treatments-xml/data/C0/33/FC/C033FC35C25B55EF8004DE6F0194772A.xml

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<mods:title id="A0D4F3D66BE574B6FCF53115FC1A73C8">East Asian Cryphalus Erichson (Curculionidae, Scolytinae): new species, new synonymy and redescriptions of species</mods:title>
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<mods:namePart id="EDBC385FE222CD3D5F5CC33F07A61772">Johnson, Andrew J.</mods:namePart>
<mods:affiliation id="6888A15F299E0F76C3BE170DFF12CE39">School of Forest Resources and Conservation, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
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<mods:namePart id="E080E631F4EFBE9599934B5AC99DF56D">Li, You</mods:namePart>
<mods:affiliation id="24B6D430D05874DCC6AC20F6D6FC1EBC">School of Forest Resources and Conservation, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
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<mods:namePart id="93427F95148510F6AC034F37711C6739">Mandelshtam, Michail Yu.</mods:namePart>
<mods:affiliation id="B119EC5221DAFFA3965880D7933079BA">Department of Forest Protection, Wood Science and Game Management, Saint-Petersburg State Forest Technical University named after S. M. Kirov, Institutskii per., 5; 194021, Saint-Petersburg, Russia</mods:affiliation>
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<mods:namePart id="46692ED1D50AAE373B9CC4D0ADEE4672">Park, Sangwook</mods:namePart>
<mods:affiliation id="0DEDF273A4598627C9D047E391E6DA01">Research Institute of Forest Insect Diversity, Namyangju, 12113, South Korea</mods:affiliation>
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<mods:namePart id="181ACCC5F6A8FD5370CD082B0066CB00">Lin, Ching-Shan</mods:namePart>
<mods:affiliation id="4EC28F9A78976E74C07C943B51687BE1">Department of Entomology, National Taiwan University, Taipei, Taiwan</mods:affiliation>
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<mods:namePart id="AD54F26D6F1EF98CDF791F3198C5D731">Gao, Lei</mods:namePart>
<mods:affiliation id="86D5806889DA9DABD0ADCFE033C84655">Shanghai Academy of Landscape Architecture Science and Planning, Key Laboratory of National Forestry and Grassland Administration on Ecological Landscaping of Challenging Urban Sites, Shanghai 200232, China</mods:affiliation>
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<mods:namePart id="B5B050914E6AF62E384115664A390751">Hulcr, Jiri</mods:namePart>
<mods:affiliation id="2BFCBDAE8798EDC098DAEF9B915148DB">School of Forest Resources and Conservation, University of Florida, Gainesville, FL, 32611, USA &amp; Department of Entomology, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
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<treatment id="C033FC35C25B55EF8004DE6F0194772A" ID-GBIF-Taxon="170399002" LSID="urn:lsid:plazi:treatment:C033FC35C25B55EF8004DE6F0194772A" httpUri="http://treatment.plazi.org/id/C033FC35C25B55EF8004DE6F0194772A" lastPageNumber="15" pageId="0" pageNumber="15" scope_family="Curculionidae" scope_subFamily="Scolytinae" scope_tribe="Cryphalini">
<subSubSection id="4AAD636BC772F8BB13C3B8B302BB280C" pageId="0" pageNumber="15" type="nomenclature">
<paragraph id="50E08E94AEB03394C0283478A2B36749" pageId="0" pageNumber="15">
<taxonomicName id="3CECACBCEDD596FD613EBB81E09FF4B1" ID-CoL="ZSQ5" LSID="C033FC35-C25B-55EF-8004-DE6F0194772A" authority="Stebbing, 1914" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">Cryphalus mangiferae Stebbing, 1914</taxonomicName>
<figureCitation id="DC04AD908CF05AD4B59D6061F128203B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Proventriculus of A Cryphalus artocarpus, UFFE: 33165 B C. dilutus, UFFE: 34684 C C. dorsalis, UFFE: 25581 D C. gnetivorus, UFFE: 31753 E C. itinerans, UFFE: 33178 F C. kyotoensis, UFFE: 33232 G C. lipingensis, UFFE: 28048 H C. mangiferae, UFFE: 34966 I C. meridionalis, UFFE: 28061 J C. morivorus, UFFE: 27604 K C. paramangiferae, UFFE: 34965." figureDoi="10.3897/zookeys.995.55981.figure2" httpUri="https://binary.pensoft.net/fig/476775" pageId="0" pageNumber="15">Figures 2H</figureCitation>
