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<mods:title id="69A986EB72F7DC451E17BC71B4719FCA">On the genus Gastrodynerus Bohart, 1984 (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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<paragraph id="8BEDF760FFD2FF9EFF2A7503FD35342F" blockId="1.[151,713,801,828]" box="[151,713,801,828]" pageId="1" pageNumber="241">
<heading id="D0A5400CFFD2FF9EFF2A7503FD35342F" bold="true" box="[151,713,801,828]" fontSize="11" level="1" pageId="1" pageNumber="241" reason="1">
<emphasis id="B9262B72FFD2FF9EFF2A7503FD35342F" bold="true" box="[151,713,801,828]" pageId="1" pageNumber="241">
Key to species of 
<taxonomicName id="4C528CE3FFD2FF9EFED37503FD35342F" ID-CoL="4MDJ" authority="Bohart, 1984" authorityName="Bohart" authorityYear="1984" box="[366,713,801,828]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="B9262B72FFD2FF9EFED37503FDDC3428" bold="true" box="[366,544,801,828]" italics="true" pageId="1" pageNumber="241">Gastrodynerus</emphasis>
<bibRefCitation id="EFC38A91FFD2FF9EFD9A7503FD35342F" author="Bohart, R. M." box="[551,713,801,827]" pageId="1" pageNumber="241" pagination="12 - 15" refId="ref6662" refString="Bohart, R. M. (1984) Gastrodynerus, a New Genus of Eumenidae from Western North America (Hymenoptera: Eumenidae). Pan-Pacific Entomologist, 61 (1), 12 - 15." type="journal article" year="1984">Bohart, 1984</bibRefCitation>
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<paragraph id="8BEDF760FFD2FF9EFF2A7547FA60346F" blockId="1.[151,1437,869,1771]" box="[151,1436,869,892]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7547FA60346F" box="[151,1436,869,892]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7547FF56346E" bold="true" box="[151,170,869,890]" pageId="1" pageNumber="241">1.</emphasis>
T2 strongly reflexed at apex (
<figureCitation id="1369EBE5FFD2FF9EFE437547FDC43468" box="[510,568,869,892]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1893,1918]" captionTargetBox="[174,1413,166,1873]" captionTargetId="figure@5.[174,1414,166,1873]" captionTargetPageId="5" captionText="FIGURES 1–8. Habitus from species of Gastrodynerus: 1: G. vanduzeei (Holotype, female); 2: G. stangei (Holotype, male); 3: G. tacubayae (Holotype, female—photo taken by Dr. Bernard Landry—Muséum d’Histoire Naturelle de Geneve [MNHN]); 4: G. searsi (Holotype, male); 5: G. yungaensis (Holotype, female); 6: G. guatemalensis (Holotype, female); 7: G. barretti (Holotype, female); 8: G. aimara (Holotype, male). Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442388" httpUri="https://zenodo.org/record/3442388/files/figure.png" pageId="1" pageNumber="241">Fig. 2</figureCitation>
); digitus long, narrower at the base (
<figureCitation id="1369EBE5FFD2FF9EFC257547FBFA346F" box="[920,1030,869,891]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figs 22–25</figureCitation>
)................................ 2
</keyLead>
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<paragraph id="8BEDF760FFD2FF9EFF2A75A1FA60348C" blockId="1.[151,1437,869,1771]" box="[151,1436,899,921]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A75A1FA60348C" box="[151,1436,899,921]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A75A1FF4D348D" bold="true" box="[151,177,899,921]" pageId="1" pageNumber="241">1’.</emphasis>
T2 not reflexed at apex, or weakly reflexed (
<figureCitation id="1369EBE5FFD2FF9EFD3075A1FD3B348D" box="[653,711,899,921]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1893,1918]" captionTargetBox="[174,1413,166,1873]" captionTargetId="figure@5.[174,1414,166,1873]" captionTargetPageId="5" captionText="FIGURES 1–8. Habitus from species of Gastrodynerus: 1: G. vanduzeei (Holotype, female); 2: G. stangei (Holotype, male); 3: G. tacubayae (Holotype, female—photo taken by Dr. Bernard Landry—Muséum d’Histoire Naturelle de Geneve [MNHN]); 4: G. searsi (Holotype, male); 5: G. yungaensis (Holotype, female); 6: G. guatemalensis (Holotype, female); 7: G. barretti (Holotype, female); 8: G. aimara (Holotype, male). Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442388" httpUri="https://zenodo.org/record/3442388/files/figure.png" pageId="1" pageNumber="241">Fig. 3</figureCitation>
); digitus shorter, broader at the base (
<figureCitation id="1369EBE5FFD2FF9EFB8C75A1FB63348D" box="[1073,1183,899,921]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figs 26–28</figureCitation>
)................... 3
</keyLead>
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<keyStep id="30A6EFC5FFD2FF9EFF2A7582FA61335D" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A7582FA6034E5" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7582FA6034E5" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7582FF5534A1" bold="true" box="[151,169,928,949]" pageId="1" pageNumber="241">2.</emphasis>
Vertex with strong polished postocellar tubercles (
<figureCitation id="1369EBE5FFD2FF9EFD7F7582FD0134A2" box="[706,765,928,950]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 9</figureCitation>
); interocellar region elevated, forming two small tubercles adjacent to lateral ocelli; distal lobe of paramere with few erect short bristles (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFC49759FFBAE34C0" box="[1012,1106,957,980]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 22</figureCitation>
).......................................................................... 
