treatments-xml/data/03/D2/3B/03D23B67FFD8D827E7C5FAC1660993E2.xml

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<mods:title id="330BE9AB06E415D1CA0EF762CA8B8413">Redescription of Notholimnophila Alexander and Acantholimnophila Alexander, poorly known New Zealand endemics (Diptera: Tipulomorpha: Limoniidae)</mods:title>
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<mods:namePart id="2849C11708A60118BCAFAA93C64D62B2">Ribeiro, Guilherme Cunha</mods:namePart>
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<treatment id="03D23B67FFD8D827E7C5FAC1660993E2" ID-DOI="http://doi.org/10.5281/zenodo.6495713" ID-GBIF-Taxon="119430709" ID-Zenodo-Dep="6495713" LSID="urn:lsid:plazi:treatment:03D23B67FFD8D827E7C5FAC1660993E2" httpUri="http://treatment.plazi.org/id/03D23B67FFD8D827E7C5FAC1660993E2" lastPageId="4" lastPageNumber="58" pageId="1" pageNumber="55">
<subSubSection id="C361D9FAFFD8D822E7C5FAC165099396" pageId="1" pageNumber="55" type="nomenclature">
<paragraph id="8BC48A71FFD8D822E7C5FAC166A09379" blockId="1.[151,660,1373,1433]" box="[151,660,1373,1398]" pageId="1" pageNumber="55">
<heading id="D08C3D1DFFD8D822E7C5FAC166A09379" bold="true" box="[151,660,1373,1398]" fontSize="10" level="2" pageId="1" pageNumber="55" reason="2">
<taxonomicName id="4C7BF1F2FFD8D822E7C5FAC166A09379" ID-CoL="47TBM" authority="Alexander, 1922" authorityName="Alexander" authorityYear="1922" box="[151,660,1373,1398]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFD8D822E7C5FAC166A09379" bold="true" box="[151,660,1373,1398]" pageId="1" pageNumber="55">
<emphasis id="B90F5663FFD8D822E7C5FAC1658E9379" bold="true" box="[151,442,1373,1398]" italics="true" pageId="1" pageNumber="55">Notholimnophila exclusa</emphasis>
(Alexander, 1922)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC48A71FFD8D822E7C5FA1D65099396" blockId="1.[151,660,1373,1433]" box="[151,317,1409,1433]" pageId="1" pageNumber="55">
(
<figureCitation id="134096F4FFD8D822E7CCFA1D65009396" box="[158,308,1409,1433]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 9" captionStart-3="FIGURES 10 – 11" captionStart-4="FIGURES 12 – 13" captionStart-5="FIGURES 14 – 16" captionStartId-0="2.[151,264,1400,1423]" captionStartId-1="3.[151,264,1457,1480]" captionStartId-2="4.[151,264,1257,1280]" captionStartId-3="5.[151,264,1386,1409]" captionStartId-4="6.[151,264,1616,1639]" captionStartId-5="7.[151,264,1933,1956]" captionTargetBox-0="[151,1436,155,1379]" captionTargetBox-1="[151,1436,193,1435]" captionTargetBox-2="[151,1436,193,1234]" captionTargetBox-3="[151,1436,193,1362]" captionTargetBox-4="[183,1382,215,1592]" captionTargetBox-5="[162,1425,202,1901]" captionTargetId-0="figure@2.[151,1436,155,1379]" captionTargetId-1="figure@3.[151,1436,193,1437]" captionTargetId-2="figure@4.[151,1436,193,1236]" captionTargetId-3="figure@5.[151,1436,193,1365]" captionTargetId-4="figure@6.[151,1436,193,1595]" captionTargetId-5="figure@7.[151,1436,193,1912]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURES 1 – 3. Notholimnophila exclusa (Alexander). Male holotype. 1: habitus. 2: head, lateral view. 3: head, dorsal view." captionText-1="FIGURES 4 – 5. Notholimnophila exclusa (Alexander). Female. 4: head and thorax, lateral view. 5: wing." captionText-2="FIGURES 6 – 9. Notholimnophila exclusa (Alexander). Male terminalia. 6: dorsal view. 7: lateral view. 8: ventral view. 9: Detail of gonostylus, dorsal view." captionText-3="FIGURES 10 – 11. Notholimnophila exclusa (Alexander). Female ovipositor. 10: lateral view. 11: dorsal view." captionText-4="FIGURES 12 – 13. Notholimnophila exclusa (Alexander). 12: wing venation. 13: thoracic sclerites, lateral view." captionText-5="FIGURES 14 – 16. Notholimnophila exclusa (Alexander). 14: male genitalia, dorsal view. 15: aedeagus and associated structures, dorsal view. 16: male terminalia, lateral view." httpUri-0="https://zenodo.org/record/279867/files/figure.png" httpUri-1="https://zenodo.org/record/279868/files/figure.png" httpUri-2="https://zenodo.org/record/279869/files/figure.png" httpUri-3="https://zenodo.org/record/279870/files/figure.png" httpUri-4="https://zenodo.org/record/279871/files/figure.png" httpUri-5="https://zenodo.org/record/279872/files/figure.png" pageId="1" pageNumber="55">Figures 1–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C361D9FAFFD8D822E7C5FA5565A69012" pageId="1" pageNumber="55" type="reference_group">
<paragraph id="8BC48A71FFD8D822E7C5FA5565A69012" blockId="1.[151,1436,1481,1565]" pageId="1" pageNumber="55">
<taxonomicName id="4C7BF1F2FFD8D822E7C5FA55666E93EF" ID-CoL="47TBM" authority="Alexander, 1922" authorityName="Alexander" authorityYear="1922" box="[151,602,1481,1504]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFD8D822E7C5FA5565A293D0" box="[151,406,1481,1503]" italics="true" pageId="1" pageNumber="55">Notholimnophila exclusa</emphasis>
(Alexander, 1922)
</taxonomicName>
. 