<figureCitation id="7C91059F644E372AB5BCE1C26F9B249F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus of A Cryphalus artocarpus, UFFE: 33165 B C. dilutus, UFFE: 34684 C C. dorsalis, UFFE: 25581 D C. gnetivorus, UFFE: 31753 E C. itinerans, UFFE: 33178 F C. kyotoensis, UFFE: 31745 G C. lipingensis, UFFE: 28048 H C. mangiferae, UFFE: 34966 I C. meridionalis, UFFE: 28061 J C. morivorus, UFFE: 27604 K C. paramangiferae, UFFE: 34965." figureDoi="10.3897/zookeys.995.55981.figure3" httpUri="https://binary.pensoft.net/fig/476776" pageId="0" pageNumber="15">, 3H</figureCitation>
<figureCitation id="B341A245FC3EE95B247FAF58729D79CD" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Cryphalus mangiferae A, C, E, G female, UFFE: 33207 B, D, F, H, I male, UFFE: 33208." figureDoi="10.3897/zookeys.995.55981.figure12" httpUri="https://binary.pensoft.net/fig/476785" pageId="0" pageNumber="15">, 12A-I</figureCitation>
</paragraph>
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<subSubSection id="08BC5D0CCF4022C13BD2B75A42BC391E" pageId="0" pageNumber="15" type="reference_group">
<paragraph id="C16215DB4AF4DF7560015F4FA9169C7D" pageId="0" pageNumber="15">
<taxonomicName id="EF7706EDAF6B07B7ED08F574D83C2FB7" authorityName="Eichhoff" authorityYear="1872" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus inops" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="inops">Cryphalus inops</taxonomicName>
Eichhoff, 1872: 131 (Guadeloupe).
</paragraph>
<paragraph id="E616BDB137C2928099AD062F8D24AA80" pageId="0" pageNumber="15">
<taxonomicName id="68A14A77266E5C12BC9B5D8834F7305B" authorityName="Blackburn" authorityYear="1885" class="Insecta" family="Curculionidae" genus="Hypothenemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypothenemus griseus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="griseus">Hypothenemus griseus</taxonomicName>
Blackburn, 1885: 194 (Hawaii).
</paragraph>
<paragraph id="85547778B33CC1766D18158D009E6818" pageId="0" pageNumber="15">
<taxonomicName id="59968CD2795ACA432B453491D02BD2BA" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">Cryphalus mangiferae</taxonomicName>
Stebbing, 1914: 542 (India).
</paragraph>
<paragraph id="6F73F640153AF6B79A1B89E62605731E" pageId="0" pageNumber="15">
<taxonomicName id="4452A611479D93573F7794AE980B9B39" class="Insecta" family="Curculionidae" genus="Hypocryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">Hypocryphalus mangiferae</taxonomicName>
Eggers, 1928: 85 (Brazi).
</paragraph>
<paragraph id="F924C575104D8586FECC1C7A647BF391" pageId="0" pageNumber="15">
<taxonomicName id="97C605B85222EDA9E960A0A0275152D3" authorityName="Schedl" authorityYear="1942" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus subcylindricus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="subcylindricus">Cryphalus subcylindricus</taxonomicName>
Schedl, 1942: 16 (Indonesia).
</paragraph>
<paragraph id="0FAD018C41903F0A7556951EEA2C5EB4" pageId="0" pageNumber="15">
<taxonomicName id="514D0FAC7A51982002017DD86AE24C78" authorityName="Schedl" authorityYear="1942" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mimicus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mimicus">Cryphalus mimicus</taxonomicName>
Schedl, 1942: 17 (Indonesia).
</paragraph>
<paragraph id="F22651BD6D70D053A6D86D0EABB97333" pageId="0" pageNumber="15">
<taxonomicName id="E327DA240E213672DF4BBF8B647B0373" authorityName="Schedl" authorityYear="1942" class="Insecta" family="Curculionidae" genus="Hypocryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocryphalus opacus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="opacus">Hypocryphalus opacus</taxonomicName>
Schedl, 1942; 20 (Indonesia).
