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<document id="619E872D2A35E53A36381173CB588D45" ID-CLB-Dataset="30756" ID-DOI="10.11646/zootaxa.4379.1.5" ID-GBIF-Dataset="2c7fe9b1-f5a7-4169-bcc0-d544c884e01f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1172356" ID-ZooBank="E860555F-9CED-4A38-AAF8-B7AB5C1A7E71" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518511506799" checkinUser="plazi" docAuthor="Rasool, Iftekhar, Abdel-Dayem, Mahmoud S., Felix, Ron F. F. L. &amp; Aldhafer, Hathal M." docDate="2018" docId="03DB41541F6CFFE2F1CBCFA5D3F0F904" docLanguage="en" docName="zootaxa.4379.1.5.pdf" docOrigin="Zootaxa 4379 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebia (Nematopeza) melanura" docType="treatment" docVersion="8" lastPageNumber="92" masterDocId="FFE2392C1F69FFE7F15CCF32D75CFF96" masterDocTitle="A review of the Subtribe Lebiina Bonelli (Lebiini, Carabidae, Coleoptera) from Southwest of Saudi Arabia" masterLastPageNumber="102" masterPageNumber="87" pageNumber="92" updateTime="1698521292142" updateUser="ExternalLinkService">
<mods:mods id="F4C7FEA48C45A2C88DA8E66FCED14531" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E807178325B3433C03B6CD0B0D7019B4">
<mods:title id="002A31DCA7FE3E2631DF7348DE8C6156">A review of the Subtribe Lebiina Bonelli (Lebiini, Carabidae, Coleoptera) from Southwest of Saudi Arabia</mods:title>
</mods:titleInfo>
<mods:name id="DB083FB2CD1C5DB4C0AD68A58BA34874" type="personal">
<mods:role id="45342F7D2CE3B49D74321CB81E3F4681">
<mods:roleTerm id="3C625F834199A022232A32830543DBE4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC3AE76E1A66E73FD7102A0E01E24AC9">Rasool, Iftekhar</mods:namePart>
</mods:name>
<mods:name id="3E6F0FB897A73C9912B05002BDF53568" type="personal">
<mods:role id="0A5FE0D6D251E4BCA4005951466F0CA4">
<mods:roleTerm id="8B8A4E497CE706B775480F1E66220208">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="899D23E9A1BB393588EBF57A861DC080">Abdel-Dayem, Mahmoud S.</mods:namePart>
</mods:name>
<mods:name id="87F586141F723295AA4CE5A14A43874F" type="personal">
<mods:role id="65C187AAAA081E9B65A62AD199F9632A">
<mods:roleTerm id="133212E7E3CC00CB87E1CEEB25FE2973">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="635F248B570AB687BEA26A69C5CA89F6">Felix, Ron F. F. L.</mods:namePart>
</mods:name>
<mods:name id="8DCC22EFB669A06BF02CF969381403B8" type="personal">
<mods:role id="A5B6ECE8CF29B69E89D8EDCBEB4BC9A3">
<mods:roleTerm id="2898169205C3D003532FF4AA7429AC23">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C889AC780AA730784AEAABB441B9ED0F">Aldhafer, Hathal M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C267BFA5B20706A9B269D116EF5B7B65">text</mods:typeOfResource>
<mods:relatedItem id="62EDEF917EF65E08B3704F0B4EB974AA" type="host">
<mods:titleInfo id="D92D21067F512E03FB1EE24FE4AFF212">
<mods:title id="180A935A8B229B883C8500EC1D8E17A8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="53A08FA6D788365ACC561534EE51141C">
<mods:date id="9ACFFFDE124EF7ED6E7A16DDF1A04000">2018</mods:date>
<mods:detail id="284055E5E8BC0DA9FD960A8550ADD38A" type="pubDate">
<mods:number id="499F4D02F14604DCCBCCF624F160BA27">2018-02-13</mods:number>
</mods:detail>
<mods:detail id="26E43A5E6D2AC2B97B44672A7451E37C" type="volume">
<mods:number id="46EAD3613C26CF24943FBD4886791197">4379</mods:number>
</mods:detail>
<mods:detail id="69A78B18A9C7DD394DC70B2AEBB18C71" type="issue">
<mods:number id="33C0EA0A711128BAD53A84AC70555310">1</mods:number>
</mods:detail>
<mods:extent id="325B8E11706533E0E0522034A1B05ACD" unit="page">
<mods:start id="F159D2250D01AD3B7C4E4756C5F92DA7">87</mods:start>
<mods:end id="39677837326CC990DC6F30DB0E0DC0A4">102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0E8911792FAB29F360F5C4D66435AB93">journal article</mods:classification>
