treatments-xml/data/03/81/87/038187A36112FFAEFDA4FC9F7ADFFB1B.xml

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<document id="860B2C6D9F49093628569D7F6BE03BB7" ID-DOI="10.1080/00222933.2010.536266" ID-ISSN="1464-5262" ID-Zenodo-Dep="5203483" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629048403400" checkinUser="felipe" docAuthor="Lee, Jimin &amp; Chang, Cheon Young" docDate="2011" docId="038187A36112FFAEFDA4FC9F7ADFFB1B" docLanguage="en" docName="JNatHist.45.13-14.855-867.pdf" docOrigin="Journal of Natural History 45 (13 - 14)" docSource="http://dx.doi.org/10.1080/00222933.2010.536266" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Lepidodasys laeviacus Lee &amp; Chang 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="861" masterDocId="FFB8FFDB6116FFA6FFA9FF867B09FFA3" masterDocTitle="Two new species of the genus Lepidodasys (Gastrotricha, Lepidodasyidae) from Tsushima Island, Japan" masterLastPageNumber="867" masterPageNumber="855" pageNumber="857" updateTime="1699057561851" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="564F35B497356DCA4D9944EBB7DACDE9">Two new species of the genus Lepidodasys (Gastrotricha, Lepidodasyidae) from Tsushima Island, Japan</mods:title>
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<mods:namePart id="648643E17F83BAD845EF13C4B602D9C4">Chang, Cheon Young</mods:namePart>
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<mods:title id="8B826E1954C47023D1226DD796BC811A">Journal of Natural History</mods:title>
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<mods:date id="4C145089C0B842805755A4693E6507E5">2011</mods:date>
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<mods:number id="E537E0C68B4A10DF3B2D865E0FB3CCBE">13 - 14</mods:number>
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<paragraph id="8B9736B56112FFA2FDA4FC9F7852FC92" blockId="4.[252,1131,729,849]" box="[525,859,793,818]" pageId="4" pageNumber="857">
<heading id="D0DF81D96112FFA2FDA4FC9F7852FC92" box="[525,859,793,818]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="857" reason="3">
<taxonomicName id="4C284D366112FFA2FDA4FC9F79F6FC91" ID-CoL="3TBPT" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[525,767,793,818]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="4" pageNumber="857" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA76112FFA2FDA4FC9F79F6FC91" bold="true" box="[525,767,793,818]" italics="true" pageId="4" pageNumber="857">Lepidodasys laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC6112FFA2FCAFFC9C7852FC92" box="[774,859,794,817]" pageId="4" pageNumber="857" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C332653E6112FFAEFDC0FCBF7F7BFCFA" lastPageId="8" lastPageNumber="861" pageId="4" pageNumber="857" type="description">
<paragraph id="8B9736B56112FFA2FDC0FCBF79F7FCF2" blockId="4.[252,1131,729,849]" box="[617,766,825,849]" pageId="4" pageNumber="857">
(
<figureCitation id="13132A306112FFA2FDD9FCBF79D4FCF2" box="[624,733,825,849]" captionStart="Figure 2" captionStartId="5.[144,211,1462,1484]" captionTargetBox="[152,1100,161,1430]" captionTargetId="figure-6@5.[144,1101,161,1430]" captionTargetPageId="5" captionText="Figure 2. Lepidodasys laeviacus sp. nov. (A) Habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) caudal region, dorsal; (E) head and pharyngeal region, ventral; (F) caudal region, ventral. Scale bars: A, B, 100 µm; CF, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5203485" httpUri="https://zenodo.org/record/5203485/files/figure.png" pageId="4" pageNumber="857">Figures 2</figureCitation>
,
<figureCitation id="13132A306112FFA2FD41FCBF79FCFCF2" box="[744,757,825,849]" captionStart="Figure 3" captionStartId="6.[213,280,1228,1250]" captionTargetBox="[213,1170,160,1195]" captionTargetId="figure-5@6.[213,1170,160,1195]" captionTargetPageId="6" captionText="Figure 3. Lepidodasys laeviacus sp. nov. (AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="4" pageNumber="857">3</figureCitation>
)
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFCF67A6DFC2B" blockId="4.[213,356,880,904]" box="[213,356,880,904]" pageId="4" pageNumber="857">
<emphasis id="B95CEAA76112FFA2FF7CFCF67A6DFC2B" box="[213,356,880,904]" italics="true" pageId="4" pageNumber="857">Type locality</emphasis>
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFC1178C3FC0C" blockId="4.[213,970,915,943]" box="[213,970,915,943]" pageId="4" pageNumber="857">
<materialsCitation id="3B403CE86112FFA2FF7CFC1178CCFC0C" box="[213,965,915,943]" country="Japan" latitude="34.672222" location="Miuda Beach" longLatPrecision="19" longitude="129.485" municipality="Tsushima Island" pageId="4" pageNumber="857" specimenCount="1">
<location id="8EF7606E6112FFA2FF7CFC117A79FC0C" LSID="urn:lsid:plazi:treatment:038187A36112FFAEFDA4FC9F7ADFFB1B:8EF7606E6112FFA2FF7CFC117A79FC0C" box="[213,368,919,943]" country="Japan" latitude="34.672222" longLatPrecision="19" longitude="129.485" municipality="Tsushima Island" name="Miuda Beach" pageId="4" pageNumber="857">Miuda Beach</location>
,
<collectingMunicipality id="6BF3ACCF6112FFA2FED2FC11793EFC0C" box="[379,567,919,943]" pageId="4" pageNumber="857">Tsushima Island</collectingMunicipality>
,
<collectingCountry id="F33F76256112FFA2FDEBFC1E798DFC0C" box="[578,644,920,943]" name="Japan" pageId="4" pageNumber="857">Japan</collectingCountry>
(
<geoCoordinate id="EE1C50726112FFA2FD3BFC117814FC0C" box="[658,797,915,943]" degrees="34" direction="north" minutes="40" orientation="latitude" pageId="4" pageNumber="857" precision="15" seconds="20" value="34.672222">