</paragraph>
<paragraph id="0FACA2AFE9744956533742C060A3EE46" pageId="0" pageNumber="15">
<taxonomicName id="DFF5CC9CE11EF666FD38C2C28A5B92C9" authorityName="Browne" authorityYear="1970" class="Insecta" family="Curculionidae" genus="Taenioglyptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taenioglyptes artestriatus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="artestriatus">Taenioglyptes artestriatus</taxonomicName>
Browne, 1970: 553 (Uganda), syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="8AA4222AB30A4CF38E019EB5FCB279D0" pageId="0" pageNumber="15" type="type material">
<paragraph id="EB329FCAB275FE153BE67B7043EB859B" pageId="0" pageNumber="15">Type material examined.</paragraph>
<paragraph id="864247A4A9B9E6CE4898E134E0B66597" pageId="0" pageNumber="15">
India • 1 ♀ 
<emphasis id="537B7540C5F8362B7EE562D89561A9E2" bold="true" italics="true" pageId="0" pageNumber="15">Lectotype</emphasis>
<taxonomicName id="D67A94011561DF1C79799C75E08076CF" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">
<emphasis id="6EAD3E895FFA232D32AF3280036A808C" italics="true" pageId="0" pageNumber="15">Cryphalus mangiferae</emphasis>
</taxonomicName>
Stebbing, 1914; 1902; labelled &quot;India// E.P.Stebing//902-309. //// mango twigs // 
<taxonomicName id="75EB941E2CD2A77B0D28EB31B8D97941" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">
<emphasis id="950207D466386B351843B4E97C96441F" italics="true" pageId="0" pageNumber="15">Cryphalus mangiferae</emphasis>
</taxonomicName>
Stb. // 
<taxonomicName id="0915F1AF2D77D4CBBFC673BF78558818" genus="C." lsidName="C." pageId="0" pageNumber="15" rank="genus">C.</taxonomicName>
Beeson det. //// Lectotype 
<taxonomicName id="278CCFE96222897FB372988E5132B982" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">
<emphasis id="1F60A2EEB3C79A6896191E9CFF4FFB07" italics="true" pageId="0" pageNumber="15">Cryphalus mangiferae</emphasis>
</taxonomicName>
// S.L.W. 1976 Stebb. //// NHMUK 010805929&quot;; UFFE:26286; (NHMUK) • 1 ♂? 
<emphasis id="A37977CF752573E6E232BCCEC0AD4BC1" bold="true" italics="true" pageId="0" pageNumber="15">Paralectotype</emphasis>
; 1902; UFFE:10426; (NHMUK).
</paragraph>
<paragraph id="E749EB1EA4079B2030F4F704C5F557F3" pageId="0" pageNumber="15">
Uganda • 1 ♀ 
<emphasis id="BCA383DC0CEC8EF39104A5A147C70EA9" bold="true" italics="true" pageId="0" pageNumber="15">Holotype</emphasis>
<taxonomicName id="A9E36282A9F9E3EABB0320D01A6BBC23" authorityName="Browne" authorityYear="1970" class="Insecta" family="Curculionidae" genus="Taenioglyptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taenioglyptes artestriatus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="artestriatus">
<emphasis id="CA9514C9811BF1D51BB2C406B5240730" italics="true" pageId="0" pageNumber="15">Taenioglyptes artestriatus</emphasis>
</taxonomicName>
Browne, 1970; Central Region, Wakiso, Zika; 
<geoCoordinate id="F3DCE1100AC6F57F2AEA804DE6BFA88E" degrees="0.12" direction="north" orientation="latitude" precision="555" value="0.12">0.12°N</geoCoordinate>
, 
<geoCoordinate id="D28D06B2CCC6A329189EF007863B16A6" degrees="32.52" direction="east" orientation="longitude" precision="555" value="32.52">32.52°E</geoCoordinate>
; 21 May 1961; K. W. Brown leg.; &quot; 
<emphasis id="4D2336FC533FB7DD14FE28EBE8F6F718" bold="true" italics="true" pageId="0" pageNumber="15">Holotype</emphasis>
////Uganda//Zika//in light trap//K. W. Brown //29/5/61 //B.1715(g) ////1666 //// 
<taxonomicName id="1E9865F0BB3974E8DAF6DA876F271639" authorityName="Browne" authorityYear="1970" class="Insecta" family="Curculionidae" genus="Taenioglyptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taenioglyptes artestriatus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="artestriatus">Taenioglyptes artestriatus</taxonomicName>
// Browne ♀ // 
<emphasis id="6D91411F09C37AEC22EE8103AA118786" bold="true" italics="true" pageId="0" pageNumber="15">Holotype</emphasis>
////Brit. Mus. 1973-70//// NHMUK 010806013&quot;; UFFE:26192; (NHMUK).
</paragraph>
</subSubSection>
<subSubSection id="D2B397C0D85825D15F3F2090FEA0B865" pageId="0" pageNumber="15" type="materials_examined">
<paragraph id="A97AB6C0510D36C1D039E46A23192130" pageId="0" pageNumber="15">Other material examined.</paragraph>
<paragraph id="5E8B83693FFEB1D5FDC312076463CEA4" pageId="0" pageNumber="15">
China • 1 ♀; Fujian, Quanzhou, 
<normalizedToken id="57BC448F83BDDB503CA1740875264062" originalValue="Nan’an">Nan'an</normalizedToken>
, Pushan village; 
<geoCoordinate id="97EBC01706CDDD2BFA83C7DE6AD3ADC6" degrees="25.1189" direction="north" orientation="latitude" precision="5" value="25.1189">25.1189°N</geoCoordinate>
, 
<geoCoordinate id="FACD2F65E33099B7E99D1C02D283A443" degrees="118.4276" direction="east" orientation="longitude" precision="5" value="118.4276">118.4276°E</geoCoordinate>
; May 2017; You Li leg.; ex. 
