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<document id="C08A639CD77340FADD9455033E9FDAB2" ID-CLB-Dataset="24431" ID-DOI="10.11646/zootaxa.4718.2.4" ID-GBIF-Dataset="8325d33c-6141-4b3d-a6d4-1804926c81f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602350" ID-ZooBank="9E4F7E47-6CF3-48CC-9558-62CF7CAFB721" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578564602617" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2020" docId="03C9D347BE65E90AFF5EB700FC1BF856" docLanguage="en" docName="zootaxa.4718.2.4.pdf" docOrigin="Zootaxa 4718 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phanoperla huanghuye Chen 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="233" masterDocId="FFF0AB3FBE64E909FFC9B12DFF9FFF9E" masterDocTitle="First record of the genus Phanoperla Banks, 1938 (Plecoptera: Perlidae) from mainland China, with description of a new species" masterLastPageNumber="238" masterPageNumber="230" pageNumber="231" updateTime="1698776522284" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B9164518DD6C43635F212A903D3B116E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2B4C624F9C2ADC0AF67B5D964147D5C2">First record of the genus Phanoperla Banks, 1938 (Plecoptera: Perlidae) from mainland China, with description of a new species</mods:title>
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<mods:name id="36EBBA8FEB2A0ACD1F3A3CF08ABFC792" type="personal">
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<mods:roleTerm id="E0B3352A80692ECCA8E561970FED0CBF">Author</mods:roleTerm>
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<mods:namePart id="A6965AF2DDD5BC28B6156984EE8799AF">Chen, Zhi-Teng</mods:namePart>
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<mods:title id="BB54223CFFD886EA008CB5572348520B">Zootaxa</mods:title>
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<mods:part id="C6E9BAEAB05D5AEA1A0FBB655E747CBD">
<mods:date id="AF142025F47EFCCA1EC3B81467A659DD">2020</mods:date>
<mods:detail id="D27E59503FC2181D087874F38733F9CB" type="pubDate">
<mods:number id="F9D9D750C775257703A376671E1CA02A">2020-01-07</mods:number>
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<mods:number id="DD51F330589094A995A537BC8CE70616">4718</mods:number>
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<mods:number id="FC2CCEBDC138032D3F6916817E9BBE23">2</mods:number>
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<mods:classification id="DAEE2F4EB26893C92EBD8BD7236C050C">journal article</mods:classification>
<mods:identifier id="3FF478BB7CC94E05C3CBBBCCE9C2EBC2" type="CLB-Dataset">24431</mods:identifier>
<mods:identifier id="7ADD754CEB9EAF0FBD8A0D24E619239F" type="DOI">10.11646/zootaxa.4718.2.4</mods:identifier>
<mods:identifier id="BF7F723273AD4A7D650CF00BD7A28455" type="GBIF-Dataset">8325d33c-6141-4b3d-a6d4-1804926c81f0</mods:identifier>
<mods:identifier id="172C5FD8076F9286B4BC91F7D551BC43" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C9D347BE65E90AFF5EB700FC1BF856" ID-DOI="http://doi.org/10.5281/zenodo.5920782" ID-GBIF-Taxon="161670826" ID-Zenodo-Dep="5920782" LSID="urn:lsid:plazi:treatment:03C9D347BE65E90AFF5EB700FC1BF856" httpUri="http://treatment.plazi.org/id/03C9D347BE65E90AFF5EB700FC1BF856" lastPageId="3" lastPageNumber="233" pageId="1" pageNumber="231">
<subSubSection id="C37A31DABE65E908FF5EB700FD85F9D9" box="[151,538,1581,1608]" pageId="1" pageNumber="231" type="nomenclature">
<paragraph id="8BDF6251BE65E908FF5EB700FD85F9D9" blockId="1.[151,538,1581,1643]" box="[151,538,1581,1608]" pageId="1" pageNumber="231">
<heading id="D097D53DBE65E908FF5EB700FD85F9D9" bold="true" box="[151,538,1581,1608]" fontSize="11" level="1" pageId="1" pageNumber="231" reason="1">
<emphasis id="B914BE43BE65E908FF5EB700FD85F9D9" bold="true" box="[151,538,1581,1608]" pageId="1" pageNumber="231">
<taxonomicName id="4C6019D2BE65E908FF5EB700FE2AF9D6" ID-CoL="8NGZH" authority="Chen, 2020" authorityName="Chen" authorityYear="2020" box="[151,437,1581,1608]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="231" phylum="Arthropoda" rank="species" species="huanghuye" status="sp. nov.">
