268 lines
32 KiB
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268 lines
32 KiB
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<document id="98D5E1DA5D4491650C4AC55F1CA02A96" ID-CLB-Dataset="54380" ID-DOI="10.11646/zootaxa.5325.4.9" ID-GBIF-Dataset="7785b7ea-e2de-4668-817f-be4f09371c63" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243787" ID-ZooBank="1E98E639-F78B-41D9-BDE9-39F075A4644F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691943785067" checkinUser="plazi" docAuthor="Guo, Liang, Huang, Si-Yao, Wang, Min & Ma, Fang-Zhou" docDate="2023" docId="038BC051FD34842EFF5139F8A0B0FE0C" docLanguage="en" docName="zootaxa.5325.4.9.pdf" docOrigin="Zootaxa 5325 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5325.4.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chonala houae Lang, Li & Deng 2017" docType="treatment" docVersion="6" lastPageNumber="599" masterDocId="FFB2B829FD378428FFC63C41A30CFFF9" masterDocTitle="Description of a new subspecies of Chonala laurae Bozano, 1999 from Southeastern China (Lepidoptera, Nymphalidae, Satyrinae)" masterLastPageNumber="600" masterPageNumber="593" pageNumber="596" updateTime="1699557162623" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FCEC2F044107D622381EAD99D960471B">Description of a new subspecies of Chonala laurae Bozano, 1999 from Southeastern China (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
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<mods:namePart id="59E49E708E1C16B0A6FDAAEDFC2B7605">Guo, Liang</mods:namePart>
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<mods:affiliation id="0B6A46CECC0BD9767C95750E6D09F88A">Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, Guangdong, China & guoljxty @ 163. com; https: // orcid. org / 0009 - 0004 - 2436 - 6151</mods:affiliation>
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<mods:namePart id="1D3A2C9F8F495E562ED9D324FC8BB086">Huang, Si-Yao</mods:namePart>
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<mods:affiliation id="B427E038A7F31F21CDDB39AD844EAB4C">Arthropoda Department, Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauer Allee 127, Bonn 53113, Germany & S. Huang @ leibniz-lib. de; huangsiyao 2007 @ aliyun. com; https: // orcid. org / 0000 - 0002 - 9859 - 9212</mods:affiliation>
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<mods:nameIdentifier id="D345B6A94D491224C570159537A625EE" type="email">uang@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="B5013FA95B52CF86134A2888B9A25524">Wang, Min</mods:namePart>
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<mods:affiliation id="6C926C909D7AC51CB3C934571AA37B7E">Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, Guangdong, China & Institute of International Rivers and Eco-Security, Yunnan University, Kunming 650500, Yunnan, China & minwang @ scau. edu. cn; https: // orcid. org / 0000 - 0001 - 5834 - 4058</mods:affiliation>
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<mods:nameIdentifier id="C01F3255769ECAC17BDB185D51F2974D" type="email">minwang@scau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="8712D9D812458EB141853C5B56CC1126">Ma, Fang-Zhou</mods:namePart>
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<mods:affiliation id="950D7975CDD1EAD7F5FDD32F30B9E16B">State Environmental Protection Scientific Observation and Research Station for Ecology and Environment of Wuyi Mountains, Ministry of Ecology and Environment, Nanjing, 210042, China & Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment; National Key Laboratory of Biosafety, Nanjing, 210042, China & mfz @ nies. org; https: // orcid. org / 0000 - 0001 - 8219 - 7967</mods:affiliation>
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<treatment id="038BC051FD34842EFF5139F8A0B0FE0C" ID-DOI="http://doi.org/10.5281/zenodo.8243803" ID-GBIF-Taxon="212430266" ID-Zenodo-Dep="8243803" LSID="urn:lsid:plazi:treatment:038BC051FD34842EFF5139F8A0B0FE0C" httpUri="http://treatment.plazi.org/id/038BC051FD34842EFF5139F8A0B0FE0C" lastPageId="6" lastPageNumber="599" pageId="3" pageNumber="596">
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<paragraph id="8B9D7147FD34842BFF5139F8A17BFA2A" blockId="3.[151,631,1465,1527]" box="[151,631,1465,1492]" pageId="3" pageNumber="596">