<taxonomicName id="9E5AB61549B5F6D4B777BEE99321B80A" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mangifera indica" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="C22A84F475680AC7AD93D8A1F5CC27E8" italics="true" pageId="0" pageNumber="15">Mangifera indica</emphasis>
</taxonomicName>
; UFFE:33207; (UFFE) • 1 ♂; same collection data; UFFE:33208; (UFFE) • 9 ♀♀, 8 ♂♂; same collection data; UFFE:33206; (NHMUK, 1♀, 1♂; FSCA, 1♀; MZB, 1♀, 1♂; NIAES, 1♀, 1♂; NMNS, 1♀, 1♂; IOZ 1♀ IOZ(E)2057932, 1♂ IOZ(E)2057933; RIFID, 1♀, 1♂; USNM, 1♀, 1♂; ZIN, 1♀, 1♂) • 1 ♂; same collection data; dissected; UFFE:34966 (UFFE); • 1 ♀; Hunan, Changsha, Yuelushan; Aug. 2019; You Li leg.; ex. 
<taxonomicName id="F5632F78CB0345D8AFCE250E6070E2DB" class="Magnoliopsida" family="Anacardiaceae" genus="Choerospondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Choerospondias axillaris" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="08793D2FF78D4DAA6A5166E9C1E7642D" italics="true" pageId="0" pageNumber="15">Choerospondias axillaris</emphasis>
</taxonomicName>
; DNA: 28S:MT431546; UFFE:33551 • 57; Hunan, Changsha, Yuelushan; 
<geoCoordinate id="94DA30CF6D4AF519266108A3EBC225A0" degrees="28.1897" direction="north" orientation="latitude" precision="5" value="28.1897">28.1897°N</geoCoordinate>
, 
<geoCoordinate id="33CF59FA07C6E8E72DD6EEBF0E64DF0D" degrees="112.9329" direction="east" orientation="longitude" precision="5" value="112.9329">112.9329°E</geoCoordinate>
; Aug. 2019; You Li leg.; ex. 
<taxonomicName id="AC42FB92981E023E62A12E8DE0BFE822" class="Magnoliopsida" family="Anacardiaceae" genus="Choerospondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Choerospondias axillaris" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="FD7211F0B2AFD1E492EDC522F843F7FE" italics="true" pageId="0" pageNumber="15">Choerospondias axillaris</emphasis>
</taxonomicName>
; UFFE:33529; (UFFE) • 1; Yunnan, Mengla XTBG; 
<taxonomicName id="53709806D791CDBE3CF6F93EDCA275E8" genus="C." lsidName="C." pageId="0" pageNumber="15" rank="genus">C.</taxonomicName>
Bateman leg.; DNA: 28S:MG051112, COI:MG051163; UFFE:14119; (UFFE) • 1; Yunnan, Menglun XTBG; 20 Jul. 2017; Craig Bateman leg.; DNA: COI:MG051162; UFFE:25419; (UFFE).
</paragraph>
<paragraph id="6DB5BA3DEE3C0F32BD0E30DA4E093DA8" pageId="0" pageNumber="15">Guadeloupe • 1 ♂; Basse Terre, Point a Lezard; 20 May 2012; R. Turnbow leg.; UFFE:12501; (FSCA).</paragraph>
<paragraph id="CEDBD31AD6539EB45B3E3E6FAE7AF210" pageId="0" pageNumber="15">Taiwan • 1; Kaohsiung, Liouguei; Tom Harrington, and Caroline Wuerst leg.; Lindgren trap with ethanol; DNA: 28S:MG051118, COI:MG051172; UFFE:15127; (UFFE).</paragraph>
<paragraph id="18CA15B525B0DD19709D425F370E3184" pageId="0" pageNumber="15">
United States • 1; Florida, Homestead; 29 Jun. 2015; Thomas H. Atkinson leg.; ex. 
<taxonomicName id="39F11E3B9F16A1ED65FD36591EFF5E72" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mangifera indica" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="B4312D6932BDB88BC552650E4424530A" italics="true" pageId="0" pageNumber="15">Mangifera indica</emphasis>
</taxonomicName>
; from vial 9444; DNA: 28S:MG051119, COI:MG051174; UFFE:25414; (UFFE) • 18; Florida, Miami Dade, Homestead, TREC; 23 Jul. 2018; Andrew J. Johnson leg.; ex. 
