagnostic for
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<taxonomicName lsidName="G. corrugata" pageId="11" pageNumber="46" rank="species" species="corrugata">G. corrugata</taxonomicName>
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.
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa similis" order="Cycloneritida" pageId="11" pageNumber="46" phylum="Mollusca" rank="species" species="similis">Georissa similis</taxonomicName>
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and
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<taxonomicName lsidName="G. borneensis" pageId="11" pageNumber="46" rank="species" species="borneensis">G. borneensis</taxonomicName>
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have a somewhat similar arrangement of radial and spiral sculpture, but do not carry the protoconch sculpture consisting of parallel lines. The shell shape and size of
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<taxonomicName lsidName="G. corrugata" pageId="11" pageNumber="46" rank="species" species="corrugata">G. corrugata</taxonomicName>
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are similar to
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<taxonomicName lsidName="G. similis" pageId="11" pageNumber="46" rank="species" species="similis">G. similis</taxonomicName>
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. Besides the difference
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<pageBreakToken pageId="12" pageNumber="47" start="start">in</pageBreakToken>
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the protoconch sculpture, the latter species also has more densely arranged radial sculpture on the teleoconch.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="47" type="distribution">
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<paragraph pageId="12" pageNumber="47">Distribution.</paragraph>
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<paragraph pageId="12" pageNumber="47">
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa corrugata" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="corrugata">Georissa corrugata</taxonomicName>
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is distributed on the limestone hills of the lower Kinabatangan valley, known to occur from Gomantong to Batu Tomanggong, but always in low densities compared to other
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
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species. The species has also been found in the Sepulut valley, ca. a hundred km further to the southwest.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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