<emphasis id="B914BE43BE65E908FF5EB700FE2AF9D6" bold="true" box="[151,437,1581,1608]" italics="true" pageId="1" pageNumber="231">Phanoperla huanghuye</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2270338BE65E908FE74B700FD85F9D9" box="[445,538,1581,1607]" pageId="1" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A31DABE65E90BFF5EB77CFB8AF878" lastPageId="2" lastPageNumber="232" pageId="1" pageNumber="231" type="description">
<paragraph id="8BDF6251BE65E908FF5EB77CFE8AF9F5" blockId="1.[151,538,1581,1643]" box="[151,277,1617,1643]" pageId="1" pageNumber="231">
<figureCitation id="135B7ED4BE65E908FF5EB77CFE8FF9F5" box="[151,272,1617,1643]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="2.[151,250,1745,1770]" captionStartId-1="3.[151,250,1427,1452]" captionStartId-2="4.[151,250,1382,1407]" captionStartId-3="5.[151,250,1030,1055]" captionStartId-4="5.[151,250,2019,2044]" captionStartId-5="6.[151,250,1128,1153]" captionStartId-6="6.[151,250,1956,1981]" captionStartId-7="7.[151,250,645,670]" captionStartId-8="7.[151,250,1595,1620]" captionTargetBox-0="[165,1427,616,1613]" captionTargetBox-1="[177,1436,301,1408]" captionTargetBox-2="[179,1409,266,1267]" captionTargetBox-3="[300,1288,174,1011]" captionTargetBox-4="[324,1264,1105,1999]" captionTargetBox-5="[151,1436,174,1073]" captionTargetBox-6="[151,1436,1195,1927]" captionTargetBox-7="[314,1274,181,620]" captionTargetBox-8="[151,1436,721,1576]" captionTargetId-0="figure@2.[151,1436,596,1624]" captionTargetId-1="figure@3.[151,1436,301,1408]" captionTargetId-2="figure@4.[151,1436,249,1288]" captionTargetId-3="figure@5.[300,1288,174,1011]" captionTargetId-4="figure@5.[324,1264,1105,1999]" captionTargetId-5="figure@6.[151,1436,174,1073]" captionTargetId-6="figure@6.[151,1436,1195,1927]" captionTargetId-7="figure@7.[314,1274,181,620]" captionTargetId-8="figure@7.[151,1436,721,1576]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="6" captionTargetPageId-7="7" captionTargetPageId-8="7" captionText-0="FIGURE 2. Phanoperla huanghuye sp. nov., A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view." captionText-1="FIGURE 3. Phanoperla huanghuye sp. nov., A. Female habitus, dorsal view; B. Female terminalia, ventral view." captionText-2="FIGURE 4. Phanoperla huanghuye sp. nov., A. Male right forewing, dorsal view; B. Male right hindwing, dorsal view." captionText-3="FIGURE 5. Phanoperla huanghuye sp. nov., A. Male terminalia with one obscure sensillum basiconicum on tergum 7, dorsal view; B. Male terminalia with about 10 sensilla basiconica on tergum 7, dorsal view." captionText-4="FIGURE 6. Phanoperla huanghuye sp. nov., A. Male terminalia, ventral view; B. Male terminalia, lateral view." captionText-5="FIGURE 7. Phanoperla huanghuye sp. nov., A. Male terminalia, dorsal view; B–C. Hemitergal lobes, dorsal view." captionText-6="FIGURE 8. Phanoperla huanghuye sp. nov., A. Aedeagus, lateral view; B–C. Aedeagus, ventrolateral view; D. Aedeagus, ventral view; E. Aedeagus, dorsal view." captionText-7="FIGURE 9. Phanoperla huanghuye sp. nov., A–B. Egg, lateral view; C. Egg with apical membrane removed, dorsolateral view." captionText-8="FIGURE 10. Phanoperla huanghuye sp. nov., A. Habitat stream in Tongmu Village; Fujian Province, China. B. Light trap used to collect the specimens." figureDoi-0="http://doi.org/10.5281/zenodo.3602354" figureDoi-1="http://doi.org/10.5281/zenodo.3602356" figureDoi-2="http://doi.org/10.5281/zenodo.3602358" figureDoi-3="http://doi.org/10.5281/zenodo.3602360" figureDoi-4="http://doi.org/10.5281/zenodo.3602362" figureDoi-5="http://doi.org/10.5281/zenodo.3602364" figureDoi-6="http://doi.org/10.5281/zenodo.3602366" figureDoi-7="http://doi.org/10.5281/zenodo.3602368" figureDoi-8="http://doi.org/10.5281/zenodo.3602370" httpUri-0="https://zenodo.org/record/3602354/files/figure.png" httpUri-1="https://zenodo.org/record/3602356/files/figure.png" httpUri-2="https://zenodo.org/record/3602358/files/figure.png" httpUri-3="https://zenodo.org/record/3602360/files/figure.png" httpUri-4="https://zenodo.org/record/3602362/files/figure.png" httpUri-5="https://zenodo.org/record/3602364/files/figure.png" httpUri-6="https://zenodo.org/record/3602366/files/figure.png" httpUri-7="https://zenodo.org/record/3602368/files/figure.png" httpUri-8="https://zenodo.org/r
.