<taxonomicName id="E9A3093BFE3B0F6FE90785116EF2096F" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mangifera indica" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="C4DCB20760877517AE8C13F3C38A6C72" italics="true" pageId="0" pageNumber="15">Mangifera indica</emphasis>
</taxonomicName>
; UFFE:29770; (UFFE). • 1; Florida, Dade County, Homestead; 2012-Apr-05; L. Bradshaw leg.; ex. 
<taxonomicName id="A3AD3B6D7F991971A807EA36F90741BA" class="Magnoliopsida" family="Euphorbiaceae" genus="Codiaeum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Codiaeum variegatum" order="Malpighiales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="variegatum">
<emphasis id="FCB2EB6BA7A25C566A0F108F24299513" italics="true" pageId="0" pageNumber="15">Codiaeum variegatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D36EAD9BAC16EE20A66E859E89188A10" pageId="0" pageNumber="15">Vietnam • 1 ♀; TamDao; Michelle Jusino, and James Skelton leg.; quercivorol and ethanol trap; DNA: 28S:MG051120, COI:MG051175; UFFE:25936; (UFFE).</paragraph>
</subSubSection>
<subSubSection id="C983F321C4CE14641720FA7E5D897818" pageId="0" pageNumber="15" type="diagnosis">
<paragraph id="1AF625D6CAEA66BB6789E78E621C8D2C" pageId="0" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="D169A3D0B621BAC424879DE3F9C15763" pageId="0" pageNumber="15">
This species is distinguished from other similar 
<taxonomicName id="D4239F3D03B28C2F2F5B39B6271779E6" authorityName="Erichson" authorityYear="1836" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B99DF281EC4970C0D385DB7292C2BC0A" italics="true" pageId="0" pageNumber="15">Cryphalus</emphasis>
</taxonomicName>
by the frons with a finely aciculate texture, the pronotal disc which is long, and has coarse hair-like setae, the elytral striae which are barely impressed but apparent by the rows without ground vestiture, and by the shape of ground vestiture which have tapered tips.
</paragraph>
<paragraph id="B2B06C2E1C7BBBECAB5E74F656B910B0" pageId="0" pageNumber="15">
See notes for diagnosing the very similar 
<taxonomicName id="3610B2AD79DE71F3363D7350ACD278F1" authorityName="Johnson" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus paramangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="paramangiferae">
<emphasis id="DD5DB914B548707138CD18480DCA04C8" italics="true" pageId="0" pageNumber="15">Cryphalus paramangiferae</emphasis>
</taxonomicName>
, described below, under the diagnosis for that species.
</paragraph>
<paragraph id="D4126592041527E140BBD1995EFDD5D5" pageId="0" pageNumber="15">
<emphasis id="C24D18D36D3952CF5A64C3F75180AC5D" bold="true" pageId="0" pageNumber="15">Female.</emphasis>
Length 1.6-2.2 mm. Proportions 2.2 
<normalizedToken id="ABB3F3A384E353BB75EA1A2293B1D5E9" originalValue="×">x</normalizedToken>
as long as wide. Frons with weak converging aciculations. Antennal funiculus with four or five funicular segments. Antennal club with three evenly procurved sutures mostly marked by coarse setae. Pronotal profile widest in line with summit. Pronotal margin armed with four to six widely spaces serrations, the median pair larger. Pronotal declivity with more than 60 asperities. Pronotal disc approximately one third of the length, gently sloped. Pronotal vestiture entirely hair-like or dagger-like, very few setae bifurcating in baso-lateral area Suture between pronotum and elytra weakly sinuate. Scutellum shaped as a rounded triangle, almost semi-circular, with sparse, pale, hair-like setae. Elytra 1.6 
<normalizedToken id="746FAE3A60B0A0BB33081DA1430256AB" originalValue="×">x</normalizedToken>
as long as pronotum, usually a similar colour to pronotum, translucent yellow-brown, sometimes to dark brown, broadly rounded with no clear transition to the declivity. Striae weakly visible as rows of punctures and hair-like setae with almost no ground vestiture, barely impressed Interstrial bristles erect, curving posteriorly, of approximately even diameter. Interstrial ground vestiture near triangular, dagger-like, tapering to a fine point, longer on declivity, ground vestiture on basal third are usually a light brown/cold colour. Mesocoxae moderately separated, much more than metacoxae. Ventrites with mostly hair-like setae. Last abdominal ventrite with margin of rounded tubercles Proventriculus sutural teeth weakly sclerotised, rounded, irregular, one indistinct row each side of suture. Apical teeth in multiple rows, extending almost the width of a segment. Closing teeth long, barely branched. Masticatory brush short, less than length or apical plate.