<typeStatus id="54C034D3FFD8D822E534FA5666AC93D0" box="[614,664,1482,1503]" pageId="1" pageNumber="55">Type</typeStatus>
species: 
<taxonomicName id="4C7BF1F2FFD8D822E5AAFA55608593D0" ID-CoL="8CKDX" authority="Alexander, 1922a: 517" authorityName="Alexander" authorityPageNumber="517" authorityYear="1922" box="[760,1201,1481,1503]" class="Insecta" family="Limoniidae" genus="Limnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="55" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFD8D822E5AAFA5567F493D0" box="[760,960,1481,1503]" italics="true" pageId="1" pageNumber="55">Limnophila exclusa</emphasis>
<bibRefCitation id="EFEAF780FFD8D822E49AFA5560B493D0" author="Alexander" box="[968,1152,1481,1503]" pageId="1" pageNumber="55" refString="Alexander, C. P. (1922 a) New or little-known Tipulidae (Diptera). X. Australasian species. Annals and Magazine of Natural History, 9, 505 - 524" type="journal article" year="1922" yearSuffix="a">Alexander, 1922a</bibRefCitation>
: 517
</taxonomicName>
–518 (original description); 
<bibRefCitation id="EFEAF780FFD8D822E657FA74658693F1" author="Alexander" box="[261,434,1512,1534]" pageId="1" pageNumber="55" refString="Alexander, C. P. (1924) New or little-known Tipulidae (Diptera). XX. Australasian species. Annals and Magazine of Natural History, 9, 177 - 194." type="journal article" year="1924">Alexander, 1924</bibRefCitation>
: 369 (taxonomy); 
<bibRefCitation id="EFEAF780FFD8D822E53DFA7466CB93F1" author="Ribeiro" box="[623,767,1512,1534]" pageId="1" pageNumber="55" refString="Ribeiro, G. C. (2008) Phylogeny of the Limnophilinae (Limoniidae) and early evolution of the Tipulomorpha (Diptera). Invertebrate Systematics, 22, 627 - 694." type="journal article" year="2008">Ribeiro, 2008</bibRefCitation>
: figs. 85, 159 (illustrations of male genitalia); 
<bibRefCitation id="EFEAF780FFD8D822E38DFA7461A893F1" author="Oosterbroek" box="[1247,1436,1512,1534]" pageId="1" pageNumber="55" refString="Oosterbroek, P. (2011) Catalogue of the Craneflies of the World, (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae). Available from: http: // ip 30. eti. uva. nl / ccw / index. php (accessed 7 th June 2011)." type="journal volume" year="2011">Oosterbroek, 2011</bibRefCitation>
(catalogue citation).
</paragraph>
</subSubSection>
<subSubSection id="C361D9FAFFD8D821E7C5F9D561A891FA" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55" type="description">
<paragraph id="8BC48A71FFD8D822E7C5F9D56713911A" blockId="1.[151,1436,1609,2030]" pageId="1" pageNumber="55">
<emphasis id="B90F5663FFD8D822E7C5F9D56570906D" bold="true" box="[151,324,1609,1634]" pageId="1" pageNumber="55">Redescription.</emphasis>
Coloration (male and female): Head and appendages brownish grey, little darker than thorax. Thorax brownish, darker dorsally, gradually turning to pale brown-yellowish toward coxae. Scutum with distinct dark brown markings: an anteromedial band plus two posterolateral strips. Wings with a brownish tinge, darker along cell c. Darker brown areas at base of Rs, along the cord, pterostigma and distal parts of wing. Male abdomen including genitalia brownish. Female abdomen brownish. Ovipositor yellowish. Legs pale brown-yellowish, with darker areas at tips of femora and tibiae. Tarsi dark brown.
