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165 lines
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<document ID-DOI="10.37520/fi.2022.022" ID-ISSN="2533-4069" ID-Zenodo-Dep="7522593" checkinTime="1673381619156" checkinUser="felipe" docAuthor="Přikryl, Tomáš, Kovalchuk, Oleksandr, Carnevale, Giorgio & Barkaszi, Zoltán" docDate="2022" docId="C0070919392AFFB314119AF0FE05E50C" docLanguage="en" docName="FossilImprint.78.2.513-518.pdf" docOrigin="Fossil Imprint 78 (2)" docSource="http://dx.doi.org/10.37520/fi.2022.022" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.3:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="3" docTitle="Archaeotetraodon winterbottomi TYLER ET BANNIKOV 1994" docType="treatment" docVersion="3" lastPageNumber="516" masterDocId="3C3E7161392BFFB017289C19FFC5E208" masterDocTitle="New Material Of The Puffer Fish Archaeotetraodon Winterbottomi Tyler Et Bannikov, 1994 (Tetraodontidae) From The Oligocene Of The Eastern Paratethys" masterLastPageNumber="518" masterPageNumber="513" pageNumber="514" updateTime="1673890994017" updateUser="juliana" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Material Of The Puffer Fish Archaeotetraodon Winterbottomi Tyler Et Bannikov, 1994 (Tetraodontidae) From The Oligocene Of The Eastern Paratethys</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Přikryl, Tomáš</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kovalchuk, Oleksandr</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Carnevale, Giorgio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barkaszi, Zoltán</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Imprint</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>78</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>513</mods:start>
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<mods:end>518</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.37520/fi.2022.022</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.37520/fi.2022.022</mods:identifier>
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<mods:identifier type="ISSN">2533-4069</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7522593</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:C0070919392AFFB314119AF0FE05E50C" httpUri="http://treatment.plazi.org/id/C0070919392AFFB314119AF0FE05E50C" lastPageId="3" lastPageNumber="516" pageId="1" pageNumber="514">
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<subSubSection box="[825,1454,1769,1793]" pageId="1" pageNumber="514" type="nomenclature">
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<paragraph blockId="1.[825,1454,1769,1793]" box="[825,1454,1769,1793]" pageId="1" pageNumber="514">
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<heading box="[825,1454,1769,1793]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="514" reason="2">
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<taxonomicName authority="TYLER et BANNIKOV, 1994" authorityName="TYLER ET BANNIKOV" authorityYear="1994" box="[825,1454,1769,1793]" family="Tetraodontidae" genus="Archaeotetraodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="514" phylum="Chordata" rank="species" species="winterbottomi">
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<emphasis bold="true" box="[825,1454,1769,1793]" pageId="1" pageNumber="514">
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<emphasis bold="true" box="[825,1175,1769,1793]" italics="true" pageId="1" pageNumber="514">Archaeotetraodon winterbottomi</emphasis>
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<bibRefCitation author="Tyler, J. C. & Bannikov, A. F." box="[1181,1250,1769,1793]" pageId="1" pageNumber="514" pagination="97 - 108" refId="ref4638" refString="Tyler, J. C., Bannikov, A. F. (1994): A new genus of fossil pufferfish (Tetraodontidae: Tetraodontiformes) based on a new species from the Oligocene of Russia and a referred species from the Miocene of Ukraine. - Proceedings of the Biological Society of Washington, 107: 97 - 108." type="journal article" year="1994">TYLER</bibRefCitation>
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et BANNIKOV, 1994
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1069,1210,1802,1826]" pageId="1" pageNumber="514" type="description">
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<paragraph blockId="1.[1069,1210,1802,1826]" box="[1069,1210,1802,1826]" pageId="1" pageNumber="514">
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<figureCitation box="[1069,1210,1802,1826]" captionStart-0="Text-fig" captionStart-1="Text-fig" captionStartId-0="2.[147,228,1310,1331]" captionStartId-1="4.[129,210,1156,1177]" captionTargetBox-0="[147,1444,151,1279]" captionTargetBox-1="[129,764,154,1141]" captionTargetId-0="figure-319@2.[-102,1638,-179,981]" captionTargetId-1="figure-574@4.[129,764,154,1141]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="Text-fig. 2. Archaeotetraodon winterbottomi. Specimen NMNHU-P PI 1995. Sheshory locality, Ukraine (Rybnytsia Member, lowermost part of the Menilite Series, Oligocene). a: photo of the specimen; b: interpretative drawing of the (a)." captionText-1="Text-fig. 3. Archaeotetraodon winterbottomi. Specimen NMN- HU-P PI 1995. Details of dermal bifid spinules." figureDoi-0="http://doi.org/10.5281/zenodo.7522597" figureDoi-1="http://doi.org/10.5281/zenodo.7522599" httpUri-0="https://zenodo.org/record/7522597/files/figure.png" httpUri-1="https://zenodo.org/record/7522599/files/figure.png" pageId="1" pageNumber="514">Text-figs 2–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="514" type="reference_group">
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<paragraph blockId="1.[820,1459,1848,2019]" pageId="1" pageNumber="514">
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1994
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<taxonomicName authority="Přikryl & Kovalchuk & Carnevale & Barkaszi, 2022" authorityName="Přikryl & Kovalchuk & Carnevale & Barkaszi" authorityYear="2022" box="[896,1215,1848,1869]" family="Tetraodontidae" genus="Archaeotetraodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="514" phylum="Chordata" rank="species" species="winterbottomi" status="new species">
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<emphasis box="[896,1215,1848,1869]" italics="true" pageId="1" pageNumber="514">Archaeotetraodon winterbottomi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1226,1349,1848,1870]" pageId="1" pageNumber="514" rank="species">new species</taxonomicNameLabel>
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; Tyler and Bannikov, p. 103, figs 1–4.