</paragraph>
<paragraph id="B5736E632B0CDD63531458B94301EC4B" pageId="0" pageNumber="15">
<emphasis id="B684D03520F6A0F85E26BEF2F217D014" bold="true" pageId="0" pageNumber="15">Male.</emphasis>
Similar to female except: Length 1.5-2.2 mm. Frons identical to females, with granulate texture on median of upper level. Pronotal profile slightly more triangular, widest nearer base. Pronotal vestiture hair like, with a few bifurcating setae on baso-lateral areas, slightly more than on female. Protibiae and protarsi with several larger, coarse curved setae on the proximal edge near the apex. Last abdominal ventrite weakly emarginated. Aedeagus long, penis body sclerotised, tapered to a point at apex, ejaculatory duct with small, evenly sized spinulae, end plates sclerotised. Penis apodemes 2.5 
<normalizedToken id="975A11B27180CAB2AD5A6D4C65DC31E6" originalValue="×">x</normalizedToken>
as long as penis body. Tegmen broad, with very small, almost obsolete paired apodemes.
</paragraph>
</subSubSection>
<subSubSection id="A92322AFD3EC51B479142E6C2C346025" pageId="0" pageNumber="15" type="distribution">
<paragraph id="1853CAFF3980179EA521D41F9FBA0A88" pageId="0" pageNumber="15">Distribution.</paragraph>
<paragraph id="341B66897FAEA662D9DC711CFC546D66" pageId="0" pageNumber="15">China (Fujian, Guangdong, Hunan, Yunnan); Taiwan; Thailand; Malaysia; Indonesia; India; Nepal; United States (Florida, Hawaii); Mexico; Puerto Rico; Cuba; Kenya, Uganda; American Samoa, Australia.</paragraph>
</subSubSection>
<subSubSection id="7A445A163334A6C4545F4BA919F8159D" pageId="0" pageNumber="15" type="suggested vernacular name">
<paragraph id="D1B3C3C35991E306C84F7AA7C24434B6" pageId="0" pageNumber="15">Suggested vernacular name.</paragraph>
<paragraph id="951006DBEAD3CDBCC10FC39AC3159BE7" pageId="0" pageNumber="15">
Chinese: 
<normalizedToken id="DB958E74AAA74FB6FF332E191C13AFE7" originalValue="芒果梢小蠹">芒果梢小蠹</normalizedToken>
[Mango twig bark beetle]; English: Mango bark beetle.
</paragraph>
</subSubSection>
<subSubSection id="26C31D0996E6A8D932AB3BDF248F9C0B" pageId="0" pageNumber="15" type="recorded plant hosts">
<paragraph id="2D6A14A8E90CC8F0D3629DE2B7EFC49D" pageId="0" pageNumber="15">Recorded plant hosts.</paragraph>
<paragraph id="855CA805AB982EEB57E7CB63B7E5DE2F" pageId="0" pageNumber="15">
<taxonomicName id="48E0A5DEE53BF8C615B24628DE4DE6EF" authorityName="Erichson" authorityYear="1836" class="Magnoliopsida" family="Anacardiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
: 
<taxonomicName id="FBE91EB457AFF2F3B3BB343D4D9CF387" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mangifera indica" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="9ED46D66B45E12B12D0EC85C6EAA05C9" italics="true" pageId="0" pageNumber="15">Mangifera indica</emphasis>
</taxonomicName>
L., 
<taxonomicName id="100ECC4D64E9CCAC9E630DA90AA823A4" lsidName="M. odorata" pageId="0" pageNumber="15" rank="species" species="odorata">
<emphasis id="4324C6A32AA8F904D0F1707D4F9D4888" italics="true" pageId="0" pageNumber="15">M. odorata</emphasis>
</taxonomicName>
Griff. (
<bibRefCitation id="A73AD17EBD0AB2EB2C5D956AC9A99D4E" author="Kalshoven, LGE" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="15" pagination="157 - 180" refId="B25" refString="Kalshoven, LGE, 1958. Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. First part. Tijdschrift voor Entomologie 101: 157 - 180" title="Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. First part." volume="101" year="1958">Kalshoven 1958</bibRefCitation>
), 
<taxonomicName id="8F243E01C467FA551408A4BC132359C5" class="Magnoliopsida" family="Anacardiaceae" genus="Choerospondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Choerospondias axillaris" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="C72290BF430898221A90CC4016E6B9FE" italics="true" pageId="0" pageNumber="15">Choerospondias axillaris</emphasis>
</taxonomicName>
(Roxb.) B.L.Burtt &amp; A.W.Hill; 
<taxonomicName id="3DDF6E9926C63764295FE918D1C21C40" authorityName="B.L.Burtt &amp; A.W.Hill" authorityYear="1937" class="Magnoliopsida" family="Euphorbiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malpighiales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
: 
<taxonomicName id="ED4D0D30B00DD832079CD4497AC4716B" class="Magnoliopsida" family="Euphorbiaceae" genus="Codiaeum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Codiaeum variegatum" order="Malpighiales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="variegatum">
<emphasis id="0C9329023D332813F52DE6BF70E40E2E" italics="true" pageId="0" pageNumber="15">Codiaeum variegatum</emphasis>
</taxonomicName>
(L.) A.Juss. [not confirmed reproductive host].