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<heading id="D0D5C62BFD34842BFF5139F8A17BFA2A" bold="true" box="[151,631,1465,1492]" fontSize="11" level="1" pageId="3" pageNumber="596" reason="1">
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<taxonomicName id="4C220AC4FD34842BFF5139F8A17BFA2A" ID-CoL="BLPCF" authority="Lang, Li & Deng, 2017" authorityName="Lang, Li & Deng" authorityYear="2017" box="[151,631,1465,1492]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="houae">
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<emphasis id="B956AD55FD34842BFF5139F8A17BFA2A" bold="true" box="[151,631,1465,1492]" pageId="3" pageNumber="596">
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<emphasis id="B956AD55FD34842BFF5139F8A25EFA2D" bold="true" box="[151,338,1465,1492]" italics="true" pageId="3" pageNumber="596">Chonala houae</emphasis>
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<bibRefCitation id="EFB30CB6FD34842BFE9F39F8A17BFA2A" author="Lang, S. Y. & Li, A. M. & Deng, H. L." box="[345,631,1465,1491]" pageId="3" pageNumber="596" pagination="35 - 37" refId="ref4926" refString="Lang, S. Y., Li, A. M. & Deng, H. L. (2017) A new species of Chonala Moore, 1893 from Chongqing, SW China (Lepidoptera: Nymphalidae, Satyrinae). Entomologische Zeitschrift, 127 (1), 35 - 37." type="journal article" year="2017">Lang, Li & Deng, 2017</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33822CCFD34842BFF51399CA256FA0E" box="[151,346,1501,1527]" pageId="3" pageNumber="596" type="description">
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<paragraph id="8B9D7147FD34842BFF51399CA256FA0E" blockId="3.[151,631,1465,1527]" box="[151,346,1501,1527]" pageId="3" pageNumber="596">
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<figureCitation id="13196DC2FD34842BFF51399CA205FA0E" box="[151,265,1501,1527]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-21@4.[203,1384,181,1919]" captionTargetPageId="4" captionText="FIGURES 9–16. Adults of Chonala spp. 9–10, C. houae, holotype, Mt. Jinfo, Chongqing, SATY1156 (CQMNH); 11–12, ditto, paratype, ditto, SATY0466 (CQMNH); 13–14, C. laurae wuyishana ssp. nov., paratype, Mt. Wuyi, Fujian (SCAU); 15–16, C. laurae laurae, Shennongjia, Hubei (CSYL); 9–12 male, 13–16 female. Scale=10 mm." figureDoi="http://doi.org/10.5281/zenodo.8243793" httpUri="https://zenodo.org/record/8243793/files/figure.png" pageId="3" pageNumber="596">Figs 9–12</figureCitation>
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,
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<figureCitation id="13196DC2FD34842BFED2399CA256FA0E" box="[276,346,1501,1527]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[197,1391,194,1907]" captionTargetId="figure-20@5.[184,1403,181,1919]" captionTargetPageId="5" captionText="FIGURES 17–20. Male genitalia of Chonala sspp. Depository of the genitalia: 17 in SCAU, 18 in CSYL, 19 & 20 in CQMNH. a=genitalia capsule with left valva and aedeagus removed; b=aedeagus in dorsal view; c=aedeagus in lateral view; d=left valva in dorsal view. Scale=1 mm." figureDoi="http://doi.org/10.5281/zenodo.8243795" httpUri="https://zenodo.org/record/8243795/files/figure.png" pageId="3" pageNumber="596">19, 20</figureCitation>
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</paragraph>
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<subSubSection id="C33822CCFD34842BFF513A63A72AF9C3" box="[151,1062,1570,1594]" pageId="3" pageNumber="596" type="reference_group">
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<paragraph id="8B9D7147FD34842BFF513A63A72AF9C3" blockId="3.[151,1062,1570,1594]" box="[151,1062,1570,1594]" pageId="3" pageNumber="596">
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<treatmentCitationGroup id="AB325669FD34842BFF513A63A72AF9C3" box="[151,1062,1570,1594]" pageId="3" pageNumber="596">
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<treatmentCitation id="0A835756FD34842BFF513A63A143F9C3" author="Lang, S. Y. & Li, A. M. & Deng, H. L." box="[151,591,1570,1594]" page="35" pageId="3" pageNumber="596" year="2017">