<mods:identifier id="B3154BBF71237A7D2F32B96A8C9603CF" type="CLB-Dataset">30756</mods:identifier>
<mods:identifier id="C633BCD43B6E52657A84ED538526FBC0" type="DOI">10.11646/zootaxa.4379.1.5</mods:identifier>
<mods:identifier id="90623EE4189382580D29E36B3438113F" type="GBIF-Dataset">2c7fe9b1-f5a7-4169-bcc0-d544c884e01f</mods:identifier>
<mods:identifier id="B3A473629E65EA59B755A8BA7E0B0524" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C0318C0791FA248209F95CC0CADD0E93" type="Zenodo-Dep">1172356</mods:identifier>
<mods:identifier id="A1EE0BD6317FCE6F2A1A0A11E073BCDB" type="ZooBank">E860555F-9CED-4A38-AAF8-B7AB5C1A7E71</mods:identifier>
</mods:mods>
<treatment id="03DB41541F6CFFE2F1CBCFA5D3F0F904" ID-DOI="http://doi.org/10.5281/zenodo.5979213" ID-GBIF-Taxon="140583187" ID-Zenodo-Dep="5979213" LSID="urn:lsid:plazi:treatment:03DB41541F6CFFE2F1CBCFA5D3F0F904" httpUri="http://treatment.plazi.org/id/03DB41541F6CFFE2F1CBCFA5D3F0F904" lastPageNumber="92" pageId="5" pageNumber="92">
<subSubSection id="C368A3C91F6CFFE2F1CBCFA5D59EFF24" box="[151,706,151,178]" pageId="5" pageNumber="92" type="nomenclature">
<paragraph id="8BCDF0421F6CFFE2F1CBCFA5D59EFF24" blockId="5.[151,706,151,178]" box="[151,706,151,178]" pageId="5" pageNumber="92">
<heading id="D085472E1F6CFFE2F1CBCFA5D59EFF24" bold="true" box="[151,706,151,178]" fontSize="11" level="1" pageId="5" pageNumber="92" reason="1">
<taxonomicName id="4C728BC11F6CFFE2F1CBCFA5D59EFF24" ID-CoL="3SQBS" authority="(Dejean, 1831)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[151,706,151,178]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura" subGenus="Nematopeza">
<emphasis id="B9062C501F6CFFE2F1CBCFA5D59EFF24" bold="true" box="[151,706,151,178]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F1CBCFA5D55AFF27" bold="true" box="[151,518,151,177]" italics="true" pageId="5" pageNumber="92">Lebia (Nematopeza) melanura</emphasis>
(
<bibRefCitation id="EFE38DB31F6CFFE2F34ACFAAD5E6FF24" box="[534,698,152,178]" pageId="5" pageNumber="92" refId="ref11429" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le comte Dejean. Tome cinquieme. Mequignon- Marviv, Paris, VIII + 883 pp." type="book">Dejean, 1831</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F6CFFE2F1CBCFD2D560FF62" box="[151,572,222,245]" pageId="5" pageNumber="92" type="reference_group">
<paragraph id="8BCDF0421F6CFFE2F1CBCFD2D560FF62" blockId="5.[151,572,222,245]" box="[151,572,222,245]" pageId="5" pageNumber="92">
<treatmentCitationGroup id="AB62D76C1F6CFFE2F1CBCFD2D560FF62" box="[151,572,222,245]" pageId="5" pageNumber="92">
<taxonomicName id="4C728BC11F6CFFE2F1CBCFD2D560FF62" ID-CoL="466YG" authority="Dejean, 1831: 370" authorityName="Dejean" authorityPageNumber="370" authorityYear="1831" box="[151,572,222,245]" class="Insecta" family="Carabidae" genus="Nematopeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F1CBCFD2D627FF62" box="[151,379,222,245]" italics="true" pageId="5" pageNumber="92">Nematopeza melanura</emphasis>
<treatmentCitation id="0AD3D6531F6CFFE2F0DECFECD560FF62" author="" box="[386,572,222,244]" page="370" pageId="5" pageNumber="92">
<bibRefCitation id="EFE38DB31F6CFFE2F0DECFECD550FF62" box="[386,524,222,244]" pageId="5" pageNumber="92" refId="ref11429" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le comte Dejean. Tome cinquieme. Mequignon- Marviv, Paris, VIII + 883 pp." type="book">Dejean, 1831</bibRefCitation>
: 370
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F6CFFE2F1CBCE13D5FEFBB8" pageId="5" pageNumber="92" type="materials_examined">
<paragraph id="8BCDF0421F6CFFE2F1CBCE13D6C5FEAC" blockId="5.[151,1437,289,1682]" box="[151,409,289,314]" pageId="5" pageNumber="92">