</paragraph>
<paragraph id="8BC48A71FFD8D822E795F8BD651D918E" blockId="1.[151,1436,1609,2030]" pageId="1" pageNumber="55">
<emphasis id="B90F5663FFD8D822E795F8BD656D9135" bold="true" box="[199,345,1825,1850]" pageId="1" pageNumber="55">Dimensions.</emphasis>
Head length, ca. 1.0 mm; Head width, ca. 1.0 mm; thorax length, 
<quantity id="4C832794FFD8D822E31AF8BD60E79136" box="[1096,1235,1825,1849]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="1" pageNumber="55" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
; thorax height ca. 
<quantity id="4C832794FFD8D822E7C5F8D964C49152" box="[151,240,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="55" unit="mm" value="2.5">2.5 mm</quantity>
; wing length, 10.0– 
<quantity id="4C832794FFD8D822E682F8D966029152" box="[464,566,1861,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="55" unit="mm" value="11.3">11.3 mm</quantity>
; wing width, 
<quantity id="4C832794FFD8D822E59DF8D9676E9152" box="[719,858,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="1" pageNumber="55" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
; gonocoxite length, ca. 
<quantity id="4C832794FFD8D822E337F8D960F99152" box="[1125,1229,1861,1885]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="55" unit="mm" value="0.63">0.63 mm</quantity>
; gonocoxite width ca. 
<quantity id="4C832794FFD8D822E7ECF8F56511918E" box="[190,293,1897,1921]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="55" unit="mm" value="0.32">0.32 mm</quantity>
.
</paragraph>
<paragraph id="8BC48A71FFD8D821E795F81261A891FA" blockId="1.[151,1436,1609,2030]" lastBlockId="2.[151,1436,1462,2037]" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55">
<emphasis id="B90F5663FFD8D822E795F812655491A9" bold="true" box="[199,352,1934,1958]" pageId="1" pageNumber="55">Morphology.</emphasis>
Head (
<figureCitation id="134096F4FFD8D822E6E6F811662F91AA" box="[436,539,1933,1957]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1400,1423]" captionTargetBox="[151,1436,155,1379]" captionTargetId="figure@2.[151,1436,155,1379]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Notholimnophila exclusa (Alexander). Male holotype. 1: habitus. 2: head, lateral view. 3: head, dorsal view." httpUri="https://zenodo.org/record/279867/files/figure.png" pageId="1" pageNumber="55">Figs 2, 3</figureCitation>
). Flagellum 14-segmented; first 3–4 flagellomeres sub globular, conspicuously stouter than remaining, bearing stout dorsal verticils; remaining flagellomeres more ovoid, bearing longer verticils; Scape ca. 1.86x longer than wide and ca. 2.7x longer than pedicel; palpus 4-segmented, all segments tubular in shape, subequal in length; eyes separated dorsally by a distance virtually equal to the length of scape, and ventrally by a much narrower space. Wing (
<figureCitation id="134096F4FFDBD821E545FA46665493FD" box="[535,608,1498,1522]" captionStart="FIGURES 4 – 5" captionStartId="3.[151,264,1457,1480]" captionTargetBox="[151,1436,193,1435]" captionTargetId="figure@3.[151,1436,193,1437]" captionTargetPageId="3" captionText="FIGURES 4 – 5. Notholimnophila exclusa (Alexander). Female. 4: head and thorax, lateral view. 5: wing." httpUri="https://zenodo.org/record/279868/files/figure.png" pageId="2" pageNumber="56">Figs 5</figureCitation>
, 
<figureCitation id="134096F4FFDBD821E53EFA4666BF93FD" box="[620,651,1498,1522]" captionStart="FIGURES 12 – 13" captionStartId="6.[151,264,1616,1639]" captionTargetBox="[183,1382,215,1592]" captionTargetId="figure@6.[151,1436,193,1595]" captionTargetPageId="6" captionText="FIGURES 12 – 13. Notholimnophila exclusa (Alexander). 12: wing venation. 13: thoracic sclerites, lateral view." httpUri="https://zenodo.org/record/279871/files/figure.png" pageId="2" pageNumber="56">12</figureCitation>
). Vein h situated at the level of midlength between origin and bifurcation of M+Cu; tip of Sc at the level of first bifurcation of M; Sc-r oblique, little longer than tip of Sc; R1 somewhat sinuous between r-r and sc-r, straight and pointed upwards after contact with r-r; Rs with or without spur at its base, mostly straight; r-r situated at level of bifurcation of R2+3; R2 parallel with longitudinal axis of wing; R3 more oblique in position, pointed downwards; cell r2 about as long as its petiole; r-m subequal in length or little longer than basal section of M1+2+3; only three branches of M reaching wing margin; discal cell large, almost as long as cel r2; m-cu situated at midpoint of discal cell; tip of vein A1 at the level of vein Sc; tip of vein A2 at the level of origin of Rs; Thorax (