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<taxonomicName id="4C220AC4FD34842BFF513A63A143F9C3" ID-CoL="BLPCF" authority="Lang, Li & Deng, 2017: 35" authorityName="Lang, Li & Deng" authorityPageNumber="35" authorityYear="2017" box="[151,591,1570,1594]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="houae">
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<emphasis id="B956AD55FD34842BFF513A63A23FF9C3" box="[151,307,1570,1594]" italics="true" pageId="3" pageNumber="596">Chonala houae</emphasis>
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<bibRefCitation id="EFB30CB6FD34842BFEFF3A63A143F9C3" author="Lang, S. Y. & Li, A. M. & Deng, H. L." box="[313,591,1570,1594]" pageId="3" pageNumber="596" pagination="35 - 37" refId="ref4926" refString="Lang, S. Y., Li, A. M. & Deng, H. L. (2017) A new species of Chonala Moore, 1893 from Chongqing, SW China (Lepidoptera: Nymphalidae, Satyrinae). Entomologische Zeitschrift, 127 (1), 35 - 37." type="journal article" year="2017">Lang, Li & Deng, 2017: 35</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 1, 2 & 9; Lang 2017: 115, pl. XIV, fig. 4.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33822CCFD34842BFF513A20A146F93A" pageId="3" pageNumber="596" type="materials_examined">
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<paragraph id="8B9D7147FD34842BFF513A20A146F93A" blockId="3.[151,1437,1633,2020]" pageId="3" pageNumber="596">
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<emphasis id="B956AD55FD34842BFF513A20A28EF982" bold="true" box="[151,386,1633,1659]" pageId="3" pageNumber="596">Material examined:</emphasis>
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<materialsCitation id="3B4A7B1AFD34842BFE4E3A20A76CF966" ID-GBIF-Occurrence="4174564301" collectingDate="2007-07-04" collectionCode="CQMNH" collectorName="Xiaodong Yang" country="China" county="Photos" elevation="1750" location="Nanchuan" municipality="Nanchuan" pageId="3" pageNumber="596" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="holotype">
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Photos:
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<typeStatus id="5499CFE5FD34842BFE243A20A14BF982" box="[482,583,1633,1659]" pageId="3" pageNumber="596" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="9D24BACEFD34842BFD963A20A185F982" box="[592,649,1633,1659]" pageId="3" pageNumber="596" type="male">male</specimenCount>
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,
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<collectingCountry id="F33531D7FD34842BFD523A20A1D6F982" box="[660,730,1633,1659]" name="China" pageId="3" pageNumber="596">China</collectingCountry>
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,
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<collectingRegion id="49E6BFA5FD34842BFD223A20A06FF982" box="[740,867,1633,1659]" country="China" name="Chongqing" pageId="3" pageNumber="596">Chongqing</collectingRegion>
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,
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<collectingMunicipality id="6BF9EB3DFD34842BFCAB3A20A0EDF982" box="[877,993,1633,1659]" pageId="3" pageNumber="596">Nanchuan</collectingMunicipality>
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, South slope of Mt. Jinfo, N29°00.367’, E107°11.858’,
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<quantity id="4CDADCA2FD34842BFE863AC4A2D1F959" box="[320,477,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="596" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">
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<elevation id="000F9674FD34842BFE863AC4A2D1F959" box="[320,477,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="596" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">1500–2000 m</elevation>
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</quantity>
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,
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<date id="FF9C5787FD34842BFE2E3AC4A169F966" box="[488,613,1669,1695]" pageId="3" pageNumber="596" value="2007-07-04">
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<collectingDate id="EFD8AE6FFD34842BFE2E3AC4A169F966" box="[488,613,1669,1695]" pageId="3" pageNumber="596" value="2007-07-04">4.VII.2007</collectingDate>
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</date>
|
|||
|
|
, leg.