<materialsCitation id="3B1AFA1F1F6CFFE2F1CBCE13D6C5FEAC" ID-GBIF-Occurrence="1806563531" box="[151,409,289,314]" country="Senegal" location="Senegal" pageId="5" pageNumber="92" specimenCount="1" typeStatus="holotype">
<emphasis id="B9062C501F6CFFE2F1CBCE13D665FEAC" bold="true" box="[151,313,289,314]" pageId="5" pageNumber="92">
<typeStatus id="54C94EE01F6CFFE2F1CBCE13D78EFEAC" box="[151,210,289,314]" pageId="5" pageNumber="92">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F365B0D21F6CFFE2F01CCE13D6C5FEAC" box="[320,409,289,314]" name="Senegal" pageId="5" pageNumber="92">Senegal</collectingCountry>
</materialsCitation>
</paragraph>
<paragraph id="8BCDF0421F6CFFE2F19BCE77D5F6FECB" blockId="5.[151,1437,289,1682]" box="[199,682,325,350]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCE77D6CCFEC8" bold="true" box="[199,400,325,350]" pageId="5" pageNumber="92">Type depository:</emphasis>
Syntype male in MNHN
</paragraph>
<paragraph id="8BCDF0421F6CFFE2F19BCE5BD3FCFE5C" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<materialsCitation id="3B1AFA1F1F6CFFE2F19BCE5BD60FFE30" ID-GBIF-Occurrence="1806563526" collectorName="Material &amp; Total" country="Saudi Arabia" latitude="18.11635" location="Al Hubail" longLatPrecision="1" longitude="42.232315" municipality="Wadi Reem" pageId="5" pageNumber="92" specimenCount="17" specimenCount-female="5" specimenCount-male="3" stateProvince="Asir">
<emphasis id="B9062C501F6CFFE2F19BCE5BD6EAFE14" bold="true" box="[199,438,361,386]" pageId="5" pageNumber="92">
<collectorName id="268795941F6CFFE2F19BCE5BD673FE14" box="[199,303,361,386]" pageId="5" pageNumber="92">Material</collectorName>
examined:
</emphasis>
<collectorName id="268795941F6CFFE2F09DCE5BD6A6FE14" box="[449,506,361,386]" pageId="5" pageNumber="92">Total</collectorName>
<specimenCount id="9D743BCB1F6CFFE2F359CE5BD5CBFE14" box="[517,663,361,386]" pageId="5" pageNumber="92" type="generic">9 specimens</specimenCount>
:
<emphasis id="B9062C501F6CFFE2F3FACE5BD5BEFE14" bold="true" box="[678,738,361,386]" pageId="5" pageNumber="92">
<collectingRegion id="49B63EA01F6CFFE2F3FACE5BD581FE14" box="[678,733,361,386]" country="Saudi Arabia" name="'Asir" pageId="5" pageNumber="92">Asir</collectingRegion>
:
</emphasis>
<specimenCount id="9D743BCB1F6CFFE2F3B1CE5BD447FE17" box="[749,795,361,385]" pageId="5" pageNumber="92" type="male">3 ♂</specimenCount>
and
<specimenCount id="9D743BCB1F6CFFE2F206CE5BD4D4FE17" box="[858,904,361,386]" pageId="5" pageNumber="92" type="female">5 ♀</specimenCount>
,
<collectingCountry id="F365B0D21F6CFFE2F2C6CE5BD489FE17" box="[922,981,361,385]" name="Saudi Arabia" pageId="5" pageNumber="92">KSA</collectingCountry>
, “
<location id="8EADA6991F6CFFE2F2B3CE5BD33AFE14" LSID="urn:lsid:plazi:treatment:03DB41541F6CFFE2F1CBCFA5D3F0F904:8EADA6991F6CFFE2F2B3CE5BD33AFE14" box="[1007,1126,361,386]" country="Saudi Arabia" latitude="18.11635" longLatPrecision="1" longitude="42.232315" municipality="Wadi Reem" name="Al Hubail" pageId="5" pageNumber="92" stateProvince="Asir">Al Hubail</location>
,
<collectingMunicipality id="6BA96A381F6CFFE2F52ACE5BD3A3FE17" box="[1142,1279,361,386]" pageId="5" pageNumber="92">Wadi Reem</collectingMunicipality>
,
<geoCoordinate id="EE4696851F6CFFE2F452CE5BD2C1FE14" box="[1294,1437,361,386]" direction="north" orientation="latitude" pageId="5" pageNumber="92" precision="1" value="18.11635">N18°06.981'</geoCoordinate>
<geoCoordinate id="EE4696851F6CFFE2F1CBCEBCD67EFE30" box="[151,290,398,422]" direction="east" orientation="longitude" pageId="5" pageNumber="92" precision="1" value="42.232315">E42°13.939'</geoCoordinate>
Alt
</materialsCitation>
.