</paragraph>
<paragraph id="8BDF6251BE65E908FF5EB7B5FCEFF8DD" blockId="1.[151,1437,1688,2039]" pageId="1" pageNumber="231">
<emphasis id="B914BE43BE65E908FF5EB7B5FED9F92D" bold="true" box="[151,326,1688,1715]" pageId="1" pageNumber="231">Adult habitus.</emphasis>
General color yellow, patterned with brown (
<figureCitation id="135B7ED4BE65E908FC96B7B4FC2DF92D" box="[863,946,1689,1715]" captionStart="FIGURE 2" captionStartId="2.[151,250,1745,1770]" captionTargetBox="[165,1427,616,1613]" captionTargetId="figure@2.[151,1436,596,1624]" captionTargetPageId="2" captionText="FIGURE 2. Phanoperla huanghuye sp. nov., A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602354" httpUri="https://zenodo.org/record/3602354/files/figure.png" pageId="1" pageNumber="231">Figs. 2</figureCitation>
,
<figureCitation id="135B7ED4BE65E908FC0BB7B4FC78F92D" box="[962,999,1689,1715]" captionStart="FIGURE 3" captionStartId="3.[151,250,1427,1452]" captionTargetBox="[177,1436,301,1408]" captionTargetId="figure@3.[151,1436,301,1408]" captionTargetPageId="3" captionText="FIGURE 3. Phanoperla huanghuye sp. nov., A. Female habitus, dorsal view; B. Female terminalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602356" httpUri="https://zenodo.org/record/3602356/files/figure.png" pageId="1" pageNumber="231">3A</figureCitation>
). Biocellate; head slightly wider than pronotum, with a dark ocellate stigma and a frontal stigma (
<figureCitation id="135B7ED4BE65E908FCFFB790FC10F949" box="[822,911,1725,1751]" captionStart="FIGURE 2" captionStartId="2.[151,250,1745,1770]" captionTargetBox="[165,1427,616,1613]" captionTargetId="figure@2.[151,1436,596,1624]" captionTargetPageId="2" captionText="FIGURE 2. Phanoperla huanghuye sp. nov., A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602354" httpUri="https://zenodo.org/record/3602354/files/figure.png" pageId="1" pageNumber="231">Fig. 2C</figureCitation>
). Antenna slender and dark brown, longer than abdomen. Pronotum pale, margins and sutures dark brown, covered with obscure rugosities. Wing membrane subhyaline, veins dark brown (
<figureCitation id="135B7ED4BE65E908FE07B628FD8BF881" box="[462,532,1797,1823]" captionStart="FIGURE 4" captionStartId="4.[151,250,1382,1407]" captionTargetBox="[179,1409,266,1267]" captionTargetId="figure@4.[151,1436,249,1288]" captionTargetPageId="4" captionText="FIGURE 4. Phanoperla huanghuye sp. nov., A. Male right forewing, dorsal view; B. Male right hindwing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602358" httpUri="https://zenodo.org/record/3602358/files/figure.png" pageId="1" pageNumber="231">Fig. 4</figureCitation>
). Coxae pale; apex of femur, apex of tibia and third tarsal segment dark brown. Cercus pale, covered with thick bristles, with about 36 segments.
</paragraph>
<paragraph id="8BDF6251BE65E90BFF0EB660FE99FE69" blockId="1.[151,1437,1688,2039]" lastBlockId="2.[151,1437,153,503]" lastPageId="2" lastPageNumber="232" pageId="1" pageNumber="231">
<emphasis id="B914BE43BE65E908FF0EB660FE94F8F9" bold="true" box="[199,267,1869,1895]" pageId="1" pageNumber="231">Male.</emphasis>
Body length ca. 19.0 mm; forewings length ca. 21.0 mm, hindwings length ca. 19.0 mm
<emphasis id="B914BE43BE65E908FADFB661FA80F8F9" bold="true" box="[1302,1311,1868,1895]" pageId="1" pageNumber="231">(</emphasis>
<figureCitation id="135B7ED4BE65E908FAD6B660FAEFF8F9" box="[1311,1392,1869,1895]" captionStart="FIGURE 2" captionStartId="2.[151,250,1745,1770]" captionTargetBox="[165,1427,616,1613]" captionTargetId="figure@2.[151,1436,596,1624]" captionTargetPageId="2" captionText="FIGURE 2. Phanoperla huanghuye sp. nov., A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602354" httpUri="https://zenodo.org/record/3602354/files/figure.png" pageId="1" pageNumber="231">Figs. 2</figureCitation>
,
<figureCitation id="135B7ED4BE65E908FAB7B660FA10F8F9" box="[1406,1423,1869,1895]" captionStart="FIGURE 4" captionStartId="4.[151,250,1382,1407]" captionTargetBox="[179,1409,266,1267]" captionTargetId="figure@4.[151,1436,249,1288]" captionTargetPageId="4" captionText="FIGURE 4. Phanoperla huanghuye sp. nov., A. Male right forewing, dorsal view; B. Male right hindwing, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602358" httpUri="https://zenodo.org/record/3602358/files/figure.png" pageId="1" pageNumber="231">4</figureCitation>
). Tergum 7 with a median patch of sensilla basiconica, number of which varies among individuals, from 1 to about 10 (
<figureCitation id="135B7ED4BE65E908FF0AB6B8FE97F831" box="[195,264,1940,1967]" captionStart="FIGURE 5" captionStartId="5.[151,250,1030,1055]" captionTargetBox="[300,1288,174,1011]" captionTargetId="figure@5.[300,1288,174,1011]" captionTargetPageId="5" captionText="FIGURE 5. Phanoperla huanghuye sp. nov., A. Male terminalia with one obscure sensillum basiconicum on tergum 7, dorsal view; B. Male terminalia with about 10 sensilla basiconica on tergum 7, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602360" httpUri="https://zenodo.org/record/3602360/files/figure.png" pageId="1" pageNumber="231">Fig. 5</figureCitation>
). Median area of tergum 8 with an oval patch of sensilla basiconica, the sensilla basiconica patch sometimes interrupted medially, divided into two smaller patches. Tergum 9 with a butterfly-shaped patch of sensilla basiconica, the patch about
<date id="FFDE4491BE65E908FE01B6F0FE74F869" box="[456,491,2013,2039]" pageId="1" pageNumber="231">2X</date>
wider than the sensilla basiconica patch on tergum 8. Anterior half of tergum 10 cleft, with a membranous median area; posterior margin of tergum 10 notched, connected with a coniform membrane. Hemitergal lobes curved and contiguous; each hemitergal lobe with an inner hump covered by tiny sensilla basiconica; apical half of each hemitergal lobe strongly compressed from dorsal view, ventroapical margin fringed with a row of long, thick spines
