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<document id="A64F398DFB07D27A8118584B2F2E70CC" ID-DOI="10.11646/zootaxa.3904.4.1" ID-GBIF-Dataset="a1c49726-9afa-4be5-85ef-9d0caabac1fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="239899" ID-ZooBank="38457A27-A15B-4E87-BE72-9A3AB007CAA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460890290884" checkinUser="plazi" docAuthor="Yoshitomi, Hiroyuki" docDate="2015" docId="03AC6D78C817215449B2E9F9301CFD70" docLanguage="en" docName="zt03904p500.pdf" docOrigin="Zootaxa 3904 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon bicolor Yoshitomi, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="474" masterDocId="FF951500C80121434925E96E356EFFBD" masterDocTitle="New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)" masterLastPageNumber="500" masterPageNumber="451" pageNumber="473" updateTime="1698359547105" updateUser="plazi">
<mods:mods id="0E28773816FD87A3D5520891203A681E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="467D2052DD82874DACB63AAA41B2633E">
<mods:title id="7154CCF746D75E6E6B65FDB222E0F8C8">New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)</mods:title>
</mods:titleInfo>
<mods:name id="34FC6111557DCA123CBF63971F8C21D3" type="personal">
<mods:role id="6BDF3D6EDC224AE8B860819DF97E5F62">
<mods:roleTerm id="26BFFF1C2381F9C475A91FC275A52645">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A80EE6461670E6B1A64E514B7DD2B11E">Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C6D31F659AC3A38C16A690FBB29EA5C">text</mods:typeOfResource>
<mods:relatedItem id="99783A811C2EC7012207BC5A38B5BA5B" type="host">
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<mods:title id="BF3FB28C13368D24BCB116ED93386489">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2C7060359EA0A94F5DC851F2A5188D3F">
<mods:date id="1652ECDBB98C4515EA91A04DD745A87C">2015</mods:date>
<mods:detail id="16C42964561DBF120848A1513282251A" type="volume">
<mods:number id="E8C56803F35BE33CDC349751C4A0EE1A">3904</mods:number>
</mods:detail>
<mods:detail id="9EF5B1709058F399BAEE846EECA6F658" type="issue">
<mods:number id="67443F600318F2552EE97588BF6EFA6E">4</mods:number>
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<mods:classification id="731885676623AA3EBAB86DBC54F20E6A">journal article</mods:classification>
<mods:identifier id="9DFF00671FAF08617A107BF8CF311554" type="DOI">10.11646/zootaxa.3904.4.1</mods:identifier>
<mods:identifier id="98A1F38542372B1FCD2007A7189AEE26" type="GBIF-Dataset">a1c49726-9afa-4be5-85ef-9d0caabac1fe</mods:identifier>
<mods:identifier id="A64CBAD9D093A93F09E5A5920C578D6B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A40357A4ED8209D987781E4E9AC295A5" type="Zenodo-Dep">239899</mods:identifier>
<mods:identifier id="1DEFB504AAEB4A16163BA27D59F6100B" type="ZooBank">38457A27-A15B-4E87-BE72-9A3AB007CAA5</mods:identifier>
</mods:mods>
<treatment id="03AC6D78C817215449B2E9F9301CFD70" ID-DOI="http://doi.org/10.5281/zenodo.6111879" ID-GBIF-Taxon="119586904" ID-Zenodo-Dep="6111879" LSID="urn:lsid:plazi:treatment:03AC6D78C817215449B2E9F9301CFD70" httpUri="http://treatment.plazi.org/id/03AC6D78C817215449B2E9F9301CFD70" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
<subSubSection id="C31F8FE5C817215549B2E9F934CEFF6E" pageId="22" pageNumber="473" type="nomenclature">
<paragraph id="8BBADC6EC817215549B2E9F93483FF0F" blockId="22.[151,493,151,211]" box="[151,493,151,178]" pageId="22" pageNumber="473">
<heading id="D0F26B02C817215549B2E9F93483FF0F" bold="true" box="[151,493,151,178]" fontSize="11" level="1" pageId="22" pageNumber="473" reason="1">
<emphasis id="B971007CC817215549B2E9F93483FF0F" bold="true" box="[151,493,151,178]" pageId="22" pageNumber="473">