<figureCitation id="134096F4FFDBD821E617F97D65BA90F6" box="[325,398,1761,1785]" captionStart="FIGURES 4 – 5" captionStartId="3.[151,264,1457,1480]" captionTargetBox="[151,1436,193,1435]" captionTargetId="figure@3.[151,1436,193,1437]" captionTargetPageId="3" captionText="FIGURES 4 – 5. Notholimnophila exclusa (Alexander). Female. 4: head and thorax, lateral view. 5: wing." httpUri="https://zenodo.org/record/279868/files/figure.png" pageId="2" pageNumber="56">Figs 4</figureCitation>
, 
<figureCitation id="134096F4FFDBD821E6CEF97D658E90F6" box="[412,442,1761,1785]" captionStart="FIGURES 12 – 13" captionStartId="6.[151,264,1616,1639]" captionTargetBox="[183,1382,215,1592]" captionTargetId="figure@6.[151,1436,193,1595]" captionTargetPageId="6" captionText="FIGURES 12 – 13. Notholimnophila exclusa (Alexander). 12: wing venation. 13: thoracic sclerites, lateral view." httpUri="https://zenodo.org/record/279871/files/figure.png" pageId="2" pageNumber="56">13</figureCitation>
). Thoracic sclerites as in 
<figureCitation id="134096F4FFDBD821E58EF97D677890F6" box="[732,844,1761,1785]" captionStart="FIGURES 12 – 13" captionStartId="6.[151,264,1616,1639]" captionTargetBox="[183,1382,215,1592]" captionTargetId="figure@6.[151,1436,193,1595]" captionTargetPageId="6" captionText="FIGURES 12 – 13. Notholimnophila exclusa (Alexander). 12: wing venation. 13: thoracic sclerites, lateral view." httpUri="https://zenodo.org/record/279871/files/figure.png" pageId="2" pageNumber="56">Figure 13</figureCitation>
; tibial spurs present and distinct; tibial spur formula 1:2:2; Tarsal claws simple and smooth. Male genitalia (
<figureCitation id="134096F4FFDBD821E448F89967B69112" box="[794,898,1797,1821]" captionStart="FIGURES 6 – 9" captionStartId="4.[151,264,1257,1280]" captionTargetBox="[151,1436,193,1234]" captionTargetId="figure@4.[151,1436,193,1236]" captionTargetPageId="4" captionText="FIGURES 6 – 9. Notholimnophila exclusa (Alexander). Male terminalia. 6: dorsal view. 7: lateral view. 8: ventral view. 9: Detail of gonostylus, dorsal view." httpUri="https://zenodo.org/record/279869/files/figure.png" pageId="2" pageNumber="56">Figs 6–9</figureCitation>
, 
<figureCitation id="134096F4FFDBD821E4C2F89967EE9112" box="[912,986,1797,1821]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1933,1956]" captionTargetBox="[162,1425,202,1901]" captionTargetId="figure@7.[151,1436,193,1912]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Notholimnophila exclusa (Alexander). 14: male genitalia, dorsal view. 15: aedeagus and associated structures, dorsal view. 16: male terminalia, lateral view." httpUri="https://zenodo.org/record/279872/files/figure.png" pageId="2" pageNumber="56">14–16</figureCitation>
). Posterior margin of tergite 9 forming extensions longer than wide. Gonocoxite cone-shaped, ca. 2.5x longer than wide; gonostylus terminal; lobe of gonostylus 
<emphasis id="B90F5663FFDBD821E64BF8D3650E9169" box="[281,314,1871,1894]" italics="true" pageId="2" pageNumber="56">ca.</emphasis>
3.4x longer than wide, gradually narrowed toward tip; clasper of gonostylus 
<emphasis id="B90F5663FFDBD821E3E4F8D360EC9169" box="[1206,1240,1871,1894]" italics="true" pageId="2" pageNumber="56">ca.</emphasis>
3.3x longer than wide, glabrous, simple, strongly serrated; aedeagus relatively short, reaching the level the tips of the interbasis; aedeagal sheath lacking processes or extensions; interbase blade-like, little serrated at apex, fused medially but with no conspicuous lateral extensions articulating with paramere. Female terminalia (
<figureCitation id="134096F4FFDBD821E37AF82560FA91DE" box="[1064,1230,1977,2001]" captionStart="FIGURES 10 – 11" captionStartId="5.[151,264,1386,1409]" captionTargetBox="[151,1436,193,1362]" captionTargetId="figure@5.[151,1436,193,1365]" captionTargetPageId="5" captionText="FIGURES 10 – 11. Notholimnophila exclusa (Alexander). Female ovipositor. 10: lateral view. 11: dorsal view." httpUri="https://zenodo.org/record/279870/files/figure.png" pageId="2" pageNumber="56">Figures 10, 11</figureCitation>
): tergite 10 ovoid, ca. 