34
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40
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20
<superScript id="7C5D9BFD6112FFA2FD5DFC1579F7FC07" attach="left" box="[756,766,915,932]" fontSize="7" pageId="4" pageNumber="857"></superScript>
N
</geoCoordinate>
,
<geoCoordinate id="EE1C50726112FFA2FC81FC1178B4FC0C" box="[808,957,915,943]" degrees="129" direction="east" minutes="29" orientation="longitude" pageId="4" pageNumber="857" precision="15" seconds="06" value="129.485">
129
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29
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06
<superScript id="7C5D9BFD6112FFA2FC3FFC1578A9FC07" attach="left" box="[918,928,915,932]" fontSize="7" pageId="4" pageNumber="857"></superScript>
E
</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFC797A89FBB5" blockId="4.[213,384,1023,1046]" box="[213,384,1023,1046]" pageId="4" pageNumber="857">
<emphasis id="B95CEAA76112FFA2FF7CFC797A89FBB5" box="[213,384,1023,1046]" italics="true" pageId="4" pageNumber="857">Type specimens</emphasis>
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFBA37A36FB3E" blockId="4.[213,1170,1061,1181]" pageId="4" pageNumber="857">
<materialsCitation id="3B403CE86112FFA2FF7CFBA37A36FB3E" collectingDate="2009-10-01" collectionCode="DBG, NHMUK" collectorName="J. M. Lee &amp; C. Y. Chang. Another" pageId="4" pageNumber="857" specimenCode="DBG1401, DBG1402-09, NHMUK 2010.1, 2010.2" specimenCount="2" typeStatus="holotype">
<typeStatus id="549388176112FFA2FF7CFBA37A49FB9E" box="[213,320,1061,1085]" pageId="4" pageNumber="857" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DB8E9ECE6112FFA2FEFBFBA37ACFFB9E" box="[338,454,1061,1085]" collectionCode="DBG" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12925" lsid="urn:lsid:biocol.org:col:12925" name="Denver Botanic Gardens" pageId="4" pageNumber="857" type="Botanical Garden (with or without associated herbarium)">DBG1401</specimenCode>
) and
<specimenCount id="9D2EFD3C6112FFA2FDA4FBA379B2FB9D" box="[525,699,1061,1086]" pageId="4" pageNumber="857" type="generic" typeStatus="paratypes">eight paratypes</specimenCount>
(
<specimenCode id="DB8E9ECE6112FFA2FD64FBA37864FB9E" box="[717,877,1061,1085]" collectionCode="DBG" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12925" lsid="urn:lsid:biocol.org:col:12925" name="Denver Botanic Gardens" pageId="4" pageNumber="857" type="Botanical Garden (with or without associated herbarium)">DBG140209</specimenCode>
), mounted in glycerine,
<date id="FF9610756112FFA2FB2DFBA37A7CFBFE" pageId="4" pageNumber="857" value="2009-10-01">
<collectingDate id="EFD2E99D6112FFA2FB2DFBA37A7CFBFE" pageId="4" pageNumber="857" value="2009-10-01">1 October 2009</collectingDate>
</date>
, leg.
<collectorName id="26DD53636112FFA2FE1EFBC07915FBFE" box="[439,540,1094,1117]" pageId="4" pageNumber="857">J.M. Lee</collectorName>
and
<collectorName id="26DD53636112FFA2FDF4FBC3785CFBFE" box="[605,853,1093,1117]" pageId="4" pageNumber="857">C.Y. Chang. Another</collectorName>
<specimenCount id="9D2EFD3C6112FFA2FCC9FBC07F0BFBFD" box="[864,1026,1094,1118]" pageId="4" pageNumber="857" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(NHM reg. nos.
<specimenCode id="DB8E9ECE6112FFA2FEA2FBE07ADBFBDE" box="[267,466,1125,1149]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="857" type="Museum">NHMUK 2010.1</specimenCode>
,
<specimenCode id="DB8E9ECE6112FFA2FE76FBE37924FBDE" box="[479,557,1125,1149]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="857" type="Museum">2010.2</specimenCode>
), mounted in glycerine, collection data same as in the
<typeStatus id="549388176112FFA2FF7CFB037A32FB3E" box="[213,315,1157,1181]" pageId="4" pageNumber="857" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFB6A7915FAA7" blockId="4.[213,540,1260,1284]" box="[213,540,1260,1284]" pageId="4" pageNumber="857">
<emphasis id="B95CEAA76112FFA2FF7CFB6A7915FAA7" box="[213,540,1260,1284]" italics="true" pageId="4" pageNumber="857">Additional material examined</emphasis>
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFA9579E1FAE8" blockId="4.[213,1170,1299,1355]" pageId="4" pageNumber="857">Five specimens with the same collection data were mounted on an aluminium stub for observation on a scanning electron microscope.</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFA1C7A48FA12" blockId="4.[213,321,1434,1457]" box="[213,321,1434,1457]" pageId="4" pageNumber="857">
<emphasis id="B95CEAA76112FFA2FF7CFA1C7A48FA12" box="[213,321,1434,1457]" italics="true" pageId="4" pageNumber="857">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B9736B56112FFA2FF7CFA4778FDF9FA" blockId="4.[213,1170,1473,1625]" pageId="4" pageNumber="857">
<taxonomicName id="4C284D366112FFA2FF7CFA477A57FA7A" authorityName="Remane" authorityYear="1926" box="[213,350,1473,1497]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="4" pageNumber="857" phylum="Gastrotricha" rank="genus">
<emphasis id="B95CEAA76112FFA2FF7CFA477A57FA7A" box="[213,350,1473,1497]" italics="true" pageId="4" pageNumber="857">Lepidodasys</emphasis>
</taxonomicName>
with strap-shaped body; dorsal cuticular armature with herring-bone pattern of scales; scales elongated oval, lacking longitudinal keel; adhesive tubes of four to seven TbA per side forming an arc, six to ten TbD arranged in zigzag in intestinal region, 914 TbL per side inserted in intestinal region, round caudum furnished with 812 TbP; testes paired, anterior end slightly exceeding middle of body.