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</paragraph>
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<paragraph blockId="1.[820,1459,1848,2019]" pageId="1" pageNumber="514">
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2010
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<taxonomicName authority="TYLER et BANNIKOV, 1994" authorityName="TYLER ET BANNIKOV" authorityYear="1994" box="[896,1455,1908,1929]" family="Tetraodontidae" genus="Archaeotetraodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="514" phylum="Chordata" rank="species" species="winterbottomi">
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<emphasis box="[896,1209,1908,1929]" italics="true" pageId="1" pageNumber="514">Archaeotetraodon winterbottomi</emphasis>
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<bibRefCitation author="Tyler, J. C. & Bannikov, A. F." box="[1212,1455,1908,1929]" pageId="1" pageNumber="514" pagination="97 - 108" refId="ref4638" refString="Tyler, J. C., Bannikov, A. F. (1994): A new genus of fossil pufferfish (Tetraodontidae: Tetraodontiformes) based on a new species from the Oligocene of Russia and a referred species from the Miocene of Ukraine. - Proceedings of the Biological Society of Washington, 107: 97 - 108." type="journal article" year="1994">TYLER et BANNIKOV, 1994</bibRefCitation>
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</taxonomicName>
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; Bannikov, p. 170, pl. XXIX, fig. 3.
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</paragraph>
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<paragraph blockId="1.[820,1459,1848,2019]" pageId="1" pageNumber="514">
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2010
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<taxonomicName authority="TYLER and BANNIKOV, 1994" authorityName="TYLER and BANNIKOV" authorityYear="1994" family="Tetraodontidae" genus="Archaeotetraodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="1" pageNumber="514" phylum="Chordata" rank="species" species="winterbottomi">
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<emphasis box="[896,1220,1968,1989]" italics="true" pageId="1" pageNumber="514">Archaeotetraodon winterbottomi</emphasis>
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<bibRefCitation author="Tyler, J. C. & Bannikov, A. F." pageId="1" pageNumber="514" pagination="97 - 108" refId="ref4638" refString="Tyler, J. C., Bannikov, A. F. (1994): A new genus of fossil pufferfish (Tetraodontidae: Tetraodontiformes) based on a new species from the Oligocene of Russia and a referred species from the Miocene of Ukraine. - Proceedings of the Biological Society of Washington, 107: 97 - 108." type="journal article" year="1994">TYLER and BANNIKOV, 1994</bibRefCitation>
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</taxonomicName>
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; Carnevale and Tyler, p. 295, fig. 13.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7522595" ID-Zenodo-Dep="7522595" httpUri="https://zenodo.org/record/7522595/files/figure.png" pageId="1" pageNumber="514" startId="1.[106,187,2019,2040]" targetBox="[106,741,1089,2004]" targetPageId="1">
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<paragraph blockId="1.[106,741,2019,2099]" pageId="1" pageNumber="514">Text-fig. 1. Iosif Vasilievich Kiselev in the later years of life. The figure from the archive of the Institute of Geography, National Academy of Sciences of Ukraine.</paragraph>
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</caption>
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<subSubSection lastPageId="2" lastPageNumber="515" pageId="1" pageNumber="514" type="materials_examined">
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<paragraph blockId="1.[820,1459,2043,2100]" lastBlockId="2.[129,767,1431,1488]" lastPageId="2" lastPageNumber="515" pageId="1" pageNumber="514">
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M a t e r i a l. The specimen NMNHU-P PI 1995 is deposited in the Department of Palaeontology of the National Museum of Natural History, National Academy of Sciences of
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<collectingCountry box="[257,343,1464,1488]" name="Ukraine" pageId="2" pageNumber="515">Ukraine</collectingCountry>
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in
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<collectingRegion box="[377,432,1464,1488]" country="Ukraine" name="Kyiv" pageId="2" pageNumber="515">Kyiv</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7522597" ID-Zenodo-Dep="7522597" httpUri="https://zenodo.org/record/7522597/files/figure.png" pageId="2" pageNumber="515" startId="2.[147,228,1310,1331]" targetBox="[147,1444,151,1279]" targetPageId="2">
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<paragraph blockId="2.[147,1462,1310,1360]" pageId="2" pageNumber="515">
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Text-fig. 2.