<taxonomicName id="4C05A7EDC817215549B2E9F934CAFF0C" box="[151,420,151,177]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="B971007CC817215549B2E9F934CAFF0C" bold="true" box="[151,420,151,177]" italics="true" pageId="22" pageNumber="473">Ypsiloncyphon bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C8172155488EE9F63483FF0F" box="[427,493,152,178]" pageId="22" pageNumber="473" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBADC6EC817215549B2E9D434CEFF6E" blockId="22.[151,493,151,211]" box="[151,416,186,211]" pageId="22" pageNumber="473">
(
<figureCitation id="133EC0EBC817215549BAE9D43594FF6E" box="[159,250,186,211]" captionStart="FIGURE 13" captionStartId="17.[151,250,1610,1632]" captionTargetBox="[155,1420,202,1578]" captionTargetId="figure@17.[151,1430,193,1583]" captionTargetPageId="17" captionText="FIGURE 13. Habitus of Ypsiloncyphon spp., holotype, male (except for F). A) Y. guamensis sp. n., B) Y. philippinensis sp. n., C) Y. panayensis sp. n., D) Y. chlorizanoides sp. n., E) Y. bicolor sp. n., F) Y. bicolor sp. n., paratype, female, G) Y. sumatranus sp. n., H) Y. acer sp. n., I) Y. nokrekensis sp. n., J) Y. boukali sp. n., K) Y. javanicus sp. n., L) Y. nepalensis sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239912/files/figure.png" pageId="22" pageNumber="473">Figs. 13</figureCitation>
E, 13F, 18, 19)
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C817215549B2E86C363AFE72" pageId="22" pageNumber="473" type="materials_examined">
<paragraph id="8BBADC6EC817215549B2E86C363AFE72" blockId="22.[151,1437,258,571]" pageId="22" pageNumber="473">
<emphasis id="B971007CC817215549B2E86C3429FEA6" bold="true" box="[151,327,258,283]" pageId="22" pageNumber="473">
<typeStatus id="54BE62CCC817215549B2E86C35BCFEA6" box="[151,210,258,283]" pageId="22" pageNumber="473">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BE62CCC81721554874E86C34D7FEA6" box="[337,441,258,283]" pageId="22" pageNumber="473" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED1444ABC817215548EEE86D3774FEA6" box="[459,538,259,283]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="473">NMW</collectionCode>
): male, “S-VIETNAM Nam Cat Tien Nat. Park
<date id="FFBBFAAEC81721554D6FE86D3198FEA6" box="[1098,1270,258,283]" pageId="22" pageNumber="473" value="1994-05-01" valueMax="1994-05-15" valueMin="1994-05-01">1. -15. 5. 1994</date>
Pacholatko &amp; Dembicky”, genit. s. no. HY 933.
<typeStatus id="54BE62CCC81721554B3CE84937E8FE82" box="[537,646,295,319]" pageId="22" pageNumber="473" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED1444ABC81721554BB0E84937B1FE82" box="[661,735,295,319]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="473">NMW</collectionCode>
,
<collectionCode id="ED1444ABC81721554BCEE8493659FE83" box="[747,823,295,318]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="22" pageNumber="473">EUMJ</collectionCode>
,
<collectionCode id="ED1444ABC81721554A67E84936FBFE82" box="[834,917,295,319]" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="22" pageNumber="473">NSMT</collectionCode>
):
<specimenCount id="9D0317E7C81721554A8CE8493163FE82" box="[937,1037,294,319]" pageId="22" pageNumber="473" type="male">14 males</specimenCount>
&amp;
<specimenCount id="9D0317E7C81721554D17E84931DEFE82" box="[1074,1200,294,319]" pageId="22" pageNumber="473" type="female">17 females</specimenCount>
, same data as for the
<typeStatus id="54BE62CCC817215549B2E8243593FEDE" box="[151,253,330,355]" pageId="22" pageNumber="473" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D0317E7C81721554828E8253409FEDE" box="[269,359,330,355]" pageId="22" pageNumber="473" type="male">2 males</specimenCount>
&amp;
<specimenCount id="9D0317E7C817215548B0E8253490FEDE" box="[405,510,330,355]" pageId="22" pageNumber="473" type="female">1 female</specimenCount>