</paragraph>
<caption id="DF57663D6113FFA3FF39FA30782FF9A4" ID-DOI="http://doi.org/10.5281/zenodo.5203485" ID-Zenodo-Dep="5203485" httpUri="https://zenodo.org/record/5203485/files/figure.png" pageId="5" pageNumber="858" startId="5.[144,211,1462,1484]" targetBox="[152,1100,161,1430]" targetPageId="5">
<paragraph id="8B9736B56113FFA3FF39FA30782FF9A4" blockId="5.[144,1101,1462,1543]" pageId="5" pageNumber="858">
Figure 2.
<taxonomicName id="4C284D366113FFA3FF51FA317ADBFA6E" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[248,466,1463,1485]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="5" pageNumber="858" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA76113FFA3FF51FA317ADBFA6E" box="[248,466,1463,1485]" italics="true" pageId="5" pageNumber="858">Lepidodasys laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC6113FFA3FE71FA31792DFA6F" box="[472,548,1463,1484]" pageId="5" pageNumber="858" rank="species">sp. nov.</taxonomicNameLabel>
(A) Habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) caudal region, dorsal; (E) head and pharyngeal region, ventral; (F) caudal region, ventral. Scale bars: A, B, 100 µm; CF, 50 µm.
</paragraph>
</caption>
<caption id="DF57663D6110FFA0FF7CFB4A7A41FAF5" ID-DOI="http://doi.org/10.5281/zenodo.5203489" ID-Zenodo-Dep="5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="6" pageNumber="859" startId="6.[213,280,1228,1250]" targetBox="[213,1170,160,1195]" targetPageId="6">
<paragraph id="8B9736B56110FFA0FF7CFB4A7A41FAF5" blockId="6.[213,1170,1228,1367]" pageId="6" pageNumber="859">
Figure 3.
<taxonomicName id="4C284D366110FFA0FEE9FB4A7917FB41" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[320,542,1228,1250]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="6" pageNumber="859" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA76110FFA0FEE9FB4A7917FB41" box="[320,542,1228,1250]" italics="true" pageId="6" pageNumber="859">Lepidodasys laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC6110FFA0FD8EFB4A797EFB42" box="[551,631,1228,1249]" pageId="6" pageNumber="859" rank="species">sp. nov.</taxonomicNameLabel>
(AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm.
</paragraph>
</caption>
<paragraph id="8B9736B56110FFA0FF7CFAF17AF7FA2C" blockId="6.[213,510,1399,1423]" box="[213,510,1399,1423]" pageId="6" pageNumber="859">
<emphasis id="B95CEAA76110FFA0FF7CFAF17AF7FA2C" box="[213,510,1399,1423]" italics="true" pageId="6" pageNumber="859">
Description of the
<typeStatus id="549388176110FFA0FE09FAF17AF7FA2C" box="[416,510,1399,1423]" pageId="6" pageNumber="859" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9736B56110FFA0FF7CFA187F76F995" blockId="6.[213,1170,1438,1590]" pageId="6" pageNumber="859">
<emphasis id="B95CEAA76110FFA0FF7CFA187A06FA15" box="[213,271,1438,1462]" italics="true" pageId="6" pageNumber="859">Body</emphasis>
(
<figureCitation id="13132A306110FFA0FEB7FA187AB6FA15" box="[286,447,1438,1462]" captionStart="Figure 2" captionStartId="5.[144,211,1462,1484]" captionTargetBox="[152,1100,161,1430]" captionTargetId="figure-6@5.[144,1101,161,1430]" captionTargetPageId="5" captionText="Figure 2. Lepidodasys laeviacus sp. nov. (A) Habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) caudal region, dorsal; (E) head and pharyngeal region, ventral; (F) caudal region, ventral. Scale bars: A, B, 100 µm; CF, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5203485" httpUri="https://zenodo.org/record/5203485/files/figure.png" pageId="6" pageNumber="859">Figures 2A, B</figureCitation>
,
<figureCitation id="13132A306110FFA0FE65FA187907FA15" box="[460,526,1438,1462]" captionStart="Figure 3" captionStartId="6.[213,280,1228,1250]" captionTargetBox="[213,1170,160,1195]" captionTargetId="figure-5@6.[213,1170,160,1195]" captionTargetPageId="6" captionText="Figure 3. Lepidodasys laeviacus sp. nov. (AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="6" pageNumber="859">3A, B</figureCitation>
). Elongated, vermiform, vaulted dorsally, with indistinct head, Lt 550 µm long; a little swollen at posterior part of intestinal region, then gradually narrowing in caudal region; caudum with simple rounded end; pharynx 100 µm long, PhJIn at U19, pharyngeal pores not seen; widths of anterior head
<emphasis id="B95CEAA76110FFA0FBAEFA787F1AF9B5" bold="true" box="[1031,1043,1534,1558]" italics="true" pageId="6" pageNumber="859">/</emphasis>
pharyngeal region
<emphasis id="B95CEAA76110FFA0FEB4F9987A20F995" bold="true" box="[285,297,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
trunk
<emphasis id="B95CEAA76110FFA0FEC1F9987A7DF995" bold="true" box="[360,372,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
caudal portion 28
<emphasis id="B95CEAA76110FFA0FD95F9987941F995" bold="true" box="[572,584,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
40
<emphasis id="B95CEAA76110FFA0FDCAF9987966F995" bold="true" box="[611,623,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
50
<emphasis id="B95CEAA76110FFA0FD23F998799FF995" bold="true" box="[650,662,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
26 µm at U02
<emphasis id="B95CEAA76110FFA0FC9AF9987836F995" bold="true" box="[819,831,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
U16
<emphasis id="B95CEAA76110FFA0FCD9F9987875F995" bold="true" box="[880,892,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
U63
<emphasis id="B95CEAA76110FFA0FC04F99878B0F995" bold="true" box="[941,953,1566,1590]" italics="true" pageId="6" pageNumber="859">/</emphasis>
U95, respectively.