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<taxonomicName authorityName="TYLER and BANNIKOV" authorityYear="1994" box="[273,601,1310,1331]" family="Tetraodontidae" genus="Archaeotetraodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tetraodontiformes" pageId="2" pageNumber="515" phylum="Chordata" rank="species" species="winterbottomi">
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<emphasis bold="true" box="[273,601,1310,1331]" italics="true" pageId="2" pageNumber="515">Archaeotetraodon winterbottomi</emphasis>
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</taxonomicName>
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. Specimen NMNHU-P PI 1995. Sheshory locality, Ukraine (Rybnytsia Member, lowermost part of the Menilite Series, Oligocene). a: photo of the specimen; b: interpretative drawing of the (a).
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</paragraph>
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</caption>
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<subSubSection lastPageId="3" lastPageNumber="516" pageId="2" pageNumber="515" type="description">
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<paragraph blockId="2.[129,768,1512,2092]" pageId="2" pageNumber="515">
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D e s c r i p t i o n. The specimen is preserved in left lateral view (
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<figureCitation box="[268,382,1544,1568]" captionStart="Text-fig" captionStartId="2.[147,228,1310,1331]" captionTargetBox="[147,1444,151,1279]" captionTargetId="figure-319@2.[-102,1638,-179,981]" captionTargetPageId="2" captionText="Text-fig. 2. Archaeotetraodon winterbottomi. Specimen NMNHU-P PI 1995. Sheshory locality, Ukraine (Rybnytsia Member, lowermost part of the Menilite Series, Oligocene). a: photo of the specimen; b: interpretative drawing of the (a)." figureDoi="http://doi.org/10.5281/zenodo.7522597" httpUri="https://zenodo.org/record/7522597/files/figure.png" pageId="2" pageNumber="515">Text-fig. 2</figureCitation>
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), with the neurocranium exposed in dorso-lateral view.
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</paragraph>
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<paragraph blockId="2.[129,768,1512,2092]" pageId="2" pageNumber="515">The skull roof is almost hourglass-shaped. Although the anterior portion of the neurocranium is not completely clear, it is possible to recognize the crescent-shaped mesethmoid posteriorly articulated with the poorly preserved lateral ethmoids.The frontals have concave lateral margins, widened posteriorly and contact the sphenotics posterolaterally. The outline of the pterotic is partially recognizable. The ventral part of the neurocranium is extensively fragmented and partially covered by overlying bones.</paragraph>
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<paragraph blockId="2.[129,768,1512,2092]" lastBlockId="2.[843,1482,1431,2092]" pageId="2" pageNumber="515">The beak-like jaws are massive. The premaxillae are more or less triangular. Remains of the left maxilla suggest the presence of a relatively large, dorsally oriented process forming a right angle with the main shaft of the bone. The left mandible is triangular in outline and anteroposteriorly elongate; the bone in its postero-ventral corner is articulated with the quadrate. There is a distinctive notch with a rounded profile along the posterior margin of the mandible. Bony fragments located just below the ethmoid region of the skull probably represent what remains of the palatine.</paragraph>
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<paragraph blockId="2.[843,1482,1431,2092]" pageId="2" pageNumber="515">The ceratohyal is rod-like anteriorly and broadly expanded posteriorly. Along the ventral margin of its expanded portion, it is possible to observe five partially complete branchiostegal rays, possibly representing the second to sixth rays of the series. The first ray was probably enlarged, as in other tetraodontids, and it is poorly preserved, displaced behind the hyoid arch. Remains of three other contralateral branchiostegal rays are preserved below the neurocranium. The left hyomandibula is displaced from its original position; it has a straight vertical shaft and a gently concave anterior margin.</paragraph>
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<paragraph blockId="2.[843,1482,1431,2092]" pageId="2" pageNumber="515">Several isolated fragments of the branchial skeleton (pharyngobranchials and, possibly, a portion of an epibranchial) are disarticulated and dispersed above the ceratohyal.</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">The cleithrum is sigmoid in outline and bears a posterodorsally-directed spine and a relatively large posterior lamina, partially covering the remains of a single pectoralfin radial, while a second one is exposed just below it. The supracleithra are extensively fragmented. There are two welldeveloped postcleithra, of which the ventral one is thin, riblike and slightly bent. Its length slightly exceeds 20 % of SL, while the dorsal one is slightly more massive than the ventral one and its length is about half of the ventral one. Remains of the scapula, coracoid and pectoral fins are not recognizable.