|
|||
|
|
<collectorName id="26D71491FD34842BFD663AC4A05CF959" box="[672,848,1669,1696]" pageId="3" pageNumber="596">Xiaodong Yang</collectorName>
|
|||
|
|
, SATY1156 (
|
|||
|
|
<collectionCode id="ED33E982FD34842BFC203AC4A759F966" box="[998,1109,1669,1695]" pageId="3" pageNumber="596">CQMNH</collectionCode>
|
|||
|
|
);
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="3B4A7B1AFD34842BFBA13AC7A14AF93A" ID-GBIF-Occurrence="4174564302" collectingDate="2007-07-04" collectionCode="CQMNH" collectorName="Mt. Jinfo, N & Xiaodong Yang" country="China" county="Photos" elevation="1750" location="South" municipality="Nanchuan" pageId="3" pageNumber="596" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="paratype">
|
|||
|
|
<typeStatus id="5499CFE5FD34842BFBA13AC7A7C0F959" box="[1127,1228,1670,1696]" pageId="3" pageNumber="596" type="paratype">paratype</typeStatus>
|
|||
|
|
:
|
|||
|
|
<specimenCount id="9D24BACEFD34842BFB133AC4A62FF966" box="[1237,1315,1669,1695]" pageId="3" pageNumber="596" type="male">1 male</specimenCount>
|
|||
|
|
, same data as in holotype, SATY0466 (
|
|||
|
|
<collectionCode id="ED33E982FD34842BFE093AE8A132F93A" box="[463,574,1705,1731]" pageId="3" pageNumber="596">CQMNH</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C33822CCFD34842EFF013A8CA16FFE28" lastPageId="6" lastPageNumber="599" pageId="3" pageNumber="596" type="discussion">
|
|||
|
|
<paragraph id="8B9D7147FD34842EFF013A8CA16FFE28" blockId="3.[151,1437,1633,2020]" lastBlockId="6.[151,1437,151,501]" lastPageId="6" lastPageNumber="599" pageId="3" pageNumber="596">
|
|||
|
|
<emphasis id="B956AD55FD34842BFF013A8CA237F91E" bold="true" box="[199,315,1741,1767]" pageId="3" pageNumber="596">Remarks.</emphasis>
|
|||
|
|
Before the discovery of
|
|||
|
|
<taxonomicName id="4C220AC4FD34842BFD9B3A8CA09EF91E" authority="Guo & Huang & Wang & Ma, 2023" authorityName="Guo & Huang & Wang & Ma" authorityYear="2023" box="[605,914,1741,1767]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="subSpecies" species="laurae" status="subsp. nov." subSpecies="wuyishana">
|
|||
|
|
<emphasis id="B956AD55FD34842BFD9B3A8CA09EF91E" box="[605,914,1741,1767]" italics="true" pageId="3" pageNumber="596">Chonala laurae wuyishana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B956AD55FD34842BFC5D3A8CA0E4F91E" bold="true" box="[923,1000,1741,1767]" pageId="3" pageNumber="596">
|
|||
|
|
<taxonomicNameLabel id="A265102EFD34842BFC5D3A8CA0E4F91E" box="[923,1000,1741,1767]" pageId="3" pageNumber="596" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, it is considered that this species was the closest relative of
|
|||
|
|
<taxonomicName id="4C220AC4FD34842BFE493AB3A2F5F8F2" authorityName="Bozano" authorityYear="1999" box="[399,505,1777,1804]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD34842BFE493AB3A2F5F8F2" box="[399,505,1777,1804]" italics="true" pageId="3" pageNumber="596">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and might even only represent a southwestern subspecies of the latter (Lang, pers. comm.) due to the similarities in the forewing pattern and the male genitalia. However, after a thorough survey of these three taxa, we consider that
|
|||
|
|
<taxonomicName id="4C220AC4FD34842BFDD73B7BA17AF8AA" authorityName="Lang, Li & Deng" authorityYear="2017" box="[529,630,1849,1876]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD34842BFDD73B7BA17AF8AA" box="[529,630,1849,1876]" italics="true" pageId="3" pageNumber="596">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
should still be treated as a bona species for the following reasons. First, the markings on both sides of the forewing are more ochreous than those of the both subspecies of
|
|||
|
|