<materialsCitation id="3B1AFA1F1F6CFFE2F03DCEBFD6B8FE5C" ID-GBIF-Occurrence="1806563546" collectingDate="2014-10-20" collectionCode="KSMA" collectorName="I. Rasool &amp; M. Al Harbi" country="Saudi Arabia" county="Alt" elevation="451" latitude="18.11635" location="Al Hubail" longLatPrecision="1" longitude="42.232315" municipality="Wadi Reem" pageId="5" pageNumber="92" specimenCount="1" specimenCount-male="1" stateProvince="Asir">
<elevation id="005F17711F6CFFE2F03DCEBFD6F2FE33" box="[353,430,397,422]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="5" pageNumber="92" unit="m" value="451.0">
<quantity id="4C8A5DA71F6CFFE2F03DCEBFD6F2FE33" box="[353,430,397,422]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="5" pageNumber="92" unit="m" value="451.0">451 m</quantity>
</elevation>
,
<collectingDate id="EF882F6A1F6CFFE2F0E1CEBCD56BFE30" box="[445,567,398,422]" pageId="5" pageNumber="92" value="2014-10-20">
<date id="FFCCD6821F6CFFE2F0E1CEBCD56BFE30" box="[445,567,398,422]" pageId="5" pageNumber="92" value="2014-10-20">20.X.2014</date>
</collectingDate>
, (LT),
<collectorName id="268795941F6CFFE2F3D5CEBCD5AFFE30" box="[649,755,397,422]" pageId="5" pageNumber="92">I. Rasool</collectorName>
&amp;
<collectorName id="268795941F6CFFE2F242CEBCD4EDFE30" box="[798,945,397,422]" pageId="5" pageNumber="92">M. Al Harbi</collectorName>
”
<emphasis id="B9062C501F6CFFE2F299CEBFD36FFE30" bold="true" box="[965,1075,397,422]" pageId="5" pageNumber="92">
[
<collectionCode id="ED6368871F6CFFE2F291CEBFD37BFE30" box="[973,1063,397,422]" pageId="5" pageNumber="92">KSMA</collectionCode>
].
</emphasis>
<specimenCount id="9D743BCB1F6CFFE2F562CEBCD332FE33" box="[1086,1134,398,422]" pageId="5" pageNumber="92" type="male">1 ♂</specimenCount>
, “
<location id="8EADA6991F6CFFE2F5D4CEBFD25CFE30" LSID="urn:lsid:plazi:treatment:03DB41541F6CFFE2F1CBCFA5D3F0F904:8EADA6991F6CFFE2F5D4CEBFD25CFE30" box="[1160,1280,397,422]" country="Saudi Arabia" county="Alt" latitude="18.11635" longLatPrecision="1" longitude="42.232315" municipality="Wadi Reem" name="Al Hubail" pageId="5" pageNumber="92" stateProvince="Asir">Al Hubail</location>
,
<collectingMunicipality id="6BA96A381F6CFFE2F452CEBFD2C4FE33" box="[1294,1432,397,422]" pageId="5" pageNumber="92">Wadi Reem</collectingMunicipality>
,
<geoCoordinate id="EE4696851F6CFFE2F1CBCE83D679FE5C" box="[151,293,433,458]" direction="north" orientation="latitude" pageId="5" pageNumber="92" precision="1" value="18.11635">N18°06.981'</geoCoordinate>
<geoCoordinate id="EE4696851F6CFFE2F077CE83D6EBFE5F" box="[299,439,433,457]" direction="east" orientation="longitude" pageId="5" pageNumber="92" precision="1" value="42.232315">E42°13.939'</geoCoordinate>
<collectingCounty id="62AC88CE1F6CFFE2F0E1CE83D6B8FE5C" box="[445,484,433,458]" pageId="5" pageNumber="92">Alt</collectingCounty>
</materialsCitation>
.
<quantity id="4C8A5DA71F6CFFE2F0B2CE83D56BFE5F" box="[494,567,433,458]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="5" pageNumber="92" unit="m" value="451.0">451 m</quantity>
,
<date id="FFCCD6821F6CFFE2F31FCE83D5E0FE5F" box="[579,700,433,457]" pageId="5" pageNumber="92" value="2014-10-20">20.X.2014</date>
, (LT), I. Rasool &amp; M. Al Harbi”
<emphasis id="B9062C501F6CFFE2F571CE83D3FCFE5C" bold="true" box="[1069,1184,433,458]" pageId="5" pageNumber="92">[RMNH].</emphasis>
</paragraph>
<paragraph id="8BCDF0421F6CFFE2F19BCEE7D5FEFBB8" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCEE7D601FE78" bold="true" box="[199,349,469,494]" pageId="5" pageNumber="92">Recognition.</emphasis>
Small sized beetles (
<figureCitation id="1349ECC71F6CFFE2F314CEE7D5C0FE78" box="[584,668,469,494]" captionStart="FIGURES 18–26" captionStartId="10.[151,264,1882,1904]" captionTargetBox="[159,1430,200,1857]" captionTargetId="figure@10.[151,1436,193,1860]" captionTargetPageId="10" captionText="FIGURES 18–26. Habitus of Lebia: (18) L. auberti (Fairmaire, 1892), (19) L. melanacra (Chaudoir, 1878), (20) L. melanura (Dejean, 1831), (21, 22) L. raeesae sp. n. (23) Holotype. L. nilotica Chaudoir, 1871, (24, 25) L. nilotica Chaudoir, 1871, (26) Syntype.L.senegalensis Chaudoir, 1871" httpUri="https://zenodo.org/record/1172364/files/figure.png" pageId="5" pageNumber="92">Fig. 20</figureCitation>
), TBL
<quantity id="4C8A5DA71F6CFFE2F3B7CEE7D4CDFE7B" box="[747,913,469,494]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="6.25" metricValueMin="4.65" pageId="5" pageNumber="92" unit="mm" value="5.45" valueMax="6.25" valueMin="4.65">4.65–6.25 mm</quantity>
.