, “NW-THAIL.: Chiang
<date id="FFBBFAAEC81721554A39E824361CFEDE" box="[796,882,330,355]" pageId="22" pageNumber="473" value="1998-05">
Mai
<geoCoordinate id="EE31BAA9C81721554A70E824361CFEDE" box="[853,882,330,355]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="925" value="98.95">98</geoCoordinate>
</date>
º57’E
<geoCoordinate id="EE31BAA9C81721554A99E8253173FEDE" box="[956,1053,331,355]" direction="north" orientation="latitude" pageId="22" pageNumber="473" precision="925" value="18.816668">18º49’N</geoCoordinate>
, Zoo
<date id="FFBBFAAEC81721554D43E8253063FEDE" box="[1126,1293,330,355]" pageId="22" pageNumber="473" value="1989-05-01" valueMax="1989-05-08" valueMin="1989-05-01">1. -8. 5. 1989</date>
, Malicky &amp; Chantaramongkol LF”;
<specimenCount id="9D0317E7C8172155488EE801377DFE3A" box="[427,531,366,391]" pageId="22" pageNumber="473" type="female">1 female</specimenCount>
, “N-THAILAND,
<date id="FFBBFAAEC81721554BD5E80136FFFE3A" box="[752,913,367,391]" pageId="22" pageNumber="473" value="1918-04-10" valueMax="1918-04-17" valueMin="1918-04-10">
10. -17. 4.
<geoCoordinate id="EE31BAA9C81721554A53E80136FFFE3A" box="[886,913,367,391]" direction="north" orientation="latitude" pageId="22" pageNumber="473" precision="925" value="18.816668">18</geoCoordinate>
</date>
º49’N
<geoCoordinate id="EE31BAA9C81721554AC5E8003155FE3A" box="[992,1083,366,391]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="925" value="98.95">98º57’E</geoCoordinate>
Chiang Mai, Zoo (Licht) leg. Chantaramongkol &amp; Malicky 1989”;
<specimenCount id="9D0317E7C81721554B19E8FD37E5FE16" box="[572,651,402,427]" pageId="22" pageNumber="473" type="male">1 male</specimenCount>
, “N-THAILAND,
<date id="FFBBFAAEC81721554A45E8FD36B7FE16" box="[864,985,402,427]" pageId="22" pageNumber="473" value="1918-01-06" valueMax="1918-08-06" valueMin="1918-01-06">
1. -8. 6,
<geoCoordinate id="EE31BAA9C81721554A9BE8FD36B7FE16" box="[958,985,403,427]" direction="north" orientation="latitude" pageId="22" pageNumber="473" precision="925" value="18.816668">18</geoCoordinate>
</date>
º49’N
<geoCoordinate id="EE31BAA9C81721554D01E8FC3111FE16" box="[1060,1151,402,427]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="925" value="98.95">98º57’E</geoCoordinate>
Chiang Mai, Zoo (Licht) leg. Chantaramongkol &amp; Malicky 1989”, genit. s. no. HY 844.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C817215449E2E8B431E2FD81" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473" type="description">
<paragraph id="8BBADC6EC817215549E2E8B43416FD86" blockId="22.[151,1437,258,571]" pageId="22" pageNumber="473">
<emphasis id="B971007CC817215549E2E8B43437FE4E" bold="true" box="[199,345,474,499]" pageId="22" pageNumber="473">Description.</emphasis>
Male. Body (
<figureCitation id="133EC0EBC81721554B27E8B43738FE4E" box="[514,598,474,499]" captionStart="FIGURE 13" captionStartId="17.[151,250,1610,1632]" captionTargetBox="[155,1420,202,1578]" captionTargetId="figure@17.[151,1430,193,1583]" captionTargetPageId="17" captionText="FIGURE 13. Habitus of Ypsiloncyphon spp., holotype, male (except for F). A) Y. guamensis sp. n., B) Y. philippinensis sp. n., C) Y. panayensis sp. n., D) Y. chlorizanoides sp. n., E) Y. bicolor sp. n., F) Y. bicolor sp. n., paratype, female, G) Y. sumatranus sp. n., H) Y. acer sp. n., I) Y. nokrekensis sp. n., J) Y. boukali sp. n., K) Y. javanicus sp. n., L) Y. nepalensis sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239912/files/figure.png" pageId="22" pageNumber="473">Fig. 13</figureCitation>
E) oval, well convex dorsally, shiny, closely covered with silver setae. Coloration of head, antennae, pronotum and legs yellowish orange; scutellar shield, elytra, meso- and metaventrites and abdomen black.
