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<document id="D8058BC3A20ECFBEF5E4CD19848796B7" ID-CLB-Dataset="3791" ID-DOI="10.11646/zootaxa.5060.1.1" ID-GBIF-Dataset="40f89bbe-7f7b-4abd-aa02-a45e410da232" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606671" ID-ZooBank="14126B35-AEEA-40F5-A892-FE6A8D64E5D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1635406161957" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2021" docId="038F87D5FF86FF97FF6D269DFF759DF4" docLanguage="en" docName="zootaxa.5060.1.1.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diplomaragna budilovi Mikhaljova 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFB6FFADFF8FFF9DFFFA2217FF8C9A76" masterDocTitle="The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="10" updateTime="1699419728057" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E21EF608CAAC3641450D09ABC1874F74">The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:namePart id="7BE175FB02670F3895F778516D2A0F67">Mikhaljova, Elena V.</mods:namePart>
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<mods:typeOfResource id="8DEFD03F0D207A2E22661DF7C47824C3">text</mods:typeOfResource>
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<mods:title id="13D94347F26966885A23E77C778C7335">Zootaxa</mods:title>
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<mods:part id="12E221514C580AA20DE1EFF4F4DE8834">
<mods:date id="C1A9EA2B62F6B7B21F3C8EEE2066B0D3">2021</mods:date>
<mods:detail id="3CC6F377151DECBF98B0F5DCCD9E4B15" type="pubDate">
<mods:number id="BB850EE726C3FBA3C41CF89E7BA6EB87">2021-10-28</mods:number>
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<mods:number id="24C313F2C1875CD85AD2A7272B428387">5060</mods:number>
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<mods:classification id="91B7CBF871259BCF25C16A05310B69A4">journal article</mods:classification>
<mods:identifier id="66B11974F9014C25A43043478A987BE4" type="CLB-Dataset">3791</mods:identifier>
<mods:identifier id="02D6A775B046A0B92930581C7863B84B" type="DOI">10.11646/zootaxa.5060.1.1</mods:identifier>
<mods:identifier id="891BFF88417285B015615D9D5FA285C8" type="GBIF-Dataset">40f89bbe-7f7b-4abd-aa02-a45e410da232</mods:identifier>
<mods:identifier id="F8BB0596CBE527900ECF56D7420192EC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038F87D5FF86FF97FF6D269DFF759DF4" ID-DOI="http://doi.org/10.5281/zenodo.5606657" ID-GBIF-Taxon="189548925" ID-Zenodo-Dep="5606657" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4" httpUri="http://treatment.plazi.org/id/038F87D5FF86FF97FF6D269DFF759DF4" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C33C6548FF86FF94FF6D269DFD999ED3" box="[151,533,1162,1189]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9936C3FF86FF94FF6D269DFD999ED3" blockId="9.[151,533,1162,1225]" box="[151,533,1162,1189]" pageId="9" pageNumber="10">
<heading id="D0D181AFFF86FF94FF6D269DFD999ED3" bold="true" box="[151,533,1162,1189]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B952EAD1FF86FF94FF6D269DFD999ED3" bold="true" box="[151,533,1162,1189]" pageId="9" pageNumber="10">
<taxonomicName id="4C264D40FF86FF94FF6D269DFE3D9ED3" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[151,433,1162,1189]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="budilovi" status="sp. nov.">
<emphasis id="B952EAD1FF86FF94FF6D269DFE3D9ED3" bold="true" box="[151,433,1162,1189]" italics="true" pageId="9" pageNumber="10">Diplomaragna budilovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157AAFF86FF94FE42269CFD999ED3" box="[440,533,1163,1189]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF86FF94FF6D26B8FF759EBF" box="[151,249,1199,1225]" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9936C3FF86FF94FF6D26B8FF759EBF" blockId="9.[151,533,1162,1225]" box="[151,249,1199,1225]" pageId="9" pageNumber="10">
