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111 lines
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<document id="8B889C31AC03E6F79022E536A478C86F" ID-DOI="http://dx.doi.org/10.3897/zookeys.423.7536" ID-GBIF-Dataset="1f17876d-2290-4d36-aa1f-10b951701679" ID-PMC="PMC4106094" ID-Pensoft-Pub="1313-2970-423-1" ID-PubMed="25061393" ID-ZooBank="55B4F9C858934F88841660FF730E8872" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-423-1" ModsDocOrigin="ZooKeys 423" ModsDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" checkinTime="1451245649604" checkinUser="pensoft" docAuthor="Brunke, Adam J. & Solodovnikov, Alexey" docDate="2014" docId="346268A36A1CD71BB62E8921BFA3E758" docLanguage="en" docName="ZooKeys 423: 1-113" docOrigin="ZooKeys 423" docSource="http://dx.doi.org/10.3897/zookeys.423.7536" docTitle="Bolitogyrus costaricensis Wendeler 1927" docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="FFAB12065F4CFFED925DFFB46A5FFFFB" masterDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" masterLastPageNumber="113" masterPageNumber="1" pageNumber="21" updateTime="1701318337016" updateUser="plazi">
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<mods:titleInfo id="70D8168713D75E4A9819B49F5D932E09">
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<mods:title id="02DA7BE75D6931A1DBA594B73B31B698">A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)</mods:title>
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<mods:name id="902D2F04BB0EAC7E9EE0625EC3873621" type="personal">
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<mods:roleTerm id="E82A7FC82DF5D6B9864BD4A76890FA9F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D9E4C6DD55E9B5C6111014FAAD64656A">Brunke, Adam J.</mods:namePart>
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</mods:name>
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<mods:name id="00F656180337A72162243CE4DA2CFA56" type="personal">
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</mods:role>
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<mods:namePart id="1CEFB20DABE936633FC34541C0CCE152">Solodovnikov, Alexey</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="24C6630610BAF1FC6D59E261EA809EDE">text</mods:typeOfResource>
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<mods:titleInfo id="2BB2A898409A63026981E86042511209">
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<mods:title id="B22B0143947F5217DF0F4ED92A3E5748">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="73E0CDD52C39272E0F657CB04971E0B2">
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<mods:date id="81A5AC4B6261514C7CE054F233B536AC">2014</mods:date>
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<mods:detail id="5CFE33071504A470DD2AD53F3F1B7F10" type="volume">
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<mods:number id="A1BE2BFC323290949DAC0906B98776B2">423</mods:number>
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</mods:detail>
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<mods:start id="D49C90360C37FFCD10CD959F413B0748">1</mods:start>
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<mods:end id="26B053EAEB4E14BF6B7BCF4DF7B6C845">113</mods:end>
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<mods:location id="04B186680A359BB00DC468B1A0D0E21D">
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<mods:url id="0A353E07F5BAB946D20794E371EB6ACD">http://dx.doi.org/10.3897/zookeys.423.7536</mods:url>
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</mods:location>
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<mods:classification id="9B808B11A616A9BF07C41CAD55DB64B1">journal article</mods:classification>
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<mods:identifier id="FC56F52DB8BFA0F171CF9A51DFA2E3C0" type="DOI">http://dx.doi.org/10.3897/zookeys.423.7536</mods:identifier>
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<mods:identifier id="54E6B77542B40D87C20B0D0A14FF84F5" type="Pensoft-Pub">1313-2970-423-1</mods:identifier>
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<mods:identifier id="A9BA4231B68C06A6B4C471F4BE3B169C" type="ZooBank">55B4F9C858934F88841660FF730E8872</mods:identifier>
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</mods:mods>
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<treatment id="346268A36A1CD71BB62E8921BFA3E758" ID-GBIF-Taxon="152054115" LSID="urn:lsid:plazi:treatment:346268A36A1CD71BB62E8921BFA3E758" httpUri="http://treatment.plazi.org/id/346268A36A1CD71BB62E8921BFA3E758" lastPageNumber="21" pageId="20" pageNumber="21">
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<subSubSection id="7C126106C54E710C9112805CAF3FF9AB" pageId="20" pageNumber="21" type="multiple">
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<paragraph id="C545FC36E80D1F851052F1333B3ADD11" pageId="20" pageNumber="21">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
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</subSubSection>
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<subSubSection id="59553939EC123F2804E6EB341C4078B0" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="39537CFE28D85378005188C20FBD7B2F" pageId="20" pageNumber="21">
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<taxonomicName id="888DA5D3C01A46DE75ED7A036E6A3DF0" ID-CoL="MDTH" authority="Wendeler, 1927" authorityName="Wendeler" authorityYear="1927" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis (Wendeler, 1927)</taxonomicName>
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Figs 5A, 6B, 7
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<normalizedToken id="4CB50D27DDF026E2E6ED79BE19C2E668" originalValue="E–F">E-F</normalizedToken>
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, 11A, 12C, 13A, D, 15
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<normalizedToken id="26963D176BE9C5E152FD21D6B71ECBF1" originalValue="E–H">E-H</normalizedToken>
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, 23B, 25
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<normalizedToken id="9C882860894D093D4F23924F7BE97F3D" originalValue="A–D">A-D</normalizedToken>
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, 29
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<normalizedToken id="E6CDA22A8D1D70D4C2832DC468B686BE" originalValue="A–B">A-B</normalizedToken>
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(maps)
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</paragraph>
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</subSubSection>
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<subSubSection id="DCFFFC252E02E1BF99C2C9CC20C3D850" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="8B8FD8A5F9F7386185E0DDD839FF7D89" pageId="20" pageNumber="21">