<emphasis id="B914BE43BE66E90BFE01B029FE4EFE81" bold="true" box="[456,465,260,287]" pageId="2" pageNumber="232">(</emphasis>
<figureCitation id="135B7ED4BE66E90BFE18B028FDBFFE81" box="[465,544,260,287]" captionStart="FIGURE 5" captionStartId="5.[151,250,1030,1055]" captionTargetBox="[300,1288,174,1011]" captionTargetId="figure@5.[300,1288,174,1011]" captionTargetPageId="5" captionText="FIGURE 5. Phanoperla huanghuye sp. nov., A. Male terminalia with one obscure sensillum basiconicum on tergum 7, dorsal view; B. Male terminalia with about 10 sensilla basiconica on tergum 7, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602360" httpUri="https://zenodo.org/record/3602360/files/figure.png" pageId="2" pageNumber="232">Figs. 5</figureCitation>
,
<figureCitation id="135B7ED4BE66E90BFDE5B028FDD0FE81" box="[556,591,261,287]" captionStart="FIGURE 6" captionStartId="5.[151,250,2019,2044]" captionTargetBox="[324,1264,1105,1999]" captionTargetId="figure@5.[324,1264,1105,1999]" captionTargetPageId="5" captionText="FIGURE 6. Phanoperla huanghuye sp. nov., A. Male terminalia, ventral view; B. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3602362" httpUri="https://zenodo.org/record/3602362/files/figure.png" pageId="2" pageNumber="232">6B</figureCitation>
,
<figureCitation id="135B7ED4BE66E90BFD93B028FDF5FE81" box="[602,618,261,287]" captionStart="FIGURE 7" captionStartId="6.[151,250,1128,1153]" captionTargetBox="[151,1436,174,1073]" captionTargetId="figure@6.[151,1436,174,1073]" captionTargetPageId="6" captionText="FIGURE 7. Phanoperla huanghuye sp. nov., A. Male terminalia, dorsal view; B–C. Hemitergal lobes, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602364" httpUri="https://zenodo.org/record/3602364/files/figure.png" pageId="2" pageNumber="232">7</figureCitation>
). Metasternum, abdominal sterna 5–7 each with a median patch of dense bristles (
<figureCitation id="135B7ED4BE66E90BFF30B004FECCFEDD" box="[249,339,297,323]" captionStart="FIGURE 2" captionStartId="2.[151,250,1745,1770]" captionTargetBox="[165,1427,616,1613]" captionTargetId="figure@2.[151,1436,596,1624]" captionTargetPageId="2" captionText="FIGURE 2. Phanoperla huanghuye sp. nov., A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602354" httpUri="https://zenodo.org/record/3602354/files/figure.png" pageId="2" pageNumber="232">Fig. 2B</figureCitation>
). Paraprocts unmodified, apices dark (
<figureCitation id="135B7ED4BE66E90BFCC5B004FCF7FEDA" box="[780,872,297,324]" captionStart="FIGURE 6" captionStartId="5.[151,250,2019,2044]" captionTargetBox="[324,1264,1105,1999]" captionTargetId="figure@5.[324,1264,1105,1999]" captionTargetPageId="5" captionText="FIGURE 6. Phanoperla huanghuye sp. nov., A. Male terminalia, ventral view; B. Male terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3602362" httpUri="https://zenodo.org/record/3602362/files/figure.png" pageId="2" pageNumber="232">Fig. 6A</figureCitation>
). Aedeagus (
<figureCitation id="135B7ED4BE66E90BFC35B004FBDFFEDD" box="[1020,1088,297,323]" captionStart="FIGURE 8" captionStartId="6.[151,250,1956,1981]" captionTargetBox="[151,1436,1195,1927]" captionTargetId="figure@6.[151,1436,1195,1927]" captionTargetPageId="6" captionText="FIGURE 8. Phanoperla huanghuye sp. nov., A. Aedeagus, lateral view; B–C. Aedeagus, ventrolateral view; D. Aedeagus, ventral view; E. Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602366" httpUri="https://zenodo.org/record/3602366/files/figure.png" pageId="2" pageNumber="232">Fig. 8</figureCitation>
) mostly membranous, slightly curved ventrally, and constricted at mid-length; base of ventral aspect strongly sclerotized, connected with a small oval sclerite (
<figureCitation id="135B7ED4BE66E90BFEFBB05CFE12FE15" box="[306,397,369,395]" captionStart="FIGURE 8" captionStartId="6.[151,250,1956,1981]" captionTargetBox="[151,1436,1195,1927]" captionTargetId="figure@6.[151,1436,1195,1927]" captionTargetPageId="6" captionText="FIGURE 8. Phanoperla huanghuye sp. nov., A. Aedeagus, lateral view; B–C. Aedeagus, ventrolateral view; D. Aedeagus, ventral view; E. Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602366" httpUri="https://zenodo.org/record/3602366/files/figure.png" pageId="2" pageNumber="232">Fig. 8C</figureCitation>
); dorsal aspect with a subapical lobe; apex membranous, irregular, and concave; basal half of aedeagus covered with spinules on dorsal surface (
<figureCitation id="135B7ED4BE66E90BFD3CB0B8FCD2FE31" box="[757,845,405,431]" captionStart="FIGURE 8" captionStartId="6.[151,250,1956,1981]" captionTargetBox="[151,1436,1195,1927]" captionTargetId="figure@6.[151,1436,1195,1927]" captionTargetPageId="6" captionText="FIGURE 8. Phanoperla huanghuye sp. nov., A. Aedeagus, lateral view; B–C. Aedeagus, ventrolateral view; D. Aedeagus, ventral view; E. Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602366" httpUri="https://zenodo.org/record/3602366/files/figure.png" pageId="2" pageNumber="232">Fig. 8E</figureCitation>
); apical half of aedeagus with a continuous patch of stout spines surrounding lateral and ventral surfaces, the spine patch forming a triangular shape on ventral surface (
<figureCitation id="135B7ED4BE66E90BFF69B0F0FF65FE69" box="[160,250,477,503]" captionStart="FIGURE 8" captionStartId="6.[151,250,1956,1981]" captionTargetBox="[151,1436,1195,1927]" captionTargetId="figure@6.[151,1436,1195,1927]" captionTargetPageId="6" captionText="FIGURE 8. Phanoperla huanghuye sp. nov., A. Aedeagus, lateral view; B–C. Aedeagus, ventrolateral view; D. Aedeagus, ventral view; E. Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3602366" httpUri="https://zenodo.org/record/3602366/files/figure.png" pageId="2" pageNumber="232">Fig. 8D</figureCitation>
).