<emphasis id="B9062C501F6CFFE2F2C7CEE7D4AFFE7B" box="[923,1011,469,493]" italics="true" pageId="5" pageNumber="92">Colour:</emphasis>
Head black, pronotum, femora, tibiae and antennomeres I–III testaceous; rest of the antennae, mouth parts and tarsi dark brown; elytra testaceous with a trilobed black patch at posterior third, widened at apex to the margin of elytra, median lobe extending along suture to middle, occupying two inner intervals, lateral lobes prolonged up to apical third, occupying all intervals and left lateral margin.
<emphasis id="B9062C501F6CFFE2F012CD57D55EFDEB" box="[334,514,613,637]" italics="true" pageId="5" pageNumber="92">Microsculpture:</emphasis>
Elytra, head, clypeus and labrum with isodiametric mesh pattern; pronotum smooth; abdomen with transverse microlines.
<emphasis id="B9062C501F6CFFE2F3F8CDBBD5B6FD37" box="[676,746,649,673]" italics="true" pageId="5" pageNumber="92">Head:</emphasis>
Slightly wider than long, HL
<quantity id="4C8A5DA71F6CFFE2F51BCDB8D3B0FD37" box="[1095,1260,650,674]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.28" metricValueMin="0.92" pageId="5" pageNumber="92" unit="mm" value="1.1" valueMax="1.28" valueMin="0.92">0.92–1.28 mm</quantity>
and HW
<quantity id="4C8A5DA71F6CFFE2F401CDB8D7A3FD53" metricMagnitude="-3" metricUnit="m" metricValue="1.17" metricValueMax="1.35" metricValueMin="0.99" pageId="5" pageNumber="92" unit="mm" value="1.17" valueMax="1.35" valueMin="0.99">0.99– 1.35 mm</quantity>
, narrower than pronotum; frons and vertex sculptured, vertex with weak longitudinal irregular furrows, frons with irregular coarse punctures (
<figureCitation id="1349ECC71F6CFFE2F31CCDE3D5C8FD7C" box="[576,660,721,746]" captionStart="FIGURES30–37" captionStartId="11.[151,264,1938,1960]" captionTargetBox="[347,1211,836,1903]" captionTargetId="figure@11.[347,1211,824,1910]" captionTargetPageId="11" captionText="FIGURES30–37. Head and pronotum of Lebia: (30) L.auberti (Fairmaire,1892), (31) L.melanacra (Chaudoir,1878), (32) L. melanura (Dejean, 1831), (33) L. raeesae sp. n. (34) L. natalensis Chaudoir, 1871, (35) L. nilotica Chaudoir, 1871, (36) L. spec.(37) Matabele arabica Mateu, 1986." httpUri="https://zenodo.org/record/1172368/files/figure.png" pageId="5" pageNumber="92">Fig. 32</figureCitation>
); labrum truncate at anterior margin.
<emphasis id="B9062C501F6CFFE2F56ECDE0D3F5FD7F" box="[1074,1193,722,745]" italics="true" pageId="5" pageNumber="92">Pronotum.</emphasis>
Transverse, PL
<quantity id="4C8A5DA71F6CFFE2F401CDE0D7A6FC9B" metricMagnitude="-4" metricUnit="m" metricValue="9.3" metricValueMax="10.6" metricValueMin="8.0" pageId="5" pageNumber="92" unit="mm" value="0.93" valueMax="1.06" valueMin="0.8">0.80– 1.06 mm</quantity>
and PW
<quantity id="4C8A5DA71F6CFFE2F03DCDC7D559FC9B" box="[353,517,757,782]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" metricValueMax="1.68" metricValueMin="1.24" pageId="5" pageNumber="92" unit="mm" value="1.46" valueMax="1.68" valueMin="1.24">1.24–1.68 mm</quantity>
, anterior margin slightly concave, anterior angles strongly rounded, lateral margin anteriorly rounded, obliquely straight in posterior half, sinuate in front of posterior angles, posterior angles almost right, pronotal base strongly incised towards posterior angles and lobate in middle (
<figureCitation id="1349ECC71F6CFFE2F52ACC0FD393FCC0" box="[1142,1231,829,854]" captionStart="FIGURES30–37" captionStartId="11.[151,264,1938,1960]" captionTargetBox="[347,1211,836,1903]" captionTargetId="figure@11.[347,1211,824,1910]" captionTargetPageId="11" captionText="FIGURES30–37. Head and pronotum of Lebia: (30) L.auberti (Fairmaire,1892), (31) L.melanacra (Chaudoir,1878), (32) L. melanura (Dejean, 1831), (33) L. raeesae sp. n. (34) L. natalensis Chaudoir, 1871, (35) L. nilotica Chaudoir, 1871, (36) L. spec.(37) Matabele arabica Mateu, 1986." httpUri="https://zenodo.org/record/1172368/files/figure.png" pageId="5" pageNumber="92">Fig. 32</figureCitation>
); disc sculptured, median longitudinal impression deep.