</paragraph>
<caption id="DF7A8CE6C817215549B2EC133714FA0F" httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473" targetBox="[330,1260,612,1375]" targetPageId="22">
<paragraph id="8BBADC6EC817215549B2EC133714FA0F" blockId="22.[151,1435,1405,1458]" pageId="22" pageNumber="473">
<emphasis id="B971007CC817215549B2EC133474FA2F" bold="true" box="[151,282,1405,1427]" pageId="22" pageNumber="473">FIGURE 18</emphasis>
.
<taxonomicName id="4C05A7EDC8172155480CEC133762FA2E" box="[297,524,1405,1427]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="B971007CC8172155480CEC133762FA2E" box="[297,524,1405,1427]" italics="true" pageId="22" pageNumber="473">Ypsiloncyphon bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C81721554B31EC133721FA2F" box="[532,591,1405,1426]" pageId="22" pageNumber="473" rank="species">
<emphasis id="B971007CC81721554B31EC133721FA2F" bold="true" box="[532,591,1405,1426]" pageId="22" pageNumber="473">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, male. A) abdominal sternites III–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view.
</paragraph>
</caption>
<paragraph id="8BBADC6EC817215549E2ECB03690F972" blockId="22.[151,1436,1502,2031]" pageId="22" pageNumber="473">Head relatively large, flat dorsally, finely punctate, front margin of clypeus shallowly concave. Eyes large, prominent; the distance between eyes about 1.8 times as long as the maximum diameter of an eye. Pronotum punctate as on head, depressed in lateral parts; anterior and lateral margins straight; posterior margin gently arcuate; antero- and postero-lateral corners obtuse; PW/PL 1.97–2.27 (2.10). Scutellar shield moderate in size, punctate as on head, equilaterally triangular. Elytra ovate, widest at the middle, densely punctate, strongly convex dorsally in caudal 2/5; humeral parts projecting dorso-anteriorly; apices right-angled; EL/EW 1.19–1.29 (1.24); EL/PL 3.89–4.33 (4.06); EW/PW 1.53–1.60 (1.56); TL/EW 1.47–1.62 (1.54).</paragraph>
<paragraph id="8BBADC6EC817215449E2EFB43580FF69" blockId="22.[151,1436,1502,2031]" lastBlockId="23.[151,1436,151,717]" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
Sternite III (
<figureCitation id="133EC0EBC8172155486AEFB434F0F94E" box="[335,414,1754,1779]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
A) glabrous in lateral parts. Caudal margin of sternite VII (
<figureCitation id="133EC0EBC81721554D0AEFB43110F94E" box="[1071,1150,1754,1779]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
A) gently arcuate. Tergite VII lacking apodeme, lightly projecting in apical margin; tergite VIII (
<figureCitation id="133EC0EBC81721554AE1EF903179F8AA" box="[964,1047,1790,1815]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
B) lightly sclerotized, trapezoidal, bearing long setae and minute spines along apical margin, with a pair of slender apodemes; tergite IX (
<figureCitation id="133EC0EBC81721554C0BEE4C30EEF886" box="[1326,1408,1826,1851]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
C) lightly sclerotized, trapezoidal, apical margin convex, apical part sparsely punctate, with a pair of long and slender apodemes; sternite VIII small; sternite IX (
<figureCitation id="133EC0EBC81721554BA3EE0437B9F83E" box="[646,727,1898,1923]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
D) moderately sclerotized, very long, U-shaped, covered with transverse keels in apical part, pointed at apices. Tegmen (
<figureCitation id="133EC0EBC