<figureCitation id="131D2A46FF86FF94FF6D26B8FF759EBF" box="[151,249,1199,1225]" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="9" pageNumber="10">Figs 3–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF86FF94FF6D26E1FB219F43" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B9936C3FF86FF94FF6D26E1FB219F43" blockId="9.[151,1437,1270,2017]" pageId="9" pageNumber="10">
<emphasis id="B952EAD1FF86FF94FF6D26E1FE0D9F66" bold="true" box="[151,385,1270,1296]" pageId="9" pageNumber="10">Material examined.</emphasis>
<materialsCitation id="3B4E3C9EFF86FF94FE7326E0FB219F43" ID-GBIF-Occurrence="3395035348" collectingDate="2011-08-05" collectingDateMax="2011-08-09" collectingDateMin="2011-08-05" collectionCode="FSCB, V" country="Russia" elevation="338" location="Nanaysky District" municipality="Khabarovsky Krai" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549D8861FF86FF94FE7326E0FE7A9F67" box="[393,502,1271,1297]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D20FD4AFF86FF94FDFA26E0FDBA9F67" box="[512,566,1271,1297]" pageId="9" pageNumber="10" type="male">male</specimenCount>
(
<collectionCode id="ED37AE06FF86FF94FDBD26E0FD1D9F67" box="[583,657,1271,1297]" pageId="9" pageNumber="10">FSCB</collectionCode>
),
<collectingCountry id="F3317653FF86FF94FD5E26E0FD7D9F67" box="[676,753,1271,1297]" name="Russia" pageId="9" pageNumber="10">Russia</collectingCountry>
,
<collectingMunicipality id="6BFDACB9FF86FF94FD0426E0FC5C9F67" box="[766,976,1271,1297]" pageId="9" pageNumber="10">Khabarovsky Krai</collectingMunicipality>
,
<location id="8EF96018FF86FF94FC2126E0FB269F67" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4:8EF96018FF86FF94FC2126E0FB269F67" box="[987,1194,1271,1297]" country="Russia" municipality="Khabarovsky Krai" name="Nanaysky District" pageId="9" pageNumber="10">Nanaysky District</location>
, basin of
<location id="8EF96018FF86FF94FAED26E0FA149F67" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4:8EF96018FF86FF94FAED26E0FA149F67" box="[1303,1432,1271,1297]" country="Russia" municipality="Khabarovsky Krai" name="Lake Gassi" pageId="9" pageNumber="10">Lake Gassi</location>
, near
<location id="8EF96018FF86FF94FF34270CFEFC9F43" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4:8EF96018FF86FF94FF34270CFEFC9F43" box="[206,368,1307,1333]" country="Russia" municipality="Khabarovsky Krai" name="Tukhala River" pageId="9" pageNumber="10">Tukhala River</location>
,
<location id="8EF96018FF86FF94FE81270CFD069F42" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4:8EF96018FF86FF94FE81270CFD069F42" box="[379,650,1307,1333]" country="Russia" municipality="Khabarovsky Krai" name="Betula forest with Picea" pageId="9" pageNumber="10">
<taxonomicName id="4C264D40FF86FF94FE81270CFE489F43" box="[379,452,1307,1333]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAD1FF86FF94FE81270CFE489F43" box="[379,452,1307,1333]" italics="true" pageId="9" pageNumber="10">Betula</emphasis>
</taxonomicName>
forest with
<taxonomicName id="4C264D40FF86FF94FDB0270CFD069F42" box="[586,650,1307,1332]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAD1FF86FF94FDB0270CFD069F42" box="[586,650,1307,1332]" italics="true" pageId="9" pageNumber="10">Picea</emphasis>
</taxonomicName>
</location>
,
<quantity id="4CDE9B26FF86FF94FD62270CFD789F43" box="[664,756,1307,1333]" metricMagnitude="2" metricUnit="m" metricValue="3.377" pageId="9" pageNumber="10" unit="m" value="337.7">
<elevation id="000BD1F0FF86FF94FD62270CFD789F43" box="[664,756,1307,1333]" metricMagnitude="2" metricUnit="m" metricValue="3.377" pageId="9" pageNumber="10" unit="m" value="337.7">337.7 m</elevation>
</quantity>
a.s.l.,
<date id="FF981003FF86FF94FCC7270DFC529F43" box="[829,990,1306,1333]" pageId="9" pageNumber="10" value="2011-08-05" valueMax="2011-08-09" valueMin="2011-08-05">
<collectingDate id="EFDCE9EBFF86FF94FCC7270DFC529F43" box="[829,990,1306,1333]" pageId="9" pageNumber="10" value="2011-08-05" valueMax="2011-08-09" valueMin="2011-08-05">5–9.VIII.2011</collectingDate>
</date>
, leg. P.
<collectionCode id="ED37AE06FF86FF94FBD4270CFBCF9F43" box="[1070,1091,1307,1333]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="10" type="Museum">V</collectionCode>
.