<emphasis id="B9062C501F6CFFE2F315CC53D5C4FCEF" box="[585,664,865,889]" italics="true" pageId="5" pageNumber="92">Elytra:</emphasis>
Elongate subovoid, EL
<quantity id="4C8A5DA71F6CFFE2F2F1CC53D318FCEF" box="[941,1092,865,890]" metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.9" metricValueMin="3.25" pageId="5" pageNumber="92" unit="mm" value="3.575" valueMax="3.9" valueMin="3.25">3.25–3.9 mm</quantity>
and EW
<quantity id="4C8A5DA71F6CFFE2F5EDCC53D215FCEF" box="[1201,1353,865,890]" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.95" metricValueMin="2.3" pageId="5" pageNumber="92" unit="mm" value="2.625" valueMax="2.95" valueMin="2.3">2.3–2.95 mm</quantity>
, lateral margin obliquely convex; apex of elytra obliquely truncate, not serrate, striae complete, deep and punctate, intervals convex throughout.
<emphasis id="B9062C501F6CFFE2F0BECC98D54BFC57" box="[482,535,938,961]" italics="true" pageId="5" pageNumber="92">Legs</emphasis>
: Tibiae in mid leg of male with one preapical incision on inner side.
<emphasis id="B9062C501F6CFFE2F476CC9BD2C8FC57" box="[1322,1428,937,961]" italics="true" pageId="5" pageNumber="92">Abdomen</emphasis>
: Abdominal sterna moderately pubescent, last sternum rounded and tetra-setose in both sexes.
<emphasis id="B9062C501F6CFFE2F5E3CCFFD269FC73" box="[1215,1333,973,997]" italics="true" pageId="5" pageNumber="92">Aedeagus:</emphasis>
AL
<quantity id="4C8A5DA71F6CFFE2F437CCFCD79AFB9F" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="5" pageNumber="92" unit="mm" value="1.33">1.33 mm</quantity>
; in lateral view, it is slightly curved and sinuate dorsally (
<figureCitation id="1349ECC71F6CFFE2F23FCCC3D4E5FB9C" box="[867,953,1009,1034]" captionStart="FIGURES38–42" captionStartId="12.[151,264,890,912]" captionTargetBox="[154,1431,200,861]" captionTargetId="figure@12.[151,1436,193,868]" captionTargetPageId="12" captionText="FIGURES38–42. Aedeagusof Lebia: (38) L.auberti (Fairmaire,1892), (39)L.melanacra (Chaudoir, 1878), (40) L.melanura (Dejean, 1831), (41) L.raeesae sp. n. (42) L. nilotica Chaudoir,1871." httpUri="https://zenodo.org/record/1172370/files/figure.png" pageId="5" pageNumber="92">Fig. 40</figureCitation>
); with equal width along its length except apical lamina, apical lamina short and pointed.
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F6CFFE2F19BCB0BD46DFAE4" pageId="5" pageNumber="92" type="description">
<paragraph id="8BCDF0421F6CFFE2F19BCB0BD61AFB0C" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCB0BD676FBC4" bold="true" box="[199,298,1081,1106]" pageId="5" pageNumber="92">Affinity.</emphasis>
This species is close to
<taxonomicName id="4C728BC11F6CFFE2F363CB08D4D5FBC4" authority="(Alluaud, 1932)" baseAuthorityName="Alluaud" baseAuthorityYear="1932" box="[575,905,1081,1106]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="chaudoiri">
<emphasis id="B9062C501F6CFFE2F363CB08D596FBC7" box="[575,714,1081,1105]" italics="true" pageId="5" pageNumber="92">L. chaudoiri</emphasis>
(Alluaud, 1932)
</taxonomicName>
, but according to Alluaud (1932),
<taxonomicName id="4C728BC11F6CFFE2F44ECB08D2C0FBC7" authorityName="Dejean" authorityYear="1831" box="[1298,1436,1081,1105]" class="Insecta" family="Carabidae" genus="Nematopeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F44ECB08D2C0FBC7" box="[1298,1436,1081,1105]" italics="true" pageId="5" pageNumber="92">L. melanura</emphasis>
</taxonomicName>
can be differentiated by elytral black patch shorter, less protruding towards the scutellum, and finer punctuation on head and striae.