</paragraph>
<paragraph id="8B9736B56110FFA0FF7CF9E77F98F93B" blockId="6.[213,1170,1632,1689]" pageId="6" pageNumber="859">
<emphasis id="B95CEAA76110FFA0FF7CF9E77A65F9DA" box="[213,364,1633,1657]" italics="true" pageId="6" pageNumber="859">Sensory hairs</emphasis>
(ca. 69 µm long). Scattered sporadically on anterodorsal surface of head at U01U03; numerous hairs (ca. 1218 µm long) irregularly implanted on
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FF207A11FF7D" blockId="7.[144,1100,166,222]" pageId="7" pageNumber="860">subdorsal, lateral and ventrolateral surfaces throughout almost whole body length (U06U95).</paragraph>
<paragraph id="8B9736B56111FFA1FF39FE8D79FCFD21" blockId="7.[144,1101,267,642]" pageId="7" pageNumber="860">
<emphasis id="B95CEAA76111FFA1FF39FE8D7A6DFE80" box="[144,356,267,291]" italics="true" pageId="7" pageNumber="860">Cuticular armature</emphasis>
(
<figureCitation id="13132A306111FFA1FEDDFE8D7919FE80" box="[372,528,267,291]" captionStart="Figure 2" captionStartId="5.[144,211,1462,1484]" captionTargetBox="[152,1100,161,1430]" captionTargetId="figure-6@5.[144,1101,161,1430]" captionTargetPageId="5" captionText="Figure 2. Lepidodasys laeviacus sp. nov. (A) Habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) caudal region, dorsal; (E) head and pharyngeal region, ventral; (F) caudal region, ventral. Scale bars: A, B, 100 µm; CF, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5203485" httpUri="https://zenodo.org/record/5203485/files/figure.png" pageId="7" pageNumber="860">Figures 2CF</figureCitation>
,
<figureCitation id="13132A306111FFA1FDB5FE8D7935FE80" box="[540,572,267,291]" captionStart="Figure 3" captionStartId="6.[213,280,1228,1250]" captionTargetBox="[213,1170,160,1195]" captionTargetId="figure-5@6.[213,1170,160,1195]" captionTargetPageId="6" captionText="Figure 3. Lepidodasys laeviacus sp. nov. (AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="7" pageNumber="860">3E</figureCitation>
). Body densely covered with elongated oval or wholegrain-shaped scales dorsally, laterally and mid-ventrally; scales smooth, lacking longitudinal keel, with both ends a little protruded; 1214 slender scales surrounding mouth opening. Dorsal scales 46 µm in diameter and 1315 µm long in middle of trunk region; dorsal and lateral scales arranged in rather irregular herring-bone pattern. Ventral scales situated between ciliary bands, aligned in two columns showing zigzag pattern regularly, anterior ends of scales pointing outward, behind TbA to level of last pair of ventral cilia (at U03U71); anterior scales small (3 × 7 µm), increasing in size toward end of ciliary bands (5 × 15 µm at U70); ventrolateral scales beside ciliary bands much narrower and elongated (ranging from 2 × 10 µm to 4 × 20 µm) than in scales between ciliary bands; caudum covered densely and irregularly with different size of scales, as in the case of dorsal and lateral scales.
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FD297ADDFD44" blockId="7.[144,1101,687,743]" pageId="7" pageNumber="860">
<emphasis id="B95CEAA76111FFA1FF39FD297A4EFD64" box="[144,327,687,711]" italics="true" pageId="7" pageNumber="860">Ventral ciliation.</emphasis>
Aligned in a pair of ciliary bands from just behind TbA (U03) to two thirds of trunk region (U71).
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FC9278F0FC48" blockId="7.[144,1101,788,1003]" pageId="7" pageNumber="860">
<emphasis id="B95CEAA76111FFA1FF39FC927A32FC8F" box="[144,315,788,812]" italics="true" pageId="7" pageNumber="860">Adhesive tubes.</emphasis>
TbA of four or five tubes per side, ca. 57 µm long, forming an arc behind oral opening at U02U05, of which three or four tubes gathered ventrally, rearmost one a little apart from others. TbD of nine tubes, ca. 1418 µm long, irregularly arranged in intestinal region (U29U84), beside median line; not paired, except for third pair. TbL of nine to ten tubes per side, ranging from 11 to 19 µm long, situated behind PhJIn to posterior trunk region at U23U89, asymmetrically. TbP of eight tubes ranging ca. 916 µm long, inserted along round caudum unevenly.
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FB9E7A23FB13" blockId="7.[144,1101,1048,1200]" pageId="7" pageNumber="860">
<emphasis id="B95CEAA76111FFA1FF39FB9E7A90FB93" box="[144,409,1048,1072]" italics="true" pageId="7" pageNumber="860">Reproductive apparatus.</emphasis>
Five suboval eggs with different sizes situated dorsally in midintestinal region at U60U78, of which the anteriormost is largest (ca. 23 × 46 µm) and well-developed, becoming smaller posteriorly. Testes paired, their anterior ends slightly exceeding middle of body (U47); vas deferens not visible. Accessory organs not observed.