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<taxonomicName id="2FE361455109D2491C3E1DD14E3F678A" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Cyrtothorax costaricensis</taxonomicName>
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Wendeler, 1927: 8
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</paragraph>
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<paragraph id="31A4AF6F29F5EEED3F89644BB0DEC8DA" pageId="20" pageNumber="21">
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<taxonomicName id="F97397B12956DC8C9ABF7682073EDD9F" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Cyrtothorax costaricensis</taxonomicName>
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:
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<bibRefCitation id="23E06E5C660FF7C2795ED1C48C0C76A4" author="Scheerpeltz, O" journalOrPublisher="Reichenbachia" pageId="72" pageNumber="73" pagination="175 - 192" title="Studien an den Arten der Gattung Cyrtothorax Kraatz mit Beschreibung neuer Arten sowie einer Dichotomik aller bis heute bekannt gewordenen Arten dieser Gattung (Coleoptera, Staphylinindae, Staphylininae, Quediini)." volume="15" year="1974">Scheerpeltz 1974</bibRefCitation>
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: 181 (in key)
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</paragraph>
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<paragraph id="AC216CF46CB8053B59D6E027FC098CF3" pageId="20" pageNumber="21">
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<taxonomicName id="6372957426F192732F22119D71352AFB" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax nevermanni" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nevermanni">Cyrtothorax nevermanni</taxonomicName>
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Scheerpeltz, 1974: 190, syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="4A2F30668BB24A8A869E9652B17C3854" pageId="20" pageNumber="21" type="type locality">
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<paragraph id="9F9E42D53D8B6C310708B768B5CEF513" pageId="20" pageNumber="21">Type locality.</paragraph>
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<paragraph id="BD7499067BDECDA7FDC00AFC6EB031B0" pageId="20" pageNumber="21">Turrialba Volcano, Cartago, Costa Rica</paragraph>
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</subSubSection>
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<subSubSection id="140343140BFF1BC8EF35008A0B3F9084" pageId="20" pageNumber="21" type="type material">
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<paragraph id="DC29A053FF5D41385F85829C47A65DD5" pageId="20" pageNumber="21">Type material.</paragraph>
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<paragraph id="53E656917B8722964855AAD3880BCB7D" pageId="20" pageNumber="21">
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<taxonomicName id="9AB3D226E93E1CD1F0DAC542C4EE9D23" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Cyrtothorax costaricensis</taxonomicName>
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Wendeler, 1927
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</paragraph>
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</subSubSection>
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<subSubSection id="827E2BD62F62F2851D1A201A8DFB9201" pageId="20" pageNumber="21" type="holotype">
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<paragraph id="EA6C918DDEE91572C95B64B1A855194C" pageId="20" pageNumber="21">Holotype</paragraph>
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<paragraph id="783C8EEB74BA7907E6CD76D60D4A1E21" pageId="20" pageNumber="21">
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(♀, ZMHB): ZENTR. AMERIKA, Costarica, Vukan Turialba, 10 km
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<normalizedToken id="B117D2E1CAFEB703BAC536F582E7CBE6" originalValue="NÖ">NOE</normalizedToken>
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[northeast] v.[ulkan] Irazu [beige printed label]/
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<taxonomicName id="C56ED2A516885AFDCE60177B4E8CF80F" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Cyrtothorax costaricensis</taxonomicName>
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, Wendeler [white label, name in script, author in print]/costaricensis [large green label, folded, hand printed]/Holotypus [red label, printed]/
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<taxonomicName id="55144DB86F95AF0E5ED7C7B6A4CCCD33" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Bolitogyrus</taxonomicName>
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, costaricensis, Det. Chatzimanolis 2010 [white label, hand printed] / Holotype ♀,
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<taxonomicName id="E75CC97492741AB3FB04A9162DD6F2EA" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Cyrtothorax costaricensis</taxonomicName>
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Wendeler, 1927, det. A. Brunke 2013 [red printed label].
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</paragraph>
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<paragraph id="FFCB515704B7AE06DD4758411F1A1A12" pageId="20" pageNumber="21">
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The specimen located in ZMHB is interpreted as the holotype as
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<bibRefCitation id="B32D0DC15999C6AC7530FEFBF1AC53DF" author="Wendeler, H" journalOrPublisher="Neue Beitraege zur systematischen Insektenkunde" pageId="72" pageNumber="73" title="Neue exotische Staphyliniden (Col.)." volume="4" year="1927">Wendeler (1927)</bibRefCitation>
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stated that only one specimen was seen. The female holotype possesses blue-purple metallic reflections on the elytra rather than deep purple, as stated by
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<bibRefCitation id="123C6C740091C9D3C4621A69CCB93C50" author="Wendeler, H" journalOrPublisher="Neue Beitraege zur systematischen Insektenkunde" pageId="72" pageNumber="73" title="Neue exotische Staphyliniden (Col.)." volume="4" year="1927">Wendeler (1927)</bibRefCitation>
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, a dark abdominal apex, fine abdominal microsculpture, a dark apical band on the mesofemur, a characteristically shaped tergite X and entirely dark forecoxae. Therefore, it is morphologically consistent with a species with known male sexual characters that is distributed from Guatemala to Costa Rica, here considered distinct from
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<taxonomicName id="C5F5B3F12CE6A251EFA5B2BEB36BD724" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
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based on features of the male genitalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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