</paragraph>
<paragraph id="8BCDF0421F6CFFE2F19BCB97D7ABFA90" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCB97D6DAFB28" bold="true" box="[199,390,1189,1214]" pageId="5" pageNumber="92">Ecological note.</emphasis>
Members of this species were collected from elevation
<quantity id="4C8A5DA71F6CFFE2F544CB97D33FFB2B" box="[1048,1123,1189,1214]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="5" pageNumber="92" unit="m" value="451.0">451 m</quantity>
in valley with damp place having gravels and stones among different kinds of grasses. This species was collected by UV–light during October.
</paragraph>
<paragraph id="8BCDF0421F6CFFE2F19BCA23D46DFAE4" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCA23D556FABC" bold="true" box="[199,522,1297,1322]" pageId="5" pageNumber="92">Geographical distribution.</emphasis>
It is described from
<collectingCountry id="F365B0D21F6CFFE2F24ECA23D437FABC" box="[786,875,1297,1322]" name="Senegal" pageId="5" pageNumber="92">Senegal</collectingCountry>
(
<bibRefCitation id="EFE38DB31F6CFFE2F2DCCA23D342FABC" box="[896,1054,1297,1322]" pageId="5" pageNumber="92" refId="ref11429" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le comte Dejean. Tome cinquieme. Mequignon- Marviv, Paris, VIII + 883 pp." type="book">Dejean, 1831</bibRefCitation>
) also known from
<collectingCountry id="F365B0D21F6CFFE2F457CA20D2C4FABC" box="[1291,1432,1298,1322]" name="Ivory Coast" pageId="5" pageNumber="92">
Ivory
<collectingRegion id="49B63EA01F6CFFE2F409CA20D2C4FABC" box="[1365,1432,1298,1322]" country="Tanzania" name="Pwani" pageId="5" pageNumber="92">Coast</collectingRegion>
</collectingCountry>
,
<collectingCountry id="F365B0D21F6CFFE2F1CBCA07D7AFFAD8" box="[151,243,1333,1358]" name="Gambia" pageId="5" pageNumber="92">Gambia</collectingCountry>
,
<collectingCountry id="F365B0D21F6CFFE2F1A1CA07D612FAD8" box="[253,334,1333,1358]" name="Guinea-Bissau" pageId="5" pageNumber="92">Guinea</collectingCountry>
(
<bibRefCitation id="EFE38DB31F6CFFE2F000CA04D6A8FAD8" box="[348,500,1334,1358]" pageId="5" pageNumber="92" refId="ref11740" refString="Lorenz, W. (2017) CarabCat: Global database of ground beetles (version Sep 2013). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J., Penev, L. (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 30 October 2017, Species 2000: Naturalis, Leiden, ISSN 2405 - 8858. Available from: http: // www. catalogueoflife. org / col (accessed 20 January 2018)" type="journal article">Lorenz, 2017</bibRefCitation>
) and newly recorded during this study from
<collectingCountry id="F365B0D21F6CFFE2F2BFCA07D329FAD8" box="[995,1141,1333,1358]" name="Saudi Arabia" pageId="5" pageNumber="92">Saudi Arabia</collectingCountry>
(
<figureCitation id="1349ECC71F6CFFE2F5D8CA07D384FAD8" box="[1156,1240,1333,1358]" captionStart="FIGURES 43–44" captionStartId="14.[151,264,1102,1124]" captionTargetBox="[159,1412,193,1073]" captionTargetId="figure@14.[151,1436,193,1081]" captionTargetPageId="14" captionText="FIGURES 43–44. Distribution of Lebia spp. in South west of Saudi Arabia." httpUri="https://zenodo.org/record/1172372/files/figure.png" pageId="5" pageNumber="92">Fig. 43</figureCitation>
).
<taxonomicName id="4C728BC11F6CFFE2F5B6CA07D2C0FADB" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1258,1436,1333,1357]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F5B6CA07D2C0FADB" box="[1258,1436,1333,1357]" italics="true" pageId="5" pageNumber="92">Lebia melanura</emphasis>
</taxonomicName>
is an Afrotropical species that exemplifies AFA Chorotype.