</paragraph>
</subSubSection>
<subSubSection id="D99BD890D529B59BE5E893E9947B60D8" pageId="0" pageNumber="15" type="remarks">
<paragraph id="2E89A1BFEB7C940B5DE4DACC5CBE4276" pageId="0" pageNumber="15">Remarks.</paragraph>
<paragraph id="A15EA018CF99239664AA212D1B361EF7" pageId="0" pageNumber="15">
<taxonomicName id="64B4D530E7007E935528A11E8E8D7D05" class="Magnoliopsida" family="Anacardiaceae" genus="Choerospondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Choerospondias axillaris" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="axillaris">
<emphasis id="0FA166614BFC4A29E7A2153ABBE19FA0" italics="true" pageId="0" pageNumber="15">Choerospondias axillaris</emphasis>
</taxonomicName>
is a specialty crop in southern China, and it represents a new reproductive host record. A record from 
<taxonomicName id="55C728BEFC4FC41FEDC69515ED0FE761" class="Magnoliopsida" family="Euphorbiaceae" genus="Codiaeum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Codiaeum variegatum" order="Malpighiales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="variegatum">
<emphasis id="BB9AEDE1F552299E2146AE5B07A0A6F3" italics="true" pageId="0" pageNumber="15">Codiaeum variegatum</emphasis>
</taxonomicName>
from Florida suggests that the beetle might have a broader host range than previously thought, though 
<taxonomicName id="BF41A47FEFD1EA8B99808ACC77806585" class="Magnoliopsida" family="Euphorbiaceae" genus="Codiaeum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Codiaeum" order="Malpighiales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="1BF614C1AFC4291DE23BD48C68281EEA" italics="true" pageId="0" pageNumber="15">Codiaeum</emphasis>
</taxonomicName>
was not confirmed as a reproductive host. 
<bibRefCitation id="78BCAF66D5839CEBEEF510B340492268" author="Kalshoven, LGE" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="15" pagination="157 - 180" refId="B25" refString="Kalshoven, LGE, 1958. Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. First part. Tijdschrift voor Entomologie 101: 157 - 180" title="Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. First part." volume="101" year="1958">Kalshoven (1958)</bibRefCitation>
also lists 
<taxonomicName id="664EBDC9D132F251D2D4E7634552C518" class="Magnoliopsida" family="Rhamnaceae" genus="Zizyphus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Zizyphus oenopila" order="Rosales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="oenopila">
<emphasis id="85D452C24AE020AE592D8998C6EC8F91" italics="true" pageId="0" pageNumber="15">Zizyphus oenopila</emphasis>
</taxonomicName>
(L.) Mill. (
<taxonomicName id="B0863CEA4485FF31C8B007F0DD2D94C1" class="Magnoliopsida" family="Rhamnaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Rhamnaceae</taxonomicName>
), 
<taxonomicName id="CE765EEDD898DEE382DB09096CFE6A5C" class="Magnoliopsida" family="Malvaceae" genus="Theobroma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Theobroma cacao" order="Malvales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="cacao">
<emphasis id="B61733DBEF41896E4D1FE1534F4F5732" italics="true" pageId="0" pageNumber="15">Theobroma cacao</emphasis>
</taxonomicName>
L. (
<taxonomicName id="F8527AAC6848141BCA969B41A785B588" class="Magnoliopsida" family="Malvaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malvales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
), 
<taxonomicName id="F26724105AD0E646975C777A5F6F3F93" authorityName="Sahni &amp; Bennet" authorityYear="1974" baseAuthorityName="Osbeck" class="Magnoliopsida" family="Meliaceae" genus="Lansium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lansium parasiticum" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="parasiticum">
<emphasis id="2376ABBB06A2643DE3EFE3D800134183" italics="true" pageId="0" pageNumber="15">Lansium parasiticum</emphasis>
</taxonomicName>
(Osbeck) Sahni &amp; Bennet (
<taxonomicName id="F313E7BDDE3A428178D13A95DDA4F6CD" class="Magnoliopsida" family="Meliaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
) and 
<taxonomicName id="E7920D87767C06782342067E9034F101" class="Magnoliopsida" family="Burseraceae" genus="Canarium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Canarium commune" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="species" species="commune">
<emphasis id="D6C06EF501AC347396D037B786445B4B" italics="true" pageId="0" pageNumber="15">Canarium commune</emphasis>
</taxonomicName>
L. (
<taxonomicName id="50E2750FC7A6A5E2EDBC0C8031310D4F" class="Magnoliopsida" family="Burseraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
), but the associated vouchers were not examined for accuracy. 