<date id="FFC5ACB1FFDBD821E7EFF84164ED91FA" box="[189,217,2013,2037]" pageId="2" pageNumber="56">2x</date>
longer than wide; hypogynial valve well sclerotized; apex of hypogynial valve reaching midlength of cercus.
</paragraph>
</subSubSection>
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<paragraph id="8BC48A71FFDBD821E7C5FAE461B09380" blockId="2.[151,1412,1400,1423]" box="[151,1412,1400,1423]" pageId="2" pageNumber="56">
<emphasis id="B90F5663FFDBD821E7C5FAE4650E9381" bold="true" box="[151,314,1400,1423]" pageId="2" pageNumber="56">FIGURES 1–3.</emphasis>
<taxonomicName id="4C7BF1F2FFDBD821E612FAE566F69380" authority="Alexander" authorityName="Alexander" box="[320,706,1401,1423]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFDBD821E612FAE5660A9380" box="[320,574,1401,1423]" italics="true" pageId="2" pageNumber="56">Notholimnophila exclusa</emphasis>
(Alexander)
</taxonomicName>
. Male holotype. 1: habitus. 2: head, lateral view. 3: head, dorsal view.
</paragraph>
</caption>
<caption id="DF04DAF9FFDAD820E7C5FA2D60F593C7" box="[151,1217,1457,1480]" httpUri="https://zenodo.org/record/279868/files/figure.png" pageId="3" pageNumber="57" targetBox="[151,1436,193,1435]" targetPageId="3">
<paragraph id="8BC48A71FFDAD820E7C5FA2D60F593C7" blockId="3.[151,1217,1457,1480]" box="[151,1217,1457,1480]" pageId="3" pageNumber="57">
<emphasis id="B90F5663FFDAD820E7C5FA2D650E93C8" bold="true" box="[151,314,1457,1480]" pageId="3" pageNumber="57">FIGURES 4–5.</emphasis>
<taxonomicName id="4C7BF1F2FFDAD820E612FA2E66F693C7" authority="Alexander" authorityName="Alexander" box="[320,706,1458,1480]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFDAD820E612FA2E660A93C7" box="[320,574,1458,1480]" italics="true" pageId="3" pageNumber="57">Notholimnophila exclusa</emphasis>
(Alexander)
</taxonomicName>
. Female. 4: head and thorax, lateral view. 5: wing.
</paragraph>
</caption>
<subSubSection id="C361D9FAFFDAD820E795FA686642915E" pageId="3" pageNumber="57" type="materials_examined">
<paragraph id="8BC48A71FFDAD820E795FA686649903F" blockId="3.[151,1436,1524,1873]" pageId="3" pageNumber="57">
<emphasis id="B90F5663FFDAD820E795FA6865859002" bold="true" box="[199,433,1524,1549]" pageId="3" pageNumber="57">Material examined.</emphasis>
<typeStatus id="54C034D3FFDAD820E6EBFA6966119002" box="[441,549,1525,1549]" pageId="3" pageNumber="57" type="holotype">Holotype</typeStatus>
. Male. 
<emphasis id="B90F5663FFDAD820E52BFA6967D19002" box="[633,997,1525,1549]" italics="true" pageId="3" pageNumber="57">Charteris Bay, Canterbury, N.Z.</emphasis>
[ca. 