</paragraph>
<caption id="DF1F32D9BE66E90BFF5EB7FCFEEDF890" ID-DOI="http://doi.org/10.5281/zenodo.3602354" ID-Zenodo-Dep="3602354" httpUri="https://zenodo.org/record/3602354/files/figure.png" pageId="2" pageNumber="232" startId="2.[151,250,1745,1770]" targetBox="[165,1427,616,1613]" targetPageId="2">
<paragraph id="8BDF6251BE66E90BFF5EB7FCFEEDF890" blockId="2.[151,1436,1745,1806]" pageId="2" pageNumber="232">
<emphasis id="B914BE43BE66E90BFF5EB7FCFE8CF974" bold="true" box="[151,275,1745,1770]" pageId="2" pageNumber="232">FIGURE 2.</emphasis>
<taxonomicName id="4C6019D2BE66E90BFED3B7FFFD98F974" authority="Chen, 2020" authorityName="Chen" authorityYear="2020" box="[282,519,1746,1770]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="232" phylum="Arthropoda" rank="species" species="huanghuye" status="sp. nov.">
<emphasis id="B914BE43BE66E90BFED3B7FFFD98F974" box="[282,519,1746,1770]" italics="true" pageId="2" pageNumber="232">Phanoperla huanghuye</emphasis>
</taxonomicName>
<emphasis id="B914BE43BE66E90BFDC7B7FFFDC1F974" bold="true" box="[526,606,1746,1770]" pageId="2" pageNumber="232">
<taxonomicNameLabel id="A2270338BE66E90BFDC7B7FFFDC1F974" box="[526,606,1746,1770]" pageId="2" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Male habitus, dorsal view; B. Male head and pronotum, dorsal view; C. Male habitus, ventral view.
</paragraph>
</caption>
<paragraph id="8BDF6251BE66E90BFF0EB610FA93F800" blockId="2.[151,1437,1852,2022]" pageId="2" pageNumber="232">
<emphasis id="B914BE43BE66E90BFF0EB610FEBAF8C9" bold="true" box="[199,293,1853,1879]" pageId="2" pageNumber="232">Female.</emphasis>
Body length ca. 23.0 mm; forewings length ca. 27.0 mm, hindwings length ca. 24.0 mm
<emphasis id="B914BE43BE66E90BFAE2B611FAABF8C9" bold="true" box="[1323,1332,1852,1879]" pageId="2" pageNumber="232">(</emphasis>
<figureCitation id="135B7ED4BE66E90BFAFDB611FA10F8C9" box="[1332,1423,1852,1879]" captionStart="FIGURE 3" captionStartId="3.[151,250,1427,1452]" captionTargetBox="[177,1436,301,1408]" captionTargetId="figure@3.[151,1436,301,1408]" captionTargetPageId="3" captionText="FIGURE 3. Phanoperla huanghuye sp. nov., A. Female habitus, dorsal view; B. Female terminalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602356" httpUri="https://zenodo.org/record/3602356/files/figure.png" pageId="2" pageNumber="232">Fig. 3A</figureCitation>
). General color and pattern similar to male. Posterior half of sternum 8 concaved medially, with a sclerotized inner margin, embracing a pale, hexagonal subgenital plate
<emphasis id="B914BE43BE66E90BFD3AB6A9FD63F801" bold="true" box="[755,764,1924,1951]" pageId="2" pageNumber="232">(</emphasis>
<figureCitation id="135B7ED4BE66E90BFD35B6A9FCCCF801" box="[764,851,1924,1951]" captionStart="FIGURE 3" captionStartId="3.[151,250,1427,1452]" captionTargetBox="[177,1436,301,1408]" captionTargetId="figure@3.[151,1436,301,1408]" captionTargetPageId="3" captionText="FIGURE 3. Phanoperla huanghuye sp. nov., A. Female habitus, dorsal view; B. Female terminalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3602356" httpUri="https://zenodo.org/record/3602356/files/figure.png" pageId="2" pageNumber="232">Fig. 3B</figureCitation>
). Paraprocts membranous, unmodified.
</paragraph>
<paragraph id="8BDF6251BE66E90BFF0EB684FB8AF878" blockId="2.[151,1437,1852,2022]" pageId="2" pageNumber="232">
<emphasis id="B914BE43BE66E90BFF0EB684FF62F85D" bold="true" box="[199,253,1961,1987]" pageId="2" pageNumber="232">Egg.</emphasis>
Egg length ca.
<quantity id="4C98CFB4BE66E90BFE60B684FD9EF85D" box="[425,513,1961,1987]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="232" unit="mm" value="800.0">800 µm</quantity>
, equatorial width ca.