</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">The vertebral column consists of 17 vertebrae (eight abdominal plus nine caudal). The first three abdominal vertebrae bear shortened neural spines. Neural spines of the three successive vertebrae are much longer and posteriorly inclined. The neural spines of the seventh to eleventh vertebrae are not completely recognizable, while the twelfth to penultimate vertebrae have wide and relatively short neural spines. The first centrum seems to be slightly thinner compared to the following ones, but that part of the fossil is not completely preserved, suggesting that this condition may represent a preservation artifact. The subsequent centra are rectangular in outline, longer than high. The terminal section of the vertebral column comprises anteroposteriorly compact centra. Ribs and intermuscular bones are not present.</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">The caudal skeleton is characterized by an autogenous parhypural and haemal spine of the second preural vertebra fused to the centrum. The hypural plate and the dorsal portion of the caudal skeleton is not completely clear. The truncate caudal fin is only partially preserved; its preserved portion is characterized by nine or ten elongates rays (preserved portion represent 18.3 % of SL).</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">The dorsal fin is located in the posterior third of the body (predorsal length represents 79.9 % of SL) and consists of about ten rays. Its endoskeleton is partially preserved, but clearly shows relatively large pterygiophores with the anterior-most one, the so-called “rayless pterygiophore”, that inserts in the fifth interneural space.</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">The anal fin is not completely preserved, having at least six preserved rays. The anal fin is located just below the posterior part of the dorsal fin and was probably supported by eight pterygiophores, of which the anterior is the longest.</paragraph>
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<paragraph blockId="3.[106,745,151,1524]" pageId="3" pageNumber="516">
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The body is covered with bifid spinules (
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<figureCitation box="[571,684,1468,1492]" captionStart="Text-fig" captionStartId="4.[129,210,1156,1177]" captionTargetBox="[129,764,154,1141]" captionTargetId="figure-574@4.[129,764,154,1141]" captionTargetPageId="4" captionText="Text-fig. 3. Archaeotetraodon winterbottomi. Specimen NMN- HU-P PI 1995. Details of dermal bifid spinules." figureDoi="http://doi.org/10.5281/zenodo.7522599" httpUri="https://zenodo.org/record/7522599/files/figure.png" pageId="3" pageNumber="516">Text-fig. 3</figureCitation>
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). The spinules apparently lack stellate basal plates.
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</paragraph>
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<paragraph blockId="3.[106,745,1547,1796]" pageId="3" pageNumber="516">
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M e a s u r e m e n t s. Standard length:
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<quantity box="[583,678,1547,1571]" metricMagnitude="-2" metricUnit="m" metricValue="7.12" pageId="3" pageNumber="516" unit="mm" value="71.2">71.2 mm</quantity>
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; total measurable length:
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<quantity box="[319,414,1579,1603]" metricMagnitude="-2" metricUnit="m" metricValue="8.62" pageId="3" pageNumber="516" unit="mm" value="86.2">86.2 mm</quantity>
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; head length (from premaxilla to posterior wall of the basioccipital):
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<quantity box="[531,622,1612,1636]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="3" pageNumber="516" unit="mm" value="27.1">27.1 mm</quantity>
|
|||
|
|
(38.1 % of SL); height/length ratio of the posteriormost abdominal centrum 0.63; rayless pterygiophore length:
|
|||
|
|
<quantity box="[577,648,1676,1700]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="516" unit="mm" value="12.0">12 mm</quantity>
|
|||
|
|
(18 % of SL); predorsal length:
|
|||
|
|
<quantity box="[350,441,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="5.6899999999999995" pageId="3" pageNumber="516" unit="mm" value="56.9">56.9 mm</quantity>
|
|||
|
|
(79.9 % of SL); measurable caudal fin:
|
|||
|
|
<quantity box="[237,308,1740,1764]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="516" unit="mm" value="13.0">13 mm</quantity>
|
|||
|
|
(18.3 % of SL); ventral postcleithrum length:
|
|||
|
|
<quantity box="[187,278,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="3" pageNumber="516" unit="mm" value="15.1">15.1 mm</quantity>
|
|||
|
|
(21.2 % of SL).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|