<geoCoordinate id="EE4FECB6FFDAD820E34FFA69604D9002" box="[1053,1145,1525,1549]" direction="south" orientation="latitude" pageId="3" pageNumber="57" precision="925" value="-43.633335">43°38’S</geoCoordinate>
<geoCoordinate id="EE4FECB6FFDAD820E3D3FA6960C39002" box="[1153,1271,1525,1549]" direction="east" orientation="longitude" pageId="3" pageNumber="57" precision="925" value="172.7">172°42’ E</geoCoordinate>
], 
<emphasis id="B90F5663FFDAD820E25FFA69649B903E" italics="true" pageId="3" pageNumber="57">
alt. 
<quantity id="4C832794FFDAD820E268FA69649F903E" metricMagnitude="1" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="3" pageNumber="57" unit="ft" value="250.0" valueMax="300.0" valueMin="200.0">200–300 ft</quantity>
.
</emphasis>
[
<quantity id="4C832794FFDAD820E7E9F9846528903F" box="[187,284,1560,1584]" metricMagnitude="1" metricUnit="m" metricValue="8.1" metricValueMax="9.2" metricValueMin="7.0" pageId="3" pageNumber="57" unit="m" value="81.0" valueMax="92.0" valueMin="70.0">70–92m</quantity>
], 
<emphasis id="B90F5663FFDAD820E662F9866642903E" box="[304,630,1561,1585]" italics="true" pageId="3" pageNumber="57">
<date id="FFC5ACB1FFDAD820E662F98665FB903E" box="[304,463,1561,1585]" pageId="3" pageNumber="57" value="1921-11-12">Nov. 12, 1921</date>
, J.W.Campbell
</emphasis>
.
</paragraph>
<paragraph id="8BC48A71FFDAD820E797F9A0660C9077" blockId="3.[151,1436,1524,1873]" pageId="3" pageNumber="57">
<emphasis id="B90F5663FFDAD820E797F9A06550905A" bold="true" box="[197,356,1596,1621]" pageId="3" pageNumber="57">Preservation.</emphasis>
The male 
<typeStatus id="54C034D3FFDAD820E68FF9A1660B905A" box="[477,575,1597,1621]" pageId="3" pageNumber="57" type="holotype">holotype</typeStatus>
is pinned and in virtual perfect condition, with only the left flagellum missing. The specimen was not dissected.
</paragraph>
<paragraph id="8BC48A71FFDAD820E797F91A64C890CE" blockId="3.[151,1436,1524,1873]" pageId="3" pageNumber="57">
<typeStatus id="54C034D3FFDAD820E797F91A651F9091" box="[197,299,1670,1694]" pageId="3" pageNumber="57" type="paratype">Paratype</typeStatus>
. Female. 
<emphasis id="B90F5663FFDAD820E6CBF919660B9092" box="[409,575,1669,1693]" italics="true" pageId="3" pageNumber="57">Ohakune, N.Z.</emphasis>
[ca. 
<geoCoordinate id="EE4FECB6FFDAD820E528F91966E19092" box="[634,725,1669,1693]" direction="south" orientation="latitude" pageId="3" pageNumber="57" precision="925" value="-39.416668">39°25’S</geoCoordinate>
<geoCoordinate id="EE4FECB6FFDAD820E58DF91967619092" box="[735,853,1669,1693]" direction="east" orientation="longitude" pageId="3" pageNumber="57" precision="925" value="175.38333">175°23’ E</geoCoordinate>
], 
<emphasis id="B90F5663FFDAD820E43FF91967C29092" box="[877,1014,1669,1693]" italics="true" pageId="3" pageNumber="57">
Alt. 
<quantity id="4C832794FFDAD820E4CFF91967C69092" box="[925,1010,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="6.27888" pageId="3" pageNumber="57" unit="ft" value="2060.0">2060 ft</quantity>
.
</emphasis>
[
<quantity id="4C832794FFDAD820E356F919607D9092" box="[1028,1097,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" pageId="3" pageNumber="57" unit="m" value="628.0">628m</quantity>
], 
<emphasis id="B90F5663FFDAD820E332F91A61B39092" box="[1120,1415,1669,1693]" italics="true" pageId="3" pageNumber="57">
<date id="FFC5ACB1FFDAD820E332F91A61369092" box="[1120,1282,1669,1693]" pageId="3" pageNumber="57" value="1921-11-12">Nov. 12, 1921</date>
, T.R.Harris
</emphasis>
. | 
<emphasis id="B90F5663FFDAD820E7C5F93564C890CE" box="[151,252,1705,1729]" italics="true" pageId="3" pageNumber="57">Allotype.</emphasis>
</paragraph>
<paragraph id="8BC48A71FFDAD820E797F95166FE9106" blockId="3.[151,1436,1524,1873]" pageId="3" pageNumber="57">
<specimenCount id="9D7D41F8FFDAD820E797F951652490EA" box="[197,272,1741,1765]" pageId="3" pageNumber="57" type="male">1 male</specimenCount>
and 
<specimenCount id="9D7D41F8FFDAD820E615F951659990EA" box="[327,429,1741,1765]" pageId="3" pageNumber="57" type="female">1 female</specimenCount>
. 