<location id="8EF96018FF86FF94FBB6270CFB249F43" LSID="urn:lsid:plazi:treatment:038F87D5FF86FF97FF6D269DFF759DF4:8EF96018FF86FF94FBB6270CFB249F43" box="[1100,1192,1307,1333]" country="Russia" municipality="Khabarovsky Krai" name="Budilov" pageId="9" pageNumber="10">Budilov</location>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF86FF94FF3D2729FA939C0F" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B9936C3FF86FF94FF3D2729FC159F0B" blockId="9.[151,1437,1270,2017]" pageId="9" pageNumber="10">
<emphasis id="B952EAD1FF86FF94FF3D2729FECE9F2E" bold="true" box="[199,322,1342,1368]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Differs from congeners mainly by the gonopod posterior angiocoxal process in the form of a thin, frontally curved plate with an elongated mesal pointed horn apically.
</paragraph>
<paragraph id="8B9936C3FF86FF94FF3D2790FA939C0F" blockId="9.[151,1437,1270,2017]" pageId="9" pageNumber="10">
The species seems to be particularly similar to
<taxonomicName id="4C264D40FF86FF94FD2D2790FB839FD6" authorityName="Mikhaljova" authorityYear="1997" box="[727,1039,1415,1441]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zimoveinaya">
<emphasis id="B952EAD1FF86FF94FD2D2790FB839FD6" box="[727,1039,1415,1441]" italics="true" pageId="9" pageNumber="10">Diplomaragna zimoveinaya</emphasis>
</taxonomicName>
and
<taxonomicName id="4C264D40FF86FF94FBBD2790FB4C9FD7" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[1095,1216,1415,1441]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentifer" status="sp. nov.">
<emphasis id="B952EAD1FF86FF94FBBD2790FB4C9FD7" box="[1095,1216,1415,1441]" italics="true" pageId="9" pageNumber="10">D. dentifer</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF86FF94FB3D2790FAAD9FD7" bold="true" box="[1223,1313,1415,1441]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A26157AAFF86FF94FB3D2790FAAD9FD7" box="[1223,1313,1415,1441]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but differs in the configuration of the gonopod posterior angiocoxal processes (
<figureCitation id="131D2A46FF86FF94FC7727BCFC7C9FB3" box="[909,1008,1451,1477]" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="9" pageNumber="10">Figs 4, 7</figureCitation>
) with an elongated mesal horn (
<emphasis id="B952EAD1FF86FF94FAAC27BCFAEE9FB3" bold="true" box="[1366,1378,1451,1477]" pageId="9" pageNumber="10">c</emphasis>
,
<figureCitation id="131D2A46FF86FF94FA8A27BCFF2A9F9F" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
) apically (vs the mesal horn apically not elongated in both) and an even distal edge (vs uneven distal edge in
<taxonomicName id="4C264D40FF86FF94FA7B27D8FEAF9C7A" authorityName="Mikhaljova" authorityYear="1997" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zimoveinaya">
<emphasis id="B952EAD1FF86FF94FA7B27D8FEAF9C7A" italics="true" pageId="9" pageNumber="10">D. zimoveinaya</emphasis>
</taxonomicName>
, but straight (without recesses) distal edge covered with ribbs in
<taxonomicName id="4C264D40FF86FF94FBF127E4FB099C7B" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[1035,1157,1523,1549]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentifer" status="sp. nov.">
<emphasis id="B952EAD1FF86FF94FBF127E4FB099C7B" box="[1035,1157,1523,1549]" italics="true" pageId="9" pageNumber="10">D. dentifer</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF86FF94FB7427E4FB659C7B" bold="true" box="[1166,1257,1523,1549]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A26157AAFF86FF94FB7427E4FB659C7B" box="[1166,1257,1523,1549]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), by having the distal part of the gonopod colpocoxite not flagelliform (vs flagelliform distal part of the gonopod colpocoxite in
<taxonomicName id="4C264D40FF86FF94FF6D242CFECA9C22" authorityName="Mikhaljova" authorityYear="1997" box="[151,326,1595,1620]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zimoveinaya">
<emphasis id="B952EAD1FF86FF94FF6D242CFECA9C22" box="[151,326,1595,1620]" italics="true" pageId="9" pageNumber="10">D. zimoveinaya</emphasis>
</taxonomicName>
but not flagelliform in
<taxonomicName id="4C264D40FF86FF94FDAA242CFD469C23" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[592,714,1595,1621]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentifer" status="sp. nov.">
<emphasis id="B952EAD1FF86FF94FDAA242CFD469C23" box="[592,714,1595,1621]" italics="true" pageId="9" pageNumber="10">D. dentifer</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF86FF94FD2B242CFCA09C23" bold="true" box="[721,812,1595,1621]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A26157AAFF86FF94FD2B242CFCA09C23" box="[721,812,1595,1621]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) as well as by the relatively short distal segment of the posterior gonopod telopodite (vs elongate in
<taxonomicName id="4C264D40FF86FF94FD742448FCB09C0E" authorityName="Mikhaljova" authorityYear="1997" box="[654,828,1631,1656]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zimoveinaya">