<taxonomicName id="4C220AC4FD34842BFB103B1FA633F88E" authorityName="Bozano" authorityYear="1999" box="[1238,1343,1885,1912]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD34842BFB103B1FA633F88E" box="[1238,1343,1885,1912]" italics="true" pageId="3" pageNumber="596">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and the rectangular patch in space 2 on the underside has its inner half nearly fused with the large diffused patch covering the cubital vein and the basal section of spaces 2 and 3, and the boundary between them is obscure. Whereas in both subspecies of
|
|||
|
|
<taxonomicName id="4C220AC4FD34842BFEF73B8BA295F81A" authorityName="Bozano" authorityYear="1999" box="[305,409,1993,2020]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="596" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD34842BFEF73B8BA295F81A" box="[305,409,1993,2020]" italics="true" pageId="3" pageNumber="596">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the rectangular patch is more uniform in color, and the boundary between it and the diffused patch is distinct. Second, in male genitalia, both subspecies of
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFCAA3CD6A0D4FF48" authorityName="Bozano" authorityYear="1999" box="[876,984,151,177]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFCAA3CD6A0D4FF48" box="[876,984,151,177]" italics="true" pageId="6" pageNumber="599">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have their gnathos about one fourth of the length of uncus, while in
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFE203CFAA140FF2C" authorityName="Lang, Li & Deng" authorityYear="2017" box="[486,588,187,213]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFE203CFAA140FF2C" box="[486,588,187,213]" italics="true" pageId="6" pageNumber="599">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
the gnathos is shorter, less than one fifth of the length of uncus. Although
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFF513C9EA2C2FF00" authority="Guo & Huang & Wang & Ma, 2023" authorityName="Guo & Huang & Wang & Ma" authorityYear="2023" box="[151,462,223,249]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="subSpecies" species="laurae" status="subsp. nov." subSpecies="wuyishana">
|
|||
|
|
<emphasis id="B956AD55FD31842EFF513C9EA2C2FF00" box="[151,462,223,249]" italics="true" pageId="6" pageNumber="599">Chonala laurae wuyishana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B956AD55FD31842EFE1F3C9EA12BFF00" bold="true" box="[473,551,223,249]" pageId="6" pageNumber="599">
|
|||
|
|
<taxonomicNameLabel id="A265102EFD31842EFE1F3C9EA12BFF00" box="[473,551,223,249]" pageId="6" pageNumber="599" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
has a prominent and well-defined patch in space 2 on the upperside of the forewing which is similar to that in
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFDFB3D42A1A8FEE4" authorityName="Lang, Li & Deng" authorityYear="2017" box="[573,676,259,285]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFDFB3D42A1A8FEE4" box="[573,676,259,285]" italics="true" pageId="6" pageNumber="599">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, other substantial characters, e. g. the coloration of the upperside markings, the markings in space 2 on the underside of the forewing and the gnathos in male genitalia, all indicate that it is much closer to
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFE673D0AA106FE9C" authorityName="Bozano" authorityYear="1999" box="[417,522,331,357]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFE673D0AA106FE9C" box="[417,522,331,357]" italics="true" pageId="6" pageNumber="599">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
than to