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FB7D7ADFFAB1" blockId="7.[144,470,1274,1298]" box="[144,470,1274,1298]" pageId="7" pageNumber="860">
<emphasis id="B95CEAA76111FFA1FF39FB7D7ADFFAB1" box="[144,470,1274,1298]" italics="true" pageId="7" pageNumber="860">Measurements and variability</emphasis>
</paragraph>
<paragraph id="8B9736B56111FFA1FF39FAA47AF8FA39" blockId="7.[144,1101,1314,1689]" pageId="7" pageNumber="860">
Among
<specimenCount id="9D2EFD3C6111FFA1FF47FAA47A48FA98" box="[238,321,1314,1339]" pageId="7" pageNumber="860" type="generic" typeStatus="type">11 type</specimenCount>
specimens, only one was immature, 543 µm in length and 42 µm in width. The immature specimen showed two discrepancies from adults, that is, fewer dorsal and posterior adhesive tubes of three TbD and six TbP, respectively, and an absence of reproductive organs.
</paragraph>
<paragraph id="8B9736B56111FFA1FF12FA277F44FA7A" blockId="7.[144,1101,1314,1689]" pageId="7" pageNumber="860">
Body lengths of
<specimenCount id="9D2EFD3C6111FFA1FE2AFA277AEDFA1A" box="[387,484,1441,1465]" pageId="7" pageNumber="860" type="adult">10 adult</specimenCount>
<typeStatus id="549388176111FFA1FE59FA247929FA19" box="[496,544,1442,1466]" pageId="7" pageNumber="860">type</typeStatus>
specimens ranged from 515 to 675 µm (mean 573 µm, SD 43), maximum widths 5171 µm (08U13U), when mounted in glycerine.
</paragraph>
<paragraph id="8B9736B56111FFA1FF12FA67786BF93A" blockId="7.[144,1101,1314,1689]" pageId="7" pageNumber="860">
The number and arrangement of adhesive tubes were somewhat variable, ranging from four to seven TbA per side, 914 TbL per side, 710 TbD and 712 TbP, respectively. Six of
<specimenCount id="9D2EFD3C6111FFA1FE82F9A77A89F999" box="[299,384,1569,1594]" pageId="7" pageNumber="860" type="generic" typeStatus="type">11 type</specimenCount>
specimens had paired TbD, implanted usually third to sixth, excluding first tube paired in the
<specimenCount id="9D2EFD3C6111FFA1FDA1F9C779ADF9FA" box="[520,676,1601,1625]" pageId="7" pageNumber="860" type="generic">one specimen</specimenCount>
. One specimen had two pairs of TbD at fourth and fifth. TbL array was asymmetrical except in
<specimenCount id="9D2EFD3C6111FFA1FCBCF9E778A5F9DB" box="[789,940,1632,1656]" pageId="7" pageNumber="860" type="generic">one specimen</specimenCount>
and several of their tubes were located on the ventrolateral trunk near the eggs.
</paragraph>
<paragraph id="8B9736B5611EFFAEFF7CFF207A44FF1D" blockId="8.[213,333,166,190]" box="[213,333,166,190]" pageId="8" pageNumber="861">
<emphasis id="B95CEAA7611EFFAEFF7CFF207A44FF1D" box="[213,333,166,190]" italics="true" pageId="8" pageNumber="861">Etymology</emphasis>
</paragraph>
<paragraph id="8B9736B5611EFFAEFF7CFF4879E4FEA5" blockId="8.[213,1170,206,262]" pageId="8" pageNumber="861">
The specific name,
<taxonomicName id="4C284D36611EFFAEFE02FF487907FF45" authorityName="Lee &amp; Chang" authorityYear="2011" box="[427,526,206,230]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="laeviacus">
<emphasis id="B95CEAA7611EFFAEFE02FF487907FF45" box="[427,526,206,230]" italics="true" pageId="8" pageNumber="861">laeviacus</emphasis>
</taxonomicName>
, alludes to smooth (L.
<emphasis id="B95CEAA7611EFFAEFCBDFF487858FF45" box="[788,849,206,230]" italics="true" pageId="8" pageNumber="861">laevis</emphasis>
) husk (L.
<emphasis id="B95CEAA7611EFFAEFC6FFF4978FFFF45" box="[966,1014,207,230]" italics="true" pageId="8" pageNumber="861">acus</emphasis>
), which is the characteristic shape of scales of the new species.