<emphasis id="B9262B72FFD2FF9EFC9575F9FA6034E5" bold="true" box="[808,1436,987,1009]" pageId="1" pageNumber="241">
<taxonomicName id="4C528CE3FFD2FF9EFC9575F9FB4134E5" ID-CoL="3FDKM" authority="(Bohart, 1948)" authorityName="Bohart" authorityYear="1948" baseAuthorityName="Bohart" baseAuthorityYear="1948" box="[808,1213,987,1009]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="vanduzeei">
<emphasis id="B9262B72FFD2FF9EFC9575F9FBDF34E5" bold="true" box="[808,1059,987,1009]" italics="true" pageId="1" pageNumber="241">Gastrodynerus vanduzeei</emphasis>
(Bohart, 1948)
</taxonomicName>
(
<collectingCountry id="F345B7F0FFD2FF9EFB7775F9FAEB34E4" box="[1226,1303,987,1009]" name="United States of America" pageId="1" pageNumber="241">U. S. A.</collectingCountry>
and 
<collectingCountry id="F345B7F0FFD2FF9EFAF775F9FA6A34E5" box="[1354,1430,987,1009]" name="Mexico" pageId="1" pageNumber="241">Mexico</collectingCountry>
)
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A75DAFA61335D" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A75DAFA61335D" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A75DAFF4D331A" bold="true" box="[151,177,1016,1038]" pageId="1" pageNumber="241">2’.</emphasis>
Vertex with weak protuberances, but not polished tubercles (
<figureCitation id="1369EBE5FFD2FF9EFC9475DAFC8D331A" box="[809,881,1016,1038]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 10</figureCitation>
); interocellar region not elevated, without small tubercles adjacent to the lateral ocelli; distal lobe with more erect short bristles (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFBA77237FB843338" box="[1050,1144,1045,1068]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 23</figureCitation>
)..................................................................................... 
<taxonomicName id="4C528CE3FFD2FF9EFC697211FA61335D" ID-CoL="3FDKK" authority="Bohart, 1984" authorityName="Bohart" authorityYear="1984" box="[980,1437,1075,1097]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="stangei">
<emphasis id="B9262B72FFD2FF9EFC697211FA61335D" bold="true" box="[980,1437,1075,1097]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFC697211FB4E335C" bold="true" box="[980,1202,1075,1097]" italics="true" pageId="1" pageNumber="241">Gastrodynerus stangei</emphasis>
<bibRefCitation id="EFC38A91FFD2FF9EFB057211FAC0335C" author="Bohart, R. M." box="[1208,1340,1075,1096]" pageId="1" pageNumber="241" pagination="12 - 15" refId="ref6662" refString="Bohart, R. M. (1984) Gastrodynerus, a New Genus of Eumenidae from Western North America (Hymenoptera: Eumenidae). Pan-Pacific Entomologist, 61 (1), 12 - 15." type="journal article" year="1984">Bohart, 1984</bibRefCitation>
(
<collectingCountry id="F345B7F0FFD2FF9EFAF57211FA6A335D" box="[1352,1430,1075,1097]" name="Mexico" pageId="1" pageNumber="241">Mexico</collectingCountry>
)
</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30A6EFC5FFD2FF9EFF2A7272FA603397" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A7272FA603371" blockId="1.[151,1437,869,1771]" box="[151,1436,1104,1126]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7272FA603371" box="[151,1436,1104,1126]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7272FF553371" bold="true" box="[151,169,1104,1125]" pageId="1" pageNumber="241">3.</emphasis>
S3 with strong and deep punctures, some bigger than ocelli (
<figureCitation id="1369EBE5FFD2FF9EFC9A7272FC903372" box="[807,876,1104,1126]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 13</figureCitation>
)............................................ 