<emphasis id="B90F5663FFDAD820E6E5F951666890EA" box="[439,604,1741,1765]" italics="true" pageId="3" pageNumber="57">Ohakune, N.Z.</emphasis>
[ca. 
<geoCoordinate id="EE4FECB6FFDAD820E5C1F95166DA90EA" box="[659,750,1741,1765]" direction="south" orientation="latitude" pageId="3" pageNumber="57" precision="925" value="-39.416668">39°25’S</geoCoordinate>
<geoCoordinate id="EE4FECB6FFDAD820E5A4F951675F90EA" box="[758,875,1741,1765]" direction="east" orientation="longitude" pageId="3" pageNumber="57" precision="925" value="175.38333">175°23’ E</geoCoordinate>
], 
<emphasis id="B90F5663FFDAD820E4D2F951603790EA" box="[896,1027,1741,1765]" italics="true" pageId="3" pageNumber="57">
alt. 
<quantity id="4C832794FFDAD820E4FEF95167CB90EA" box="[940,1023,1741,1765]" metricMagnitude="2" metricUnit="m" metricValue="6.27888" pageId="3" pageNumber="57" unit="ft" value="2060.0">2060 ft</quantity>
.
</emphasis>
[
<quantity id="4C832794FFDAD820E342F951606190EA" box="[1040,1109,1741,1765]" metricMagnitude="2" metricUnit="m" metricValue="6.279999999999999" pageId="3" pageNumber="57" unit="m" value="628.0">628m</quantity>
], 
<emphasis id="B90F5663FFDAD820E338F95266FE9106" italics="true" pageId="3" pageNumber="57">
<date id="FFC5ACB1FFDAD820E338F952613E90EA" box="[1130,1290,1741,1765]" pageId="3" pageNumber="57" value="1922-12-10">Dec. 10, 1922</date>
, T.R.Harris | 
<taxonomicName id="4C7BF1F2FFDAD820E7C5F96D66F29106" authority="Al., Det. C.P.Alexander, 1923" authorityName="Al., Det. C.P.Alexander" authorityYear="1923" box="[151,710,1777,1801]" class="Insecta" family="Limoniidae" genus="Limnophila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="exclusa">Limnophila exclusa Al., Det. C.P.Alexander, 1923</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BC48A71FFDAD820E797F8896642915E" blockId="3.[151,1436,1524,1873]" pageId="3" pageNumber="57">
<specimenCount id="9D7D41F8FFDAD820E797F88965209122" box="[197,276,1813,1837]" pageId="3" pageNumber="57" type="male">1 male</specimenCount>
. 
<emphasis id="B90F5663FFDAD820E64CF88A66519122" box="[286,613,1813,1837]" italics="true" pageId="3" pageNumber="57">
<collectingCountry id="F36CCAE1FFDAD820E64CF88A65829122" box="[286,438,1813,1837]" name="New Zealand" pageId="3" pageNumber="57">New Zealand</collectingCountry>
: S. Is., Kumara
</emphasis>
[
<geoCoordinate id="EE4FECB6FFDAD820E526F88966E59122" box="[628,721,1813,1837]" direction="south" orientation="latitude" pageId="3" pageNumber="57" precision="925" value="-42.616665">42°37’S</geoCoordinate>
<geoCoordinate id="EE4FECB6FFDAD820E58BF88967729122" box="[729,838,1813,1837]" direction="east" orientation="longitude" pageId="3" pageNumber="57" precision="925" value="171.18333">171°11’E</geoCoordinate>
], 
<emphasis id="B90F5663FFDAD820E409F8896642915E" italics="true" pageId="3" pageNumber="57">
<date id="FFC5ACB1FFDAD820E409F88960159122" box="[859,1057,1813,1837]" pageId="3" pageNumber="57" value="1929-02-23">23 february 1929</date>
, J.W.Campbell | 
<taxonomicName id="4C7BF1F2FFDAD820E38CF8896646915E" authority="Alexander. Det. G.C.Ribeiro, 2011" authorityName="Alexander. Det. G.C.Ribeiro" authorityYear="2011" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="exclusa">Notholimnophila exclusa Alexander. Det. G.C.Ribeiro, 2011</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF04DAF9FFDDD827E7C5FB7565D89311" httpUri="https://zenodo.org/record/279869/files/figure.png" pageId="4" pageNumber="58" targetBox="[151,1436,193,1234]" targetPageId="4">
<paragraph id="8BC48A71FFDDD827E7C5FB7565D89311" blockId="4.[151,1436,1257,1310]" pageId="4" pageNumber="58">
<emphasis id="B90F5663FFDDD827E7C5FB75650A92F1" bold="true" box="[151,318,1257,1280]" pageId="4" pageNumber="58">FIGURES 6–9.</emphasis>
<taxonomicName id="4C7BF1F2FFDDD827E615FB7566FB92F0" authority="Alexander" authorityName="Alexander" box="[327,719,1257,1279]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="B90F5663FFDDD827E615FB75667C92F0" box="[327,584,1257,1279]" italics="true" pageId="4" pageNumber="58">Notholimnophila exclusa</emphasis>
(Alexander)
</taxonomicName>
. Male terminalia. 6: dorsal view. 7: lateral view. 8: ventral view. 9: Detail of gonostylus, dorsal view.