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFDA23D0AA1C4FE9C" authorityName="Lang, Li & Deng" authorityYear="2017" box="[612,712,331,357]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFDA23D0AA1C4FE9C" box="[612,712,331,357]" italics="true" pageId="6" pageNumber="599">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Hence
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFCE23D0AA75FFE9C" authority="Guo & Huang & Wang & Ma, 2023" authorityName="Guo & Huang & Wang & Ma" authorityYear="2023" box="[804,1107,331,357]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="subSpecies" species="laurae" status="subsp. nov." subSpecies="wuyishana">
|
|||
|
|
<emphasis id="B956AD55FD31842EFCE23D0AA75FFE9C" box="[804,1107,331,357]" italics="true" pageId="6" pageNumber="599">Chonala laurae wuyishana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B956AD55FD31842EFB9F3D0AA7AFFE9C" bold="true" box="[1113,1187,331,357]" pageId="6" pageNumber="599">
|
|||
|
|
<taxonomicNameLabel id="A265102EFD31842EFB9F3D0AA7AFFE9C" box="[1113,1187,331,357]" pageId="6" pageNumber="599" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
does not represent the expected intermediate form connecting these two species, and, in the characters mentioned above, no continuous variation is detected among the known individuals of these three taxa. Therefore, there is no reason to downgrade
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFF513DF6A3F0FE28" authorityName="Lang, Li & Deng" authorityYear="2017" box="[151,252,439,465]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFF513DF6A3F0FE28" box="[151,252,439,465]" italics="true" pageId="6" pageNumber="599">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to be a subspecies of
|
|||
|
|
<taxonomicName id="4C220AC4FD31842EFE343DF6A150FE28" authorityName="Bozano" authorityYear="1999" box="[498,604,439,465]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="laurae">
|
|||
|
|
<emphasis id="B956AD55FD31842EFE343DF6A150FE28" box="[498,604,439,465]" italics="true" pageId="6" pageNumber="599">C. laurae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF5D21CFFD33842CFF513BD6A0F3F80E" ID-DOI="http://doi.org/10.5281/zenodo.8243793" ID-Zenodo-Dep="8243793" httpUri="https://zenodo.org/record/8243793/files/figure.png" pageId="4" pageNumber="597" startId="4.[151,264,1943,1967]" targetBox="[203,1384,181,1919]" targetPageId="4" targetType="figure">
|
|||
|
|
<paragraph id="8B9D7147FD33842CFF513BD6A0F3F80E" blockId="4.[151,1437,1943,2039]" pageId="4" pageNumber="597">
|
|||
|
|
<emphasis id="B956AD55FD33842CFF513BD6A24BF856" bold="true" box="[151,327,1943,1967]" pageId="4" pageNumber="597">FIGURES 9–16.</emphasis>
|
|||
|
|
Adults of
|
|||
|
|
<taxonomicName id="4C220AC4FD33842CFE763BD6A145F856" box="[432,585,1943,1967]" class="Insecta" family="Nymphalidae" genus="Chonala" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="undefined-9">
|
|||
|
|
<emphasis id="B956AD55FD33842CFE763BD6A10BF856" box="[432,519,1943,1967]" italics="true" pageId="4" pageNumber="597">Chonala</emphasis>
|
|||
|
|
spp. 9
|
|||
|
|
</taxonomicName>
|
|||
|
|
–10,
|
|||
|
|
<taxonomicName id="4C220AC4FD33842CFDBD3BD9A1D9F856" authorityName="Lang, Li & Deng" authorityYear="2017" box="[635,725,1943,1967]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="houae">
|
|||
|
|
<emphasis id="B956AD55FD33842CFDBD3BD9A1D9F856" box="[635,725,1943,1967]" italics="true" pageId="4" pageNumber="597">C. houae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, holotype, Mt. Jinfo, Chongqing, SATY1156 (CQMNH); 11–12,
|
|||
|
|
<emphasis id="B956AD55FD33842CFAA13BD6A699F856" box="[1383,1429,1943,1967]" italics="true" pageId="4" pageNumber="597">ditto</emphasis>
|
|||
|
|
, paratype,
|
|||
|
|
<emphasis id="B956AD55FD33842CFF3A3BFAA226F82A" box="[252,298,1979,2003]" italics="true" pageId="4" pageNumber="597">ditto</emphasis>
|
|||
|
|
, SATY0466 (CQMNH); 13–14,