<emphasis id="B9262B72FFD2FF9EFA2D7272FA603371" bold="true" box="[1424,1436,1104,1125]" pageId="1" pageNumber="241">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A724FFA603397" blockId="1.[151,1437,869,1771]" box="[151,1436,1133,1156]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A724FFA603397" box="[151,1436,1133,1156]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A724FFF4D3397" bold="true" box="[151,177,1133,1155]" pageId="1" pageNumber="241">3’.</emphasis>
S3 with strong, but sparse punctures, never bigger than ocelli (
<figureCitation id="1369EBE5FFD2FF9EFC87724FFC7C3390" box="[826,896,1133,1156]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 14</figureCitation>
)........................................... 
<emphasis id="B9262B72FFD2FF9EFA2D724CFA603397" bold="true" box="[1424,1436,1134,1155]" pageId="1" pageNumber="241">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6EFC5FFD2FF9EFF2A72A9FA613227" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A72A9FA6133CF" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A72A9FA6133CF" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A72A9FF5533B4" bold="true" box="[151,169,1163,1184]" pageId="1" pageNumber="241">4.</emphasis>
Punctation of clypeus more evident, deeper, homogeneously distributed; humeral region weakly projected (more evident in males); T1 with coarse punctures that are clearly separated from each other (easiest to observe in dorsal view) (
<figureCitation id="1369EBE5FFD2FF9EFAAA728AFAA233AA" box="[1303,1374,1192,1214]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 17</figureCitation>
); base of digitus with erect and sparse bristles (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFD4772E7FCAB33CF" box="[762,855,1221,1243]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 25</figureCitation>
). 
<taxonomicName id="4C528CE3FFD2FF9EFCC872E7FA6133CF" ID-CoL="3FDKL" authority="(de Saussure, 1857) (Mexico)" authorityName="de Saussure" authorityYear="1857" baseAuthorityName="de Saussure" baseAuthorityYear="1857" box="[885,1437,1221,1243]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="tacubayae">
<emphasis id="B9262B72FFD2FF9EFCC872E7FA6133CF" bold="true" box="[885,1437,1221,1243]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFCC872E7FB8C33CF" bold="true" box="[885,1136,1221,1243]" italics="true" pageId="1" pageNumber="241">Gastrodynerus tacubayae</emphasis>
(de Saussure, 1857) (
<collectingCountry id="F345B7F0FFD2FF9EFAF572E7FA6A33CF" box="[1352,1430,1221,1243]" name="Mexico" pageId="1" pageNumber="241">Mexico</collectingCountry>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A72C1FA613227" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A72C1FA613227" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A72C1FF4D33ED" bold="true" box="[151,177,1251,1273]" pageId="1" pageNumber="241">4’.</emphasis>
Punctation of clypeus weaker, smooth, more evident at superior portion; humeral region more projected (more evident in males); T1 with coarser, wider and reticulated punctures (easier to observe in dorsal view) (
<figureCitation id="1369EBE5FFD2FF9EFBE77322FB5E3202" box="[1114,1186,1280,1302]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 16</figureCitation>
); base of digitus with tuft (denser) of erect bristles (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFDD1733FFD353220" box="[620,713,1309,1332]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 24</figureCitation>
)...................... 