<emphasis id="B952EAD1FF86FF94FD742448FCB09C0E" box="[654,828,1631,1656]" italics="true" pageId="9" pageNumber="10">D. zimoveinaya</emphasis>
</taxonomicName>
, of medium length in
<taxonomicName id="4C264D40FF86FF94FBCF2448FB229C0F" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[1077,1198,1631,1657]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentifer" status="sp. nov.">
<emphasis id="B952EAD1FF86FF94FBCF2448FB229C0F" box="[1077,1198,1631,1657]" italics="true" pageId="9" pageNumber="10">D. dentifer</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF86FF94FB4F2448FA829C0F" bold="true" box="[1205,1294,1631,1657]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A26157AAFF86FF94FB4F2448FA829C0F" box="[1205,1294,1631,1657]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF86FF97FF3D2495FE1D9D4C" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9936C3FF86FF94FF3D2495FC279C93" blockId="9.[151,1437,1270,2017]" pageId="9" pageNumber="10">
<emphasis id="B952EAD1FF86FF94FF3D2495FED59CEA" bold="true" box="[199,345,1666,1692]" pageId="9" pageNumber="10">Description.</emphasis>
Male (two fragments of the anterior body part). Length of these two fragments about
<quantity id="4CDE9B26FF86FF94FACD2494FA149CEB" box="[1335,1432,1667,1693]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="9" pageNumber="10" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
, width 1.5–1.7 mm with paraterga. Coloration in alcohol brown (discoloration during fixation is possible). Legs dark brown. Antennae dark brown. Eye patches black, with about 40 ocelli.
</paragraph>
<paragraph id="8B9936C3FF86FF94FF3D24F8FB869DEF" blockId="9.[151,1437,1270,2017]" pageId="9" pageNumber="10">The exact number of body rings is unknown. There are 2 fragments (7 and 10 rings) of the front part; the posterior part is broken off and lost. Collum semicircular. Both collum and ring 2 narrower than head with genae. Ring 2 somewhat wider than collum. Paraterga beginning on ring 2–3, well developed on rings 5 and on the remaining rings. Anterolateral macrochaetae shortest, pointed, but not very sharply so. Caudolateral macrochaetae longest. Medial and caudolateral macrochaetae pointed. Axial suture poorly developed.</paragraph>
<paragraph id="8B9936C3FF86FF97FF3D25B4FC7C9A8F" blockId="9.[151,1437,1270,2017]" lastBlockId="10.[151,1437,151,321]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">Legs long and slender. Leg pairs 1 and 2 typically reduced in size, with usual tarsal brushes. Other pregonopodal legs practically not enlarged. Leg pairs 3–7 with a group of funnel–shaped tarsal papillae apically near claw. Claw of leg pairs 3–7 at base with a setoid outgrowth ventrally but without additional claws dorsally. Postgonopodal legs (including leg pairs 10 and 11) without tarsal papillae. Claw of midbody legs (including leg pairs 10 and 11) at base with a long setoid outgrowth ventrally but without additional claws dorsally.</paragraph>
<paragraph id="8B9936C3FF85FF97FF3D2314FE379B37" blockId="10.[151,1437,151,321]" pageId="10" pageNumber="11">Legs 10 and 11 with coxal glands. Coxae and trochanters 10 unmodified. Coxa 11 with very low outgrowth; trochanter 11 unmodified.</paragraph>
<caption id="DF59664BFF85FF97FF6D2759FEF59FA4" ID-DOI="http://doi.org/10.5281/zenodo.5606675" ID-Zenodo-Dep="5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1358,1382]" targetBox="[151,1435,345,1331]" targetPageId="10">
<paragraph id="8B9936C3FF85FF97FF6D2759FEF59FA4" blockId="10.[151,1436,1358,1490]" pageId="10" pageNumber="11">
<emphasis id="B952EAD1FF85FF97FF6D2759FEB69F10" bold="true" box="[151,314,1358,1382]" pageId="10" pageNumber="11">FIGURES 3–7.</emphasis>
<taxonomicName id="4C264D40FF85FF97FEC52759FDA19F10" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[319,557,1358,1382]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="budilovi" status="sp. nov.">
<emphasis id="B952EAD1FF85FF97FEC52759FDA19F10" box="[319,557,1358,1382]" italics="true" pageId="10" pageNumber="11">Diplomaragna budilovi</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF85FF97FDC82759FD0E9F10" bold="true" box="[562,642,1358,1382]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A26157AAFF85FF97FDC82759FD0E9F10" box="[562,642,1358,1382]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view;
<emphasis id="B952EAD1FF85FF97FBA52765FBE69FFC" bold="true" box="[1119,1130,1394,1418]" pageId="10" pageNumber="11">c</emphasis>
, horn;
<emphasis id="B952EAD1FF85FF97FB482765FB429FFC" bold="true" box="[1202,1230,1394,1418]" pageId="10" pageNumber="11">pp</emphasis>
, posterior angicoxal process;
<emphasis id="B952EAD1FF85FF97FF0A2781FE879FD8" bold="true" box="[240,267,1430,1454]" pageId="10" pageNumber="11">ap</emphasis>
, anterior angiocoxal process;
<emphasis id="B952EAD1FF85FF97FDCD2781FDC39FD8" bold="true" box="[567,591,1430,1454]" pageId="10" pageNumber="11">cx</emphasis>
, coxosternum;
<emphasis id="B952EAD1FF85FF97FD102781FD739FD8" bold="true" box="[746,767,1430,1454]" pageId="10" pageNumber="11">at</emphasis>
, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7).