</paragraph>
</caption>
<subSubSection id="C361D9FAFFDDD827E797FAD9660993E2" pageId="4" pageNumber="58" type="discussion">
<paragraph id="8BC48A71FFDDD827E797FAD9660993E2" blockId="4.[151,1437,1349,1522]" pageId="4" pageNumber="58">
<emphasis id="B90F5663FFDDD827E797FAD9650D9352" bold="true" box="[197,313,1349,1373]" pageId="4" pageNumber="58">Remarks.</emphasis>
The two specimens mentioned in the original description – one male and one female – were designated as 
<typeStatus id="54C034D3FFDDD827E7ABFAF5656F938E" box="[249,347,1385,1409]" pageId="4" pageNumber="58" type="holotype">holotype</typeStatus>
and allotype, respectively. The study of 
<bibRefCitation id="EFEAF780FFDDD827E473FAF567F3938E" author="Ribeiro" box="[801,967,1385,1409]" pageId="4" pageNumber="58" refString="Ribeiro, G. C. (2008) Phylogeny of the Limnophilinae (Limoniidae) and early evolution of the Tipulomorpha (Diptera). Invertebrate Systematics, 22, 627 - 694." type="journal article" year="2008">Ribeiro (2008)</bibRefCitation>
indicated 
<taxonomicName id="4C7BF1F2FFDDD827E36EFAF560D7938E" box="[1084,1251,1385,1409]" class="Insecta" family="Limoniidae" genus="Bergrothomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B90F5663FFDDD827E36EFAF560D7938E" box="[1084,1251,1385,1409]" italics="true" pageId="4" pageNumber="58">Bergrothomyia</emphasis>
</taxonomicName>
, a genus including three species in 
<collectingCountry id="F36CCAE1FFDDD827E6F5FA11663A93AA" box="[423,526,1421,1445]" name="Australia" pageId="4" pageNumber="58">Australia</collectingCountry>
(Victoria, New South 
<collectingCountry id="F36CCAE1FFDDD827E46EFA1167B693AA" box="[828,898,1421,1445]" name="United Kingdom" pageId="4" pageNumber="58">Wales</collectingCountry>
) and Tasmania, as the closest relative of 
<taxonomicName id="4C7BF1F2FFDDD827E7C5FA2D656193C6" box="[151,341,1457,1481]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B90F5663FFDDD827E7C5FA2D656193C6" box="[151,341,1457,1481]" italics="true" pageId="4" pageNumber="58">Notholimnophila</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C7BF1F2FFDDD827E630FA2D663D93C6" box="[354,521,1457,1481]" class="Insecta" family="Limoniidae" genus="Bergrothomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B90F5663FFDDD827E630FA2D663D93C6" box="[354,521,1457,1481]" italics="true" pageId="4" pageNumber="58">Bergrothomyia</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7BF1F2FFDDD827E5CFFA2D676F93C6" box="[669,859,1457,1481]" class="Insecta" family="Limoniidae" genus="Notholimnophila" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B90F5663FFDDD827E5CFFA2D676F93C6" box="[669,859,1457,1481]" italics="true" pageId="4" pageNumber="58">Notholimnophila</emphasis>
</taxonomicName>
mostly by having a rostrum as long as the head and the vein M bifurcated into M1 and M2.
</paragraph>
</subSubSection>
</treatment>
</document>