<taxonomicName id="4C528CE3FFD2FF9EFC5C733FFA613227" ID-CoL="3FDKJ" authority="Bohart, 1984" authorityName="Bohart" authorityYear="1984" box="[993,1437,1309,1331]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="searsi">
<emphasis id="B9262B72FFD2FF9EFC5C733FFA613227" bold="true" box="[993,1437,1309,1331]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFC5C733FFB4D3227" bold="true" box="[993,1201,1309,1331]" italics="true" pageId="1" pageNumber="241">Gastrodynerus searsi</emphasis>
<bibRefCitation id="EFC38A91FFD2FF9EFB05733CFAC03226" author="Bohart, R. M." box="[1208,1340,1309,1331]" pageId="1" pageNumber="241" pagination="12 - 15" refId="ref6662" refString="Bohart, R. M. (1984) Gastrodynerus, a New Genus of Eumenidae from Western North America (Hymenoptera: Eumenidae). Pan-Pacific Entomologist, 61 (1), 12 - 15." type="journal article" year="1984">Bohart, 1984</bibRefCitation>
(
<collectingCountry id="F345B7F0FFD2FF9EFAF5733FFA6A3227" box="[1352,1430,1309,1331]" name="Mexico" pageId="1" pageNumber="241">Mexico</collectingCountry>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6EFC5FFD2FF9EFF2A7319FA60329E" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A7319FA60327A" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7319FA60327A" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7319FF5F3244" bold="true" box="[151,163,1339,1360]" pageId="1" pageNumber="241">5</emphasis>
Dorsal surface of propodeum with coarse, wide and reticulated punctures (almost same size as ocelli) (
<figureCitation id="1369EBE5FFD2FF9EFB5B7319FAD33245" box="[1254,1327,1339,1361]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 11</figureCitation>
); male unknown............................. 
<emphasis id="B9262B72FFD2FF9EFD29737AFA60327A" bold="true" box="[660,1436,1368,1390]" pageId="1" pageNumber="241">
<taxonomicName id="4C528CE3FFD2FF9EFD29737AFB42327A" authority="Grandinete &amp; Carpenter" authorityName="Grandinete &amp; Cortes &amp; Carpenter" authorityYear="2019" box="[660,1214,1368,1390]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="guatemalensis" status="sp. nov.">
<emphasis id="B9262B72FFD2FF9EFD29737AFC44327A" bold="true" box="[660,952,1368,1390]" italics="true" pageId="1" pageNumber="241">Gastrodynerus guatemalensis</emphasis>
Grandinete &amp; Carpenter
</taxonomicName>
, 
<taxonomicNameLabel id="A2159609FFD2FF9EFB75737AFAE93279" box="[1224,1301,1368,1389]" pageId="1" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
(
<collectingCountry id="F345B7F0FFD2FF9EFA9C737AFA69327A" box="[1313,1429,1368,1390]" name="Guatemala" pageId="1" pageNumber="241">Guatemala</collectingCountry>
)
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A7357FA60329E" blockId="1.[151,1437,869,1771]" box="[151,1436,1397,1420]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7357FA60329E" box="[151,1436,1397,1420]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7357FF55329F" bold="true" box="[151,169,1397,1419]" pageId="1" pageNumber="241">5’</emphasis>
Dorsal surface of propodeum with coarse punctures, clearly smaller and denser, but not reticulated (
<figureCitation id="1369EBE5FFD2FF9EFB1D7357FB1A3298" box="[1184,1254,1397,1420]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1929,1954]" captionTargetBox="[151,1436,181,1864]" captionTargetId="figure@6.[151,1436,181,1864]" captionTargetPageId="6" captionText="FIGURES 9–14. 9: Vertex of G. vanduzeei with the two polished tubercles; 10: Vertex of G. stangei, with no tubercles; 11: Posterior surface of propodeum from G. guatemalensis; 12: Posterior surface of propodeum from G. barretti; 13: S3 from G. vanduzeei (Holotype, female); 14: S3 from G. guatemalensis. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442390" httpUri="https://zenodo.org/record/3442390/files/figure.png" pageId="1" pageNumber="241">Fig. 12</figureCitation>
)............. 