</paragraph>
<paragraph id="8B9736B5611EFFAEFF7CFEDA7A3EFED7" blockId="8.[213,311,348,372]" box="[213,311,348,372]" pageId="8" pageNumber="861">
<emphasis id="B95CEAA7611EFFAEFF7CFEDA7A3EFED7" box="[213,311,348,372]" italics="true" pageId="8" pageNumber="861">Remarks</emphasis>
</paragraph>
<paragraph id="8B9736B5611EFFAEFF7CFE057AC9FDF9" blockId="8.[213,1170,387,1208]" pageId="8" pageNumber="861">
Since the genus
<taxonomicName id="4C284D36611EFFAEFE08FE057923FE38" authorityName="Remane" authorityYear="1926" box="[417,554,387,411]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="genus">
<emphasis id="B95CEAA7611EFFAEFE08FE057923FE38" box="[417,554,387,411]" italics="true" pageId="8" pageNumber="861">Lepidodasys</emphasis>
</taxonomicName>
was first established by
<bibRefCitation id="EFB94B44611EFFAEFCCBFE027F1EFE38" author="Remane A" box="[866,1047,387,411]" pageId="8" pageNumber="861" pagination="625 - 754" refId="ref5312" refString="Remane A. 1926. Morphologie und Verwandtschaftsbeziehungen der aberranten Gastrotrichen I. Z Morph Oekol Tiere. 5: 625 - 754." type="journal article" year="1926">Remane (1926)</bibRefCitation>
, only five species have been recorded:
<taxonomicName id="4C284D36611EFFAEFDB6FE22784AFE18" authority="Remane, 1926" authorityName="Remane" authorityYear="1926" box="[543,835,419,443]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFDB6FE22799AFE18" box="[543,659,420,443]" italics="true" pageId="8" pageNumber="861">L. martini</emphasis>
<bibRefCitation id="EFB94B44611EFFAEFD37FE22784AFE18" author="Remane A" box="[670,835,419,443]" pageId="8" pageNumber="861" pagination="625 - 754" refId="ref5312" refString="Remane A. 1926. Morphologie und Verwandtschaftsbeziehungen der aberranten Gastrotrichen I. Z Morph Oekol Tiere. 5: 625 - 754." type="journal article" year="1926">Remane, 1926</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C284D36611EFFAEFCFAFE227F84FE18" authority="Remane, 1927" authorityName="Remane" authorityYear="1927" box="[851,1165,419,443]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="platyurus">
<emphasis id="B95CEAA7611EFFAEFCFAFE2278D7FE18" box="[851,990,419,443]" italics="true" pageId="8" pageNumber="861">L. platyurus</emphasis>
<bibRefCitation id="EFB94B44611EFFAEFC41FE227F84FE18" author="Remane A" box="[1000,1165,419,443]" pageId="8" pageNumber="861" pagination="203 - 242" refId="ref5336" refString="Remane A. 1927. Neue Gastrotricha Macrodasyoidea. Zool Jahrb Abt Syst. 54: 203 - 242." type="journal article" year="1927">Remane, 1927</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C284D36611EFFAEFF7CFE427824FE78" authority="Balsamo, Fregni and Tongiorgi, 1994" authorityName="Balsamo, Fregni and Tongiorgi" authorityYear="1994" box="[213,813,451,475]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="unicarenatus">
<emphasis id="B95CEAA7611EFFAEFF7CFE427A89FE78" box="[213,384,452,475]" italics="true" pageId="8" pageNumber="861">L. unicarenatus</emphasis>
<bibRefCitation id="EFB94B44611EFFAEFE2EFE457824FE78" author="Balsamo M &amp; Fregni E &amp; Tongiorgi P" box="[391,813,451,475]" pageId="8" pageNumber="861" pagination="217 - 227" refId="ref4921" refString="Balsamo M, Fregni E, Tongiorgi P. 1994. Marine and freshwater Gastrotricha from the Island of Montecristo (Tuscan Archipelago, Italy), with the description of new species. Boll Zool. 61: 217 - 227." type="journal article" year="1994">Balsamo, Fregni and Tongiorgi, 1994</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C284D36611EFFAEFC90FE427F69FE78" authority="Clausen, 2004" authorityName="Clausen" authorityYear="2004" box="[825,1120,451,475]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="arcolepis">
<emphasis id="B95CEAA7611EFFAEFC90FE4278B3FE78" box="[825,954,451,475]" italics="true" pageId="8" pageNumber="861">L. arcolepis</emphasis>
<bibRefCitation id="EFB94B44611EFFAEFC68FE457F69FE78" author="Clausen C" box="[961,1120,451,475]" pageId="8" pageNumber="861" pagination="423 - 458" refId="ref5025" refString="Clausen C. 2004. Gastrotricha from the Faroe Bank. Sarsia 89: 423 - 458." type="journal article" year="2004">Clausen, 2004</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C284D36611EFFAEFF7CFE62791EFE58" authority="Clausen, 2004" authorityName="Clausen" authorityYear="2004" box="[213,535,483,507]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="castoroides">
<emphasis id="B95CEAA7611EFFAEFF7CFE627A66FE58" box="[213,367,483,507]" italics="true" pageId="8" pageNumber="861">L. castoroides</emphasis>
<bibRefCitation id="EFB94B44611EFFAEFEDCFE65791EFE58" author="Clausen C" box="[373,535,483,507]" pageId="8" pageNumber="861" pagination="423 - 458" refId="ref5025" refString="Clausen C. 2004. Gastrotricha from the Faroe Bank. Sarsia 89: 423 - 458." type="journal article" year="2004">Clausen, 2004</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C284D36611EFFAEFD89FE6579F4FE58" box="[544,765,483,507]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="undetermined">
<emphasis id="B95CEAA7611EFFAEFD89FE6579A0FE58" box="[544,681,483,507]" italics="true" pageId="8" pageNumber="861">Lepidodasys</emphasis>
species
</taxonomicName>
have been grouped by two main criteria: whether the scales are arranged in herring-bone or parallel style, and whether the scales bear longitudinal keel(s) or not (
<bibRefCitation id="EFB94B44611EFFAEFD6EFDA578B9FD98" author="Balsamo M &amp; Fregni E &amp; Tongiorgi P" box="[711,944,547,571]" pageId="8" pageNumber="861" pagination="217 - 227" refId="ref4921" refString="Balsamo M, Fregni E, Tongiorgi P. 1994. Marine and freshwater Gastrotricha from the Island of Montecristo (Tuscan Archipelago, Italy), with the description of new species. Boll Zool. 61: 217 - 227." type="journal article" year="1994">Balsamo et al. 1994</bibRefCitation>
;
<bibRefCitation id="EFB94B44611EFFAEFC17FDA57F84FD98" author="Fregni E &amp; Faienza MG &amp; De Zio Grimaldi &amp; Tongiorgi P &amp; Balsamo M" box="[958,1165,547,571]" pageId="8" pageNumber="861" pagination="183 - 194" refId="ref5065" refString="Fregni E, Faienza MG, De Zio Grimaldi S, Tongiorgi P, Balsamo M. 1999. Marine gastrotrichs from the Tremiti Archipelago in the southern Adriatic Sea, with the description of two new species of of Urodasys. Ital J Zool. 66: 183 - 194." type="journal article" year="1999">Fregni et al. 1999</bibRefCitation>
;
<bibRefCitation id="EFB94B44611EFFAEFF7CFDC47A66FDF9" author="Clausen C" box="[213,367,578,602]" pageId="8" pageNumber="861" pagination="357 - 384" refId="ref5003" refString="Clausen C. 2000. Gastrotricha Macrodasyida from the Tromso region, northern Norway. Sarsia 85: 357 - 384." type="journal article" year="2000">Clausen 2000</bibRefCitation>
,
<bibRefCitation id="EFB94B44611EFFAEFED3FDC47ABDFDF9" author="Clausen C" box="[378,436,578,602]" pageId="8" pageNumber="861" pagination="423 - 458" refId="ref5025" refString="Clausen C. 2004. Gastrotricha from the Faroe Bank. Sarsia 89: 423 - 458." type="journal article" year="2004">2004</bibRefCitation>
).