</paragraph>
</caption>
<paragraph id="8B9936C3FF85FF97FF3D2417FB5F9D61" blockId="10.[151,1437,1536,1922]" pageId="10" pageNumber="11">
Gonopods as in figs 3–7. Anterior gonopod telopodites flagelliform, covered with tiny spinules (at least basally) positioned on posterior surface of posterior gonopod colpocoxites inside a shallow, partly open-edged sheathing groove. Colpocoxites fused medially in basal half; their distal parts slightly curved caudally, tips hooked. Each colpocoxite entire. Colpocoxite sheath groove without evident processes. Posterior gonopod angiocoxites with subglobules in posterior view, depressed centrally in anterior view. Anterior angiocoxal process (
<emphasis id="B952EAD1FF85FF97FB192487FA8D9CDC" bold="true" box="[1251,1281,1680,1706]" pageId="10" pageNumber="11">ap</emphasis>
,
<figureCitation id="131D2A46FF85FF97FAF52487FADE9CDD" box="[1295,1362,1680,1707]" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
) long, flagelliform, its distal portion positioned inside frontal fold of colpocoxites. Posterior angiocoxal process (
<emphasis id="B952EAD1FF85FF97FAB824A3FAED9CB8" bold="true" box="[1346,1377,1716,1742]" pageId="10" pageNumber="11">pp</emphasis>
,
<figureCitation id="131D2A46FF85FF97FA8A24A3FF299C84" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
) large, in the form of a thin, frontally curved plate with elongated mesal pointed horn (
<emphasis id="B952EAD1FF85FF97FB9324CFFBF99C84" bold="true" box="[1129,1141,1752,1778]" pageId="10" pageNumber="11">c</emphasis>
,
<figureCitation id="131D2A46FF85FF97FB7924CFFB489C85" box="[1155,1220,1752,1779]" captionStart="FIGURES 3–7" captionStartId="10.[151,264,1358,1382]" captionTargetBox="[151,1435,345,1331]" captionTargetId="figure-95@10.[151,1435,345,1334]" captionTargetPageId="10" captionText="FIGURES 3–7. Diplomaragna budilovi sp. nov., male holotype. 3, 7, gonopods, caudal view; 4, posterior angiocoxal processes and colpocoxites, caudal view; 5, gonopods, front view; 6, basal part of gonopods, caudal view; c, horn; pp, posterior angicoxal process; ap, anterior angiocoxal process; cx, coxosternum; at, anterior gonopod telopodite. Scales: 20 μm (Fig. 6), 100 μm (Figs 3–5), 0.5 mm (Fig. 7)." figureDoi="http://doi.org/10.5281/zenodo.5606675" httpUri="https://zenodo.org/record/5606675/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
) apically. Posterior gonopod telopodite 2-segmented, setose; length of the distal segment is more short than average.
</paragraph>
<paragraph id="8B9936C3FF85FF97FF3D2537FE1D9D4C" blockId="10.[151,1437,1536,1922]" box="[199,401,1824,1850]" pageId="10" pageNumber="11">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF85FF97FF3D2553FF759DF4" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8B9936C3FF85FF97FF3D2553FF759DF4" blockId="10.[151,1437,1536,1922]" pageId="10" pageNumber="11">
<emphasis id="B952EAD1FF85FF97FF3D2553FEC29D28" bold="true" box="[199,334,1860,1886]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species is named in honor of the collector P.V. Budilov, Russian entomologist. A noun in genitive.
</paragraph>
</subSubSection>
</treatment>
</document>
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