|
|||
|
|
<taxonomicName id="4C220AC4FD33842CFDBF3BFDA040F82A" authorityName="Guo & Huang & Wang & Ma" authorityYear="2023" box="[633,844,1979,2003]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="subSpecies" species="laurae" subSpecies="wuyishana">
|
|||
|
|
<emphasis id="B956AD55FD33842CFDBF3BFDA040F82A" box="[633,844,1979,2003]" italics="true" pageId="4" pageNumber="597">C. laurae wuyishana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B956AD55FD33842CFC953BFAA0A2F82A" bold="true" box="[851,942,1979,2003]" pageId="4" pageNumber="597">ssp. nov.</emphasis>
|
|||
|
|
, paratype, Mt. Wuyi, Fujian (SCAU); 15–16,
|
|||
|
|
<taxonomicName id="4C220AC4FD33842CFA433BFDA2A3F80E" authority=", Shennongjia" authorityName="Shennongjia" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="subSpecies" species="laurae" subSpecies="laurae">
|
|||
|
|
<emphasis id="B956AD55FD33842CFA433BFDA22EF80E" italics="true" pageId="4" pageNumber="597">C. laurae laurae</emphasis>
|
|||
|
|
, Shennongjia
|
|||
|
|
</taxonomicName>
|
|||
|
|
, Hubei (CSYL); 9–12 male, 13–16 female. Scale=10 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF5D21CFFD32842DFF513BD6A2B5F80E" ID-DOI="http://doi.org/10.5281/zenodo.8243795" ID-Zenodo-Dep="8243795" httpUri="https://zenodo.org/record/8243795/files/figure.png" pageId="5" pageNumber="598" startId="5.[151,264,1943,1967]" targetBox="[197,1391,194,1907]" targetPageId="5" targetType="figure">
|
|||
|
|
<paragraph id="8B9D7147FD32842DFF513BD6A2B5F80E" blockId="5.[151,1437,1943,2039]" pageId="5" pageNumber="598">
|
|||
|
|
<emphasis id="B956AD55FD32842DFF513BD6A25FF856" bold="true" box="[151,339,1943,1967]" pageId="5" pageNumber="598">FIGURES 17–20.</emphasis>
|
|||
|
|
Male genitalia of
|
|||
|
|
<taxonomicName id="4C220AC4FD32842DFDCC3BD6A16DF856" authorityName="Moore" authorityYear="1893" box="[522,609,1943,1967]" class="Insecta" family="Nymphalidae" genus="Chonala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B956AD55FD32842DFDCC3BD6A16DF856" box="[522,609,1943,1967]" italics="true" pageId="5" pageNumber="598">Chonala</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sspp. Depository of the genitalia: 17 in SCAU, 18 in CSYL, 19 & 20 in CQMNH. a=genitalia capsule with left valva and aedeagus removed; b=aedeagus in dorsal view; c=aedeagus in lateral view; d=left valva in dorsal view. Scale=1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C33822CCFD31842EFF013D9BA0B0FE0C" box="[199,956,474,501]" pageId="6" pageNumber="599" type="distribution">
|
|||
|
|
<paragraph id="8B9D7147FD31842EFF013D9BA0B0FE0C" blockId="6.[151,1437,151,501]" box="[199,956,474,501]" pageId="6" pageNumber="599">
|
|||
|
|
<emphasis id="B956AD55FD31842EFF013D9BA26FFE0D" bold="true" box="[199,355,474,500]" pageId="6" pageNumber="599">Distribution:</emphasis>
|
|||
|
|
<collectingRegion id="49E6BFA5FD31842EFEAC3D9AA2EAFE0C" box="[362,486,475,501]" country="China" name="Chongqing" pageId="6" pageNumber="599">Chongqing</collectingRegion>
|
|||
|
|
(Mt. Jinfo), southwestern
|
|||
|
|
<collectingCountry id="F33531D7FD31842EFCC93D9AA05EFE0C" box="[783,850,475,501]" name="China" pageId="6" pageNumber="599">China</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13196DC2FD31842EFCA43D9AA0BFFE0C" box="[866,947,475,501]" captionStart="FIGURES 21" captionStartId="6.[151,264,1435,1459]" captionTargetBox="[151,1436,525,1412]" captionTargetId="figure-195@6.[151,1436,525,1412]" captionTargetPageId="6" captionText="FIGURES 21. Distribution map of Chonala spp. in the current study." figureDoi="http://doi.org/10.5281/zenodo.8243797" httpUri="https://zenodo.org/record/8243797/files/figure.png" pageId="6" pageNumber="599">Fig. 21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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