<bibRefCitation id="82CDCDEEA975B23ABEFB3295A067C3BD" DOI="https://doi.org/10.1002/mmnd.19900370408" author="Beaver, RA" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="15" pagination="279 - 284" refId="B2" refString="Beaver, RA, 1990. New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae). Deutsche Entomologische Zeitschrift 37: 279 - 284, DOI: https://doi.org/10.1002/mmnd.19900370408" title="New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae)." url="https://doi.org/10.1002/mmnd.19900370408" volume="37" year="1990">Beaver (1990)</bibRefCitation>
also recorded this species from an unknown angiosperm not 
<taxonomicName id="52FDA6B9017087CB59666584A968489F" class="Magnoliopsida" family="Anacardiaceae" genus="Mangifera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mangifera" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="57383C2B88CAB1C98A2F44BFBFF557D1" italics="true" pageId="0" pageNumber="15">Mangifera</emphasis>
</taxonomicName>
. No host outside of 
<taxonomicName id="4FCB53098CA1F6E471250F3C5AB223D2" class="Magnoliopsida" family="Anacardiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="15" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
has been verified as a reproductive host.
</paragraph>
<paragraph id="A15E18730F3D90A302AE0089CD9DAED3" pageId="0" pageNumber="15">
The holotype of 
<taxonomicName id="D631D6189B4BE7A0CC16D9E74F1B7656" authorityName="Browne" authorityYear="1970" class="Insecta" family="Curculionidae" genus="Taenioglyptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taenioglyptes artestriatus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="artestriatus">
<emphasis id="C5EE6E2FC3F7AEA563FD4AD6314EBB81" italics="true" pageId="0" pageNumber="15">Taenioglyptes artestriatus</emphasis>
</taxonomicName>
Browne, 1970, while not from the regions of study, is recognised as a synonym. All diagnostic characters are visible. The antennae have four funicular segments, which previously distinguished the genera 
<taxonomicName id="D5FD023AB5308943F35D76A2331C016A" authorityName="Erichson" authorityYear="1836" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="C6795B418E08C2830614D60EBCACFEC4" italics="true" pageId="0" pageNumber="15">Cryphalus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3BADD4C6D1A0BEF9B2F73EC398F7557D" authorityName="Hopkins" authorityYear="1915" class="Insecta" family="Curculionidae" genus="Hypocryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocryphalus" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B18CB3E15CD496B0B635B5B10A66A4B5" italics="true" pageId="0" pageNumber="15">Hypocryphalus</emphasis>
</taxonomicName>
, probably leading to this error. Several examples from China had similar antennae, and one specimen from Guadeloupe.
</paragraph>
<caption id="85CDCB39F74CA851BC1E839A2F013227" doi="10.3897/zookeys.995.55981.figure12" httpUri="https://binary.pensoft.net/fig/476785" pageId="0" pageNumber="15" start="Figure 12" startId="F12">
<paragraph id="053FE4B5342A29948FCB851EDC2B58EC" pageId="0" pageNumber="15">
<emphasis id="6AF609FA8A50933B628FE1393AA336A1" bold="true" pageId="0" pageNumber="15">Figure 12.</emphasis>
<taxonomicName id="47E0417A0B8C94FEAC50C8845F2A349F" authorityName="Stebbing" authorityYear="1914" class="Insecta" family="Curculionidae" genus="Cryphalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryphalus mangiferae" order="Coleoptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="mangiferae">
<emphasis id="F2A5E1F956FA049B515FFAE1254B7E81" italics="true" pageId="0" pageNumber="15">Cryphalus mangiferae</emphasis>
</taxonomicName>
<emphasis id="D34F2EF4FB4B144CFF43719A4F685A1D" bold="true" pageId="0" pageNumber="15">A, C, E, G</emphasis>
female, UFFE:33207 
<emphasis id="773B9C4681FE28D56955B09E7A8C9B62" bold="true" pageId="0" pageNumber="15">B, D, F, H, I</emphasis>
male, UFFE:33208.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>