<quantity id="4C98CFB4BE66E90BFD27B685FCDBF85D" box="[750,836,1960,1987]" metricMagnitude="-1" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="232" unit="mm" value="680.0">680 µm</quantity>
<emphasis id="B914BE43BE66E90BFC83B685FCCCF85D" bold="true" box="[842,851,1960,1987]" pageId="2" pageNumber="232">(</emphasis>
<figureCitation id="135B7ED4BE66E90BFC9DB685FC09F85D" box="[852,918,1960,1987]" captionStart="FIGURE 9" captionStartId="7.[151,250,645,670]" captionTargetBox="[314,1274,181,620]" captionTargetId="figure@7.[314,1274,181,620]" captionTargetPageId="7" captionText="FIGURE 9. Phanoperla huanghuye sp. nov., A–B. Egg, lateral view; C. Egg with apical membrane removed, dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.3602368" httpUri="https://zenodo.org/record/3602368/files/figure.png" pageId="2" pageNumber="232">Fig. 9</figureCitation>
). Color dark brown, oval in shape; collar dark, wide and short. Chorion covered throughout with uniform, circular punctations.
</paragraph>
</subSubSection>
<subSubSection id="C37A31DABE67E90AFF0EB1B4FE92FE81" pageId="3" pageNumber="233" type="materials_examined">
<paragraph id="8BDF6251BE67E90AFF0EB1B4FE92FE81" blockId="3.[151,1436,152,287]" pageId="3" pageNumber="233">
<emphasis id="B914BE43BE67E90AFF0EB1B4FEEAFF2D" bold="true" box="[199,373,152,179]" pageId="3" pageNumber="233">Type material.</emphasis>
<materialsCitation id="3B08680CBE67E90AFEB2B1B5FD66FF65" ID-GBIF-Occurrence="2545448264" collectingDate="2019-07-23" country="China" county="Wuyishan City" elevation="800" latitude="27.7527" location="Tongmu Village" longLatPrecision="6" longitude="117.6882" municipality="Xingcun Town" pageId="3" pageNumber="233" specimenCount="1" stateProvince="Fujian" typeStatus="holotype">
<typeStatus id="54DBDCF3BE67E90AFEB2B1B5FE77FF2C" box="[379,488,152,178]" pageId="3" pageNumber="233" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="F37722C1BE67E90AFDFCB1B5FDE3FF2C" box="[565,636,152,178]" name="China" pageId="3" pageNumber="233">China</collectingCountry>
:
<collectingRegion id="49A4ACB3BE67E90AFD4FB1B5FCA4FF2C" box="[646,827,152,178]" country="China" name="Fujian" pageId="3" pageNumber="233">Fujian Province</collectingRegion>
,
<collectingCounty id="62BE1ADDBE67E90AFC8CB1B5FC72FF2C" box="[837,1005,152,178]" pageId="3" pageNumber="233">Wuyishan City</collectingCounty>
,
<collectingMunicipality id="6BBBF82BBE67E90AFC31B1B5FB3FFF2D" box="[1016,1184,152,179]" pageId="3" pageNumber="233">Xingcun Town</collectingMunicipality>
,
<location id="8EBF348ABE67E90AFB62B1B4FAFDFF2C" LSID="urn:lsid:plazi:treatment:03C9D347BE65E90AFF5EB700FC1BF856:8EBF348ABE67E90AFB62B1B4FAFDFF2C" box="[1195,1378,152,179]" country="China" county="Wuyishan City" latitude="27.7527" longLatPrecision="6" longitude="117.6882" municipality="Xingcun Town" name="Tongmu Village" pageId="3" pageNumber="233" stateProvince="Fujian">Tongmu Village</location>
, near
<location id="8EBF348ABE67E90AFF5EB190FEA6FF48" LSID="urn:lsid:plazi:treatment:03C9D347BE65E90AFF5EB700FC1BF856:8EBF348ABE67E90AFF5EB190FEA6FF48" box="[151,313,188,215]" country="China" county="Wuyishan City" latitude="27.7527" longLatPrecision="6" longitude="117.6882" municipality="Xingcun Town" name="Mt. Wuyishan" pageId="3" pageNumber="233" stateProvince="Fujian">Mt. Wuyishan</location>
, unnamed stream (
<figureCitation id="135B7ED4BE67E90AFDC5B191FDEBFF49" box="[524,628,188,215]" captionStart="FIGURE 10" captionStartId="7.[151,250,1595,1620]" captionTargetBox="[151,1436,721,1576]" captionTargetId="figure@7.[151,1436,721,1576]" captionTargetPageId="7" captionText="FIGURE 10. Phanoperla huanghuye sp. nov., A. Habitat stream in Tongmu Village; Fujian Province, China. B. Light trap used to collect the specimens." figureDoi="http://doi.org/10.5281/zenodo.3602370" httpUri="https://zenodo.org/record/3602370/files/figure.png" pageId="3" pageNumber="233">Fig. 10A</figureCitation>
),
<geoCoordinate id="EE540496BE67E90AFD41B191FC9DFF49" box="[648,770,188,215]" degrees="27.7527" direction="north" orientation="latitude" pageId="3" pageNumber="233" precision="5" value="27.7527">27.7527 N</geoCoordinate>
,
<geoCoordinate id="EE540496BE67E90AFCC4B191FC10FF49" box="[781,911,188,215]" degrees="117.6882" direction="east" orientation="longitude" pageId="3" pageNumber="233" precision="5" value="117.6882">117.6882 E</geoCoordinate>
,
<quantity id="4C98CFB4BE67E90AFC55B190FC41FF49" box="[924,990,189,215]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="233" unit="m" value="800.0">
<elevation id="004D8562BE67E90AFC55B190FC41FF49" box="[924,990,189,215]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="233" unit="m" value="800.0">800m</elevation>
</quantity>
,
<collectingMethod id="52211A46BE67E90AFC23B191FBECFF49" box="[1002,1139,188,215]" pageId="3" pageNumber="233">by light trap</collectingMethod>
(
<figureCitation id="135B7ED4BE67E90AFB4BB191FB76FF49" box="[1154,1257,188,215]" captionStart="FIGURE 10" captionStartId="7.[151,250,1595,1620]" captionTargetBox="[151,1436,721,1576]" captionTargetId="figure@7.[151,1436,721,1576]" captionTargetPageId="7" captionText="FIGURE 10. Phanoperla huanghuye sp. nov., A. Habitat stream in Tongmu Village; Fujian Province, China. B. Light trap used to collect the specimens." figureDoi="http://doi.org/10.5281/zenodo.3602370" httpUri="https://zenodo.org/record/3602370/files/figure.png" pageId="3" pageNumber="233">Fig. 10B</figureCitation>
),
<date id="FFDE4491BE67E90AFB34B191FA07FF49" box="[1277,1432,188,215]" pageId="3" pageNumber="233" value="2019-07-23">
<collectingDate id="EF9ABD79BE67E90AFB34B191FA07FF49" box="[1277,1432,188,215]" pageId="3" pageNumber="233" value="2019-07-23">July 23, 2019</collectingDate>
</date>
, Yi-Fei Huang, Yu-Xuan Hu, Xiao-Han Ye (ICJUST)
</materialsCitation>
.