<emphasis id="B9262B72FFD2FF9EFA2D7357FA60329E" bold="true" box="[1424,1436,1397,1418]" pageId="1" pageNumber="241">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6EFC5FFD2FF9EFF2A73B1FA60312F" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A73B1FA603115" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A73B1FA603115" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A73B1FF5532BC" bold="true" box="[151,169,1427,1448]" pageId="1" pageNumber="241">6.</emphasis>
Basal portion of T1 with strong (wider on lateral surfaces) reticulate punctures, becoming slightly separate towards apex (
<figureCitation id="1369EBE5FFD2FF9EFAC873B1FF0032D2" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 18</figureCitation>
). Males: When compared to females, the punctures on T1 even stronger and wider, becoming smaller (although still reticulate) towards apex; digitus very short, scarcely surpassing the distal lobe (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFB8773EFFB6432F0" box="[1082,1176,1485,1508]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 27</figureCitation>
)...................................................... 
<emphasis id="B9262B72FFD2FF9EFDC573C9FA603115" bold="true" box="[632,1436,1515,1537]" pageId="1" pageNumber="241">
<taxonomicName id="4C528CE3FFD2FF9EFDC573C9FBC53115" authority="Grandinete &amp; Cortes" authorityName="Grandinete &amp; Cortes &amp; Carpenter" authorityYear="2019" box="[632,1081,1515,1537]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="barretti" status="sp. nov.">
<emphasis id="B9262B72FFD2FF9EFDC573C9FCA53115" bold="true" box="[632,857,1515,1537]" italics="true" pageId="1" pageNumber="241">Gastrodynerus barretti</emphasis>
Grandinete &amp; Cortes
</taxonomicName>
, 
<taxonomicNameLabel id="A2159609FFD2FF9EFBF973C9FB6D3114" box="[1092,1169,1515,1536]" pageId="1" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
(
<collectingCountry id="F345B7F0FFD2FF9EFB2073C9FB143115" box="[1181,1256,1515,1537]" name="Bolivia" pageId="1" pageNumber="241">Bolivia</collectingCountry>
, 
<collectingCountry id="F345B7F0FFD2FF9EFB4E73C9FAA43115" box="[1267,1368,1515,1537]" name="Colombia" pageId="1" pageNumber="241">Colombia</collectingCountry>
, 
<collectingCountry id="F345B7F0FFD2FF9EFADE73C9FA6A3115" box="[1379,1430,1515,1537]" name="Peru" pageId="1" pageNumber="241">Peru</collectingCountry>
)
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A702AFA60312F" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A702AFA60312F" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A702AFF4D310A" bold="true" box="[151,177,1544,1566]" pageId="1" pageNumber="241">6’.</emphasis>
Punctures of T1 smaller, not reticulate, homogeneously distributed (
<figureCitation id="1369EBE5FFD2FF9EFCC9702AFC40310A" box="[884,956,1544,1566]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 19</figureCitation>
); digitus longer, clearly surpassing the distal lobe (male genitalia, 
<figureCitation id="1369EBE5FFD2FF9EFEC07007FDD73128" box="[381,555,1573,1596]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figures 26 and 27</figureCitation>
)....................................................................... 
<emphasis id="B9262B72FFD2FF9EFA2D7004FA60312F" bold="true" box="[1424,1436,1574,1595]" pageId="1" pageNumber="241">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6EFC5FFD2FF9EFF2A7061FA6031FF" pageId="1" pageNumber="241">
<paragraph id="8BEDF760FFD2FF9EFF2A7061FA613187" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A7061FA613187" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A7061FF55314C" bold="true" box="[151,169,1603,1624]" pageId="1" pageNumber="241">7.</emphasis>
T1 with strong and close punctures (
<figureCitation id="1369EBE5FFD2FF9EFD867061FD7D314D" box="[571,641,1603,1625]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 19</figureCitation>
); T2 with lateral margins (in dorsal view) more convex (
<figureCitation id="1369EBE5FFD2FF9EFB277061FB1D314D" box="[1178,1249,1603,1625]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 21</figureCitation>
); same tergum with weak, shallow and clearly separate punctures, becoming stronger and close to each other on pre-apical region (
<figureCitation id="1369EBE5FFD2FF9EFAAE7042FAA73162" box="[1299,1371,1632,1654]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 21</figureCitation>
); male genitalia in 
<figureCitation id="1369EBE5FFD2FF9EFEEE705FFE4C3187" box="[339,432,1661,1683]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 28</figureCitation>
................................. 