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F6CFFE2F19BCA4FD3F0F904" pageId="5" pageNumber="92" type="discussion">
<paragraph id="8BCDF0421F6CFFE2F19BCA4FD3F0F904" blockId="5.[151,1437,289,1682]" pageId="5" pageNumber="92">
<emphasis id="B9062C501F6CFFE2F19BCA4FD60DFA00" bold="true" box="[199,337,1405,1430]" pageId="5" pageNumber="92">Comments:</emphasis>
<taxonomicName id="4C728BC11F6CFFE2F007CA4FD553FA03" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[347,527,1405,1429]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F007CA4FD553FA03" box="[347,527,1405,1429]" italics="true" pageId="5" pageNumber="92">Lebia melanura</emphasis>
</taxonomicName>
and
<taxonomicName id="4C728BC11F6CFFE2F316CA4CD58BFA03" baseAuthorityName="Alluaud" baseAuthorityYear="1932" box="[586,727,1405,1429]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="chaudoiri">
<emphasis id="B9062C501F6CFFE2F316CA4CD58BFA03" box="[586,727,1405,1429]" italics="true" pageId="5" pageNumber="92">L. chaudoiri</emphasis>
</taxonomicName>
are problematic to distinguish and often wrongly recognized. The morphological characters suggested by Alluaud (1932) to differentiate
<taxonomicName id="4C728BC11F6CFFE2F2BFCA90D332FA2F" baseAuthorityName="Alluaud" baseAuthorityYear="1932" box="[995,1134,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="chaudoiri">
<emphasis id="B9062C501F6CFFE2F2BFCA90D332FA2F" box="[995,1134,1441,1465]" italics="true" pageId="5" pageNumber="92">L. chaudoiri</emphasis>
</taxonomicName>
from
<taxonomicName id="4C728BC11F6CFFE2F5EECA90D267FA2F" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1202,1339,1441,1465]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F5EECA90D267FA2F" box="[1202,1339,1441,1465]" italics="true" pageId="5" pageNumber="92">L. melanura</emphasis>
</taxonomicName>
include: black suture longer extending towards the scutellum, smaller red humeral spots, stronger punctuation on the head, the striae coarser punctuate and the posterior angles of the pronotum more elevated. In
<taxonomicName id="4C728BC11F6CFFE2F5C9CAD8D27DF997" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1173,1313,1513,1537]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F5C9CAD8D27DF997" box="[1173,1313,1513,1537]" italics="true" pageId="5" pageNumber="92">L. melanura</emphasis>
</taxonomicName>
, all above distinguishing characters overlap with those of
<taxonomicName id="4C728BC11F6CFFE2F39FC93CD40EF9B3" baseAuthorityName="Alluaud" baseAuthorityYear="1932" box="[707,850,1549,1573]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="chaudoiri">
<emphasis id="B9062C501F6CFFE2F39FC93CD40EF9B3" box="[707,850,1549,1573]" italics="true" pageId="5" pageNumber="92">L. chaudoiri</emphasis>
</taxonomicName>
in several examined specimens from
<collectingCountry id="F365B0D21F6CFFE2F44FC93CD2CBF9B3" box="[1299,1431,1550,1574]" name="Ivory Coast" pageId="5" pageNumber="92">Ivory coast</collectingCountry>
,
<collectingRegion id="49B63EA01F6CFFE2F1CBC900D7B8F9DC" box="[151,228,1586,1610]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="5" pageNumber="92">
<collectingCountry id="F365B0D21F6CFFE2F1CBC900D7B8F9DC" box="[151,228,1586,1610]" name="Democratic Republic of the Congo" pageId="5" pageNumber="92">Congo</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F365B0D21F6CFFE2F1ADC903D610F9DC" box="[241,332,1585,1610]" name="Senegal" pageId="5" pageNumber="92">Senegal</collectingCountry>
,
<collectingCountry id="F365B0D21F6CFFE2F005C903D6CBF9DC" box="[345,407,1585,1610]" name="Chad" pageId="5" pageNumber="92">Chad</collectingCountry>
,
<collectingCountry id="F365B0D21F6CFFE2F0FFC903D554F9DC" box="[419,520,1585,1610]" name="Tanzania" pageId="5" pageNumber="92">Tanzania</collectingCountry>
and Arabian Peninsula. The genitalia of both species seem also the same. These results suggest that
<taxonomicName id="4C728BC11F6CFFE2F020C964D554F9FB" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[380,520,1621,1645]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="melanura">
<emphasis id="B9062C501F6CFFE2F020C964D554F9FB" box="[380,520,1621,1645]" italics="true" pageId="5" pageNumber="92">L. melanura</emphasis>
</taxonomicName>
and
<taxonomicName id="4C728BC11F6CFFE2F318C964D58EF9FB" baseAuthorityName="Alluaud" baseAuthorityYear="1932" box="[580,722,1621,1645]" class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="chaudoiri">
<emphasis id="B9062C501F6CFFE2F318C964D58EF9FB" box="[580,722,1621,1645]" italics="true" pageId="5" pageNumber="92">L. chaudoiri</emphasis>
</taxonomicName>
are identical species. However, the
<typeStatus id="54C94EE01F6CFFE2F52BC967D3B8F9F8" box="[1143,1252,1621,1646]" pageId="5" pageNumber="92" type="holotype">holotypes</typeStatus>
of both species were not found in their
<typeStatus id="54C94EE01F6CFFE2F0C2C948D692F904" box="[414,462,1658,1682]" pageId="5" pageNumber="92">type</typeStatus>
depository, so the authors keep the current status of both species.
</paragraph>
</subSubSection>
</treatment>
</document>
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