</paragraph>
<paragraph id="8B9736B5611EFFAEFEA9FDE57980FD79" blockId="8.[213,1170,387,1208]" pageId="8" pageNumber="861">
<taxonomicName id="4C284D36611EFFAEFEA9FDE57AFFFDD8" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[256,502,611,635]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA7611EFFAEFEA9FDE57AFFFDD8" box="[256,502,611,635]" italics="true" pageId="8" pageNumber="861">Lepidodasys laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC611EFFAEFDA9FDE57951FDD9" box="[512,600,611,634]" pageId="8" pageNumber="861" rank="species">sp. nov.</taxonomicNameLabel>
shares the herring-bone disposition of the scales with
<taxonomicName id="4C284D36611EFFAEFEB8FD057A8CFD39" authorityName="Remane" authorityYear="1926" box="[273,389,643,666]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFEB8FD057A8CFD39" box="[273,389,643,666]" italics="true" pageId="8" pageNumber="861">L. martini</emphasis>
</taxonomicName>
. In addition, the new species is closely similar to
<taxonomicName id="4C284D36611EFFAEFC78FD057F4DFD39" authorityName="Remane" authorityYear="1926" box="[977,1092,643,666]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFC78FD0578EAFD39" box="[977,995,643,666]" italics="true" pageId="8" pageNumber="861">L</emphasis>
.
<emphasis id="B95CEAA7611EFFAEFC5DFD057F4DFD39" box="[1012,1092,643,666]" italics="true" pageId="8" pageNumber="861">martini</emphasis>
</taxonomicName>
in the arrangements of the ventral scales between ciliary bands, the anterior adhesive tubes, and the (ventro-)lateral adhesive tubes.
</paragraph>
<paragraph id="8B9736B5611EFFAEFEA9FD657F7BFCFA" blockId="8.[213,1170,387,1208]" pageId="8" pageNumber="861">
However,
<taxonomicName id="4C284D36611EFFAEFED9FD657AE9FD59" authorityName="Remane" authorityYear="1926" box="[368,480,739,762]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFED9FD657A8BFD59" box="[368,386,739,762]" italics="true" pageId="8" pageNumber="861">L</emphasis>
.
<emphasis id="B95CEAA7611EFFAEFE39FD657AE9FD59" box="[400,480,739,762]" italics="true" pageId="8" pageNumber="861">martini</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C284D36611EFFAEFDDAFD6579FFFD59" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[627,758,738,762]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA7611EFFAEFDDAFD6579FFFD59" box="[627,758,738,762]" italics="true" pageId="8" pageNumber="861">L. laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC611EFFAEFD54FD65785BFD59" box="[765,850,739,762]" pageId="8" pageNumber="861" rank="species">sp. nov.</taxonomicNameLabel>
by the absence of the dorsal adhesive tubes in the trunk region (six to ten TbD in
<taxonomicName id="4C284D36611EFFAEFCB2FC857895FCB9" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[795,924,770,794]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA7611EFFAEFCB2FC857824FCB9" box="[795,813,771,794]" italics="true" pageId="8" pageNumber="861">L</emphasis>
.
<emphasis id="B95CEAA7611EFFAEFC90FC847895FCB9" box="[825,924,770,794]" italics="true" pageId="8" pageNumber="861">laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC611EFFAEFC0BFC8578F1FCB9" box="[930,1016,770,794]" pageId="8" pageNumber="861" rank="species">sp. nov.</taxonomicNameLabel>
). The decisive character discriminating the two species is the shape of scales, that is, smooth scales without any keel in
<taxonomicName id="4C284D36611EFFAEFE1BFCC5793CFCF9" authority="Lee &amp; Chang, 2011" authorityName="Lee &amp; Chang" authorityYear="2011" box="[434,565,834,858]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="laeviacus" status="sp. nov.">
<emphasis id="B95CEAA7611EFFAEFE1BFCC57ACDFCF9" box="[434,452,835,858]" italics="true" pageId="8" pageNumber="861">L</emphasis>
.
<emphasis id="B95CEAA7611EFFAEFE7BFCC4793CFCF9" box="[466,565,834,858]" italics="true" pageId="8" pageNumber="861">laeviacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F57DC611EFFAEFD92FCC4799BFCFA" box="[571,658,834,857]" pageId="8" pageNumber="861" rank="species">sp. nov.</taxonomicNameLabel>
, versus “unicarinated” scales in
<taxonomicName id="4C284D36611EFFAEFC52FCC57F62FCF9" authorityName="Remane" authorityYear="1926" box="[1019,1131,834,858]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFC52FCC57F04FCF9" box="[1019,1037,835,858]" italics="true" pageId="8" pageNumber="861">L</emphasis>
.