<taxonomicName id="4C528CE3FFD2FF9EFCF8705FFA613187" authority="Grandinete &amp; Cortes, 2019" authorityName="Grandinete &amp; Cortes &amp; Carpenter" authorityYear="2019" box="[837,1437,1661,1683]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="aimara">
<emphasis id="B9262B72FFD2FF9EFCF8705FFA613187" bold="true" box="[837,1437,1661,1683]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFCF8705FFBDF3187" bold="true" box="[837,1059,1661,1683]" italics="true" pageId="1" pageNumber="241">Gastrodynerus aimara</emphasis>
Grandinete &amp; Cortes, 2019 (
<collectingCountry id="F345B7F0FFD2FF9EFAF6705FFA6A3187" box="[1355,1430,1661,1683]" name="Bolivia" pageId="1" pageNumber="241">Bolivia</collectingCountry>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEDF760FFD2FF9EFF2A70B9FA6031FF" blockId="1.[151,1437,869,1771]" pageId="1" pageNumber="241">
<keyLead id="30A35455FFD2FF9EFF2A70B9FA6031FF" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFF2A70B9FF4D31A5" bold="true" box="[151,177,1691,1713]" pageId="1" pageNumber="241">7’.</emphasis>
Punctures on T1 weaker and homogeneously distributed (
<figureCitation id="1369EBE5FFD2FF9EFCAA70B9FCA331A5" box="[791,863,1691,1713]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 15</figureCitation>
); T2 with lateral margins (in dorsal view) weakly convex, almost parallel (
<figureCitation id="1369EBE5FFD2FF9EFEC0709AFE3931DA" box="[381,453,1720,1742]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 20</figureCitation>
); with weak and shallow punctures towards pre-apical region (
<figureCitation id="1369EBE5FFD2FF9EFBA2709AFB9B31DA" box="[1055,1127,1720,1742]" captionStart="FIGURES 15–21" captionStartId="7.[151,264,1971,1996]" captionTargetBox="[151,1428,181,1920]" captionTargetId="figure@7.[151,1428,181,1920]" captionText="FIGURES 15–21. 15: T1 of G. yungaensis; 16: T1 of G. searsi; 17: T1 of G. tacubayae; 18: T1 of G. barretti; 19: T1 of G. aimara; 20: T2 of G. yungaensis; 21: T2 of G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3450550" httpUri="https://zenodo.org/record/3450550/files/figure.png" pageId="1" pageNumber="241">Fig. 20</figureCitation>
); male genitalia in 
<figureCitation id="1369EBE5FFD2FF9EFAA3709AFA8731DA" box="[1310,1403,1720,1742]" captionStart="FIGURES 22–28" captionStartId="8.[151,264,1432,1457]" captionTargetBox="[151,1434,662,1413]" captionTargetId="figure@8.[151,1435,662,1413]" captionTargetPageId="8" captionText="FIGURES 22–28. Male genitalia (aedeagus and paremere + volsella, respectively). 22: G. vanduzeei; 23: G. stangei; 24: G. searsi; 25: G. tacubayae; 26: G. yungaensis; 27: G. barretti; 28: G. aimara. Scale bar = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3442392" httpUri="https://zenodo.org/record/3442392/files/figure.png" pageId="1" pageNumber="241">Figure 26</figureCitation>
........................................... 
<taxonomicName id="4C528CE3FFD2FF9EFD6570F7FA6031FF" authority="Grandinete &amp; Carpenter, 2019" authorityName="Grandinete &amp; Cortes &amp; Carpenter" authorityYear="2019" box="[728,1436,1749,1771]" class="Insecta" family="Eumenidae" genus="Gastrodynerus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="yungaensis">
<emphasis id="B9262B72FFD2FF9EFD6570F7FA6031FF" bold="true" box="[728,1436,1749,1771]" pageId="1" pageNumber="241">
<emphasis id="B9262B72FFD2FF9EFD6570F7FC2331FF" bold="true" box="[728,991,1749,1771]" italics="true" pageId="1" pageNumber="241">Gastrodynerus yungaensis</emphasis>
Grandinete &amp; Carpenter, 2019 (
<collectingCountry id="F345B7F0FFD2FF9EFA9670F7FA6A31FF" box="[1323,1430,1749,1771]" name="Argentina" pageId="1" pageNumber="241">Argentina</collectingCountry>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>