<materialsCitation id="3B08680CBE67E90AFCCFB1CCFE96FE81" ID-GBIF-Occurrence="2545448265" collectingDate="2019-07-23" country="China" county="Wuyishan City" elevation="800" latitude="27.7527" location="Tongmu Village" longLatPrecision="6" longitude="117.6882" municipality="Xingcun Town" pageId="3" pageNumber="233" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Fujian" typeStatus="paratype">
<typeStatus id="54DBDCF3BE67E90AFCCFB1CCFCE8FF65" box="[774,887,225,251]" pageId="3" pageNumber="233" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D66A9D8BE67E90AFC4CB1CCFC41FF64" box="[901,990,224,251]" pageId="3" pageNumber="233" type="male">4 males</specimenCount>
and
<specimenCount id="9D66A9D8BE67E90AFBD2B1CDFB11FF64" box="[1051,1166,224,250]" pageId="3" pageNumber="233" type="female">6 females</specimenCount>
, same data as holotype (ICJUST)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF1F32D9BE67E90AFF5EB4BEFA86FA32" ID-DOI="http://doi.org/10.5281/zenodo.3602356" ID-Zenodo-Dep="3602356" box="[151,1305,1427,1452]" httpUri="https://zenodo.org/record/3602356/files/figure.png" pageId="3" pageNumber="233" startId="3.[151,250,1427,1452]" targetBox="[177,1436,301,1408]" targetPageId="3">
<paragraph id="8BDF6251BE67E90AFF5EB4BEFA86FA32" blockId="3.[151,1305,1427,1452]" box="[151,1305,1427,1452]" pageId="3" pageNumber="233">
<emphasis id="B914BE43BE67E90AFF5EB4BEFE8CFA32" bold="true" box="[151,275,1427,1452]" pageId="3" pageNumber="233">FIGURE 3.</emphasis>
<taxonomicName id="4C6019D2BE67E90AFED0B4B9FD99FA32" authority="Chen, 2020" authorityName="Chen" authorityYear="2020" box="[281,518,1428,1452]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="huanghuye" status="sp. nov.">
<emphasis id="B914BE43BE67E90AFED0B4B9FD99FA32" box="[281,518,1428,1452]" italics="true" pageId="3" pageNumber="233">Phanoperla huanghuye</emphasis>
</taxonomicName>
<emphasis id="B914BE43BE67E90AFDC4B4B9FDC2FA32" bold="true" box="[525,605,1428,1452]" pageId="3" pageNumber="233">
<taxonomicNameLabel id="A2270338BE67E90AFDC4B4B9FDC2FA32" box="[525,605,1428,1452]" pageId="3" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Female habitus, dorsal view; B. Female terminalia, ventral view.
</paragraph>
</caption>
<subSubSection id="C37A31DABE67E90AFF0EB4F6FC05F986" pageId="3" pageNumber="233" type="etymology">
<paragraph id="8BDF6251BE67E90AFF0EB4F6FC05F986" blockId="3.[151,1437,1498,1992]" pageId="3" pageNumber="233">
<emphasis id="B914BE43BE67E90AFF0EB4F6FED7FA6B" bold="true" box="[199,328,1499,1525]" pageId="3" pageNumber="233">Etymology</emphasis>
. The specific epithet is a combination of surnames of the three collectors, Yi-Fei Huang, Yu-Xuan Hu, Xiao-Han Ye, who generously provided the specimens for study.
</paragraph>
</subSubSection>
<subSubSection id="C37A31DABE67E90AFF0EB70EFC1BF856" pageId="3" pageNumber="233" type="discussion">
<paragraph id="8BDF6251BE67E90AFF0EB70EFC1BF856" blockId="3.[151,1437,1498,1992]" pageId="3" pageNumber="233">
<emphasis id="B914BE43BE67E90AFF0EB70EFEA4F9A3" bold="true" box="[199,315,1571,1597]" pageId="3" pageNumber="233">Remarks.</emphasis>
The new species can be distinguished from the two known Chinese congeners by the sensilla basiconica patches on abdominal terga 7–9, the specially modified hemitergal lobes, and the unique aedeagal structure (see fig.
<quantity id="4C98CFB4BE67E90AFF30B746FEADF91A" box="[249,306,1642,1669]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="233" unit="in" value="10.0">10 in</quantity>
Zwick 1982, and figs.