<emphasis id="B95CEAA7611EFFAEFBB2FCC57F62FCF9" box="[1051,1131,835,858]" italics="true" pageId="8" pageNumber="861">martini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C332653E611EFFAEFEA9FCE47ADFFB1B" pageId="8" pageNumber="861" type="reference_group">
<paragraph id="8B9736B5611EFFAEFEA9FCE47ADFFB1B" blockId="8.[213,1170,387,1208]" pageId="8" pageNumber="861">
<taxonomicName id="4C284D36611EFFAEFEA9FCE47AD6FCDA" ID-CoL="3TBPV" authorityName="Remane" authorityYear="1926" box="[256,479,866,890]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFEA9FCE47AD6FCDA" box="[256,479,866,890]" italics="true" pageId="8" pageNumber="861">Lepidodasys martini</emphasis>
</taxonomicName>
was originally described as bearing two longitudinal keels on the scales (
<bibRefCitation id="EFB94B44611EFFAEFEFDFC077AFFFC3A" author="Remane A" box="[340,502,897,921]" pageId="8" pageNumber="861" pagination="625 - 754" refId="ref5312" refString="Remane A. 1926. Morphologie und Verwandtschaftsbeziehungen der aberranten Gastrotrichen I. Z Morph Oekol Tiere. 5: 625 - 754." type="journal article" year="1926">Remane 1926</bibRefCitation>
). However, later this was corrected and reported as being furnished with only one keel following scanning electron microscopy by
<bibRefCitation id="EFB94B44611EFFAEFBA0FC277A29FC7A" author="Fregni E &amp; Faienza MG &amp; De Zio Grimaldi &amp; Tongiorgi P &amp; Balsamo M" pageId="8" pageNumber="861" pagination="183 - 194" refId="ref5065" refString="Fregni E, Faienza MG, De Zio Grimaldi S, Tongiorgi P, Balsamo M. 1999. Marine gastrotrichs from the Tremiti Archipelago in the southern Adriatic Sea, with the description of two new species of of Urodasys. Ital J Zool. 66: 183 - 194." type="journal article" year="1999">Fregni et al. (1999)</bibRefCitation>
.
<bibRefCitation id="EFB94B44611EFFAEFE85FC477ADCFC7A" author="Clausen C" box="[300,469,961,985]" pageId="8" pageNumber="861" pagination="357 - 384" refId="ref5003" refString="Clausen C. 2000. Gastrotricha Macrodasyida from the Tromso region, northern Norway. Sarsia 85: 357 - 384." type="journal article" year="2000">Clausen (2000)</bibRefCitation>
also reported
<taxonomicName id="4C284D36611EFFAEFDD5FC4479E4FC7A" ID-CoL="3TBPV" authorityName="Remane" authorityYear="1926" box="[636,749,962,985]" family="Lepidodasyidae" genus="Lepidodasys" kingdom="Animalia" order="Macrodasyida" pageId="8" pageNumber="861" phylum="Gastrotricha" rank="species" species="martini">
<emphasis id="B95CEAA7611EFFAEFDD5FC4479E4FC7A" box="[636,749,962,985]" italics="true" pageId="8" pageNumber="861">L. martini</emphasis>
</taxonomicName>
from
<collectingCountry id="F33F7625611EFFAEFC9DFC447886FC79" box="[820,911,962,986]" name="Norway" pageId="8" pageNumber="861">Norway</collectingCountry>
and
<collectingCountry id="F33F7625611EFFAEFC63FC477F41FC7A" box="[970,1096,961,985]" name="Germany" pageId="8" pageNumber="861">Heligoland</collectingCountry>
under differential interference contrast microscopy, which had “unicarinated” scales. Even in our new species, two faint longitudinal lines were sometimes observed under a light microscope at high magnification (see
<figureCitation id="13132A30611EFFAEFD68FBA7783DFB9A" box="[705,820,1057,1081]" captionStart="Figure 3" captionStartId="6.[213,280,1228,1250]" captionTargetBox="[213,1170,160,1195]" captionTargetId="figure-5@6.[213,1170,160,1195]" captionTargetPageId="6" captionText="Figure 3. Lepidodasys laeviacus sp. nov. (AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="8" pageNumber="861">Figure 3F</figureCitation>
). However, as all the scales are evidently smooth without any longitudinal keel when seen in scanning electron micrographs (see
<figureCitation id="13132A30611EFFAEFEF7FBE67ADAFBDB" box="[350,467,1120,1144]" captionStart="Figure 3" captionStartId="6.[213,280,1228,1250]" captionTargetBox="[213,1170,160,1195]" captionTargetId="figure-5@6.[213,1170,160,1195]" captionTargetPageId="6" captionText="Figure 3. Lepidodasys laeviacus sp. nov. (AE) Scanning electron micrographs: (A) habitus, dorsal; (B) habitus, ventral; (C) head and pharyngeal region, dorsal; (D) head and pharyngeal region, ventral; (E) dorsal scales in the trunk region; (F) differential interference contrast photograph: dorsal scales in the trunk region. Scale bars: A, B, 100 µm; C, D, 10 µm; E, F, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5203489" httpUri="https://zenodo.org/record/5203489/files/figure.png" pageId="8" pageNumber="861">Figure 3E</figureCitation>
), these lines may result from the inner rib(s) within the scale (not the real keels), or from traces of adherent between scales and body surface, under improper illumination.
</paragraph>
</subSubSection>
</treatment>
</document>