<quantity id="4C98CFB4BE67E90AFDFFB746FDE2F91A" box="[566,637,1642,1669]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="3" pageNumber="233" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">1–2 in</quantity>
<bibRefCitation id="EFF11FA0BE67E90AFD4DB747FCAFF91B" author="Li, W. H. &amp; Qin, X." box="[644,816,1642,1669]" pageId="3" pageNumber="233" pagination="193 - 196" refId="ref2200" refString="Li, W. H. &amp; Qin, X. (2016) The second species of Phanoperla (Plecoptera: Perlidae) from China, P. hainana sp. nov., from Hainan Island. Zootaxa, 4162 (1), 193 - 196. https: // doi. org / 10.11646 / zootaxa. 4162.1.12" type="journal article" year="2016">Li &amp; Qin 2016</bibRefCitation>
). When compared with all congeners of
<taxonomicName id="4C6019D2BE67E90AFB3CB747FAE6F91A" authorityName="Banks" authorityYear="1938" box="[1269,1401,1642,1668]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B914BE43BE67E90AFB3CB747FAE6F91A" box="[1269,1401,1642,1668]" italics="true" pageId="3" pageNumber="233">Phanoperla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6019D2BE67E90AFA41B746FE8DF936" authorityName="Chen" authorityYear="2020" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="huanghuye">
<emphasis id="B914BE43BE67E90AFA41B746FE8DF936" italics="true" pageId="3" pageNumber="233">P. huanghuye</emphasis>
</taxonomicName>
is also distinctive by the combination of following characters: 1) male terga 7–9 with sensilla basiconica; 2) male hemitergal lobes with basal spinulose callus and ventroapical fringe of long, thick spines; 3) aedeagus with a subapical dorsal lobe and a membranous, concave apex; 4) subgenital plate of the female near hexagonal; 5) egg oval with a short collar, chorion covered with circular punctations. These unique characters make it difficult to assign
<taxonomicName id="4C6019D2BE67E90AFF05B632FEFDF8A6" authorityName="Chen" authorityYear="2020" box="[204,354,1822,1848]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="huanghuye">
<emphasis id="B914BE43BE67E90AFF05B632FEFDF8A6" box="[204,354,1822,1848]" italics="true" pageId="3" pageNumber="233">P. huanghuye</emphasis>
</taxonomicName>
to any of the species groups defined by Zwick (1982). In addition, CuP of forewings of
<taxonomicName id="4C6019D2BE67E90AFAFBB632FF53F8C2" authorityName="Chen" authorityYear="2020" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="species" species="huanghuye">
<emphasis id="B914BE43BE67E90AFAFBB632FF53F8C2" italics="true" pageId="3" pageNumber="233">P. huanghuye</emphasis>
</taxonomicName>
is straight basally, rather than curved as commonly found in other congeners (Zwick 1982). Although the three
<taxonomicName id="4C6019D2BE67E90AFF5EB64BFE84F81E" authorityName="Banks" authorityYear="1938" box="[151,283,1894,1920]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B914BE43BE67E90AFF5EB64BFE84F81E" box="[151,283,1894,1920]" italics="true" pageId="3" pageNumber="233">Phanoperla</emphasis>
</taxonomicName>
species of
<collectingCountry id="F37722C1BE67E90AFE53B64BFE42F81E" box="[410,477,1894,1920]" name="China" pageId="3" pageNumber="233">China</collectingCountry>
are currently known only from
<collectingRegion id="49A4ACB3BE67E90AFC8FB64BFC11F81E" box="[838,910,1894,1920]" country="China" name="Fujian" pageId="3" pageNumber="233">Fujian</collectingRegion>
and
<collectingRegion id="49A4ACB3BE67E90AFC0CB64BFB89F81E" box="[965,1046,1894,1920]" country="China" name="Hainan" pageId="3" pageNumber="233">Hainan</collectingRegion>
provinces (
<figureCitation id="135B7ED4BE67E90AFB53B64BFB43F81F" box="[1178,1244,1894,1921]" captionStart="FIGURE 1" captionStartId="1.[151,250,1402,1427]" captionTargetBox="[151,1436,174,1380]" captionTargetId="figure@1.[151,1436,174,1383]" captionTargetPageId="1" captionText="FIGURE 1. Type localities of species of Phanoperla from China." figureDoi="http://doi.org/10.5281/zenodo.3602352" httpUri="https://zenodo.org/record/3602352/files/figure.png" pageId="3" pageNumber="233">Fig. 1</figureCitation>
), more members will certainly be found in other coastal provinces such as
<collectingRegion id="49A4ACB3BE67E90AFCD2B6A7FC3EF83A" box="[795,929,1930,1956]" country="China" name="Guangdong" pageId="3" pageNumber="233">Guangdong</collectingRegion>
,
<collectingRegion id="49A4ACB3BE67E90AFC65B6A7FB90F83A" box="[940,1039,1930,1956]" country="China" name="Guangxi" pageId="3" pageNumber="233">Guangxi</collectingRegion>
, and
<collectingRegion id="49A4ACB3BE67E90AFB80B6A6FB00F83B" box="[1097,1183,1931,1957]" country="China" name="Yunnan" pageId="3" pageNumber="233">Yunnan</collectingRegion>
provinces considering the known high biodiversity of
<taxonomicName id="4C6019D2BE67E90AFE3EB683FDE4F856" authorityName="Banks" authorityYear="1938" box="[503,635,1966,1992]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B914BE43BE67E90AFE3EB683FDE4F856" box="[503,635,1966,1992]" italics="true" pageId="3" pageNumber="233">Phanoperla</emphasis>
</taxonomicName>
in the Oriental Region.
</paragraph>
</subSubSection>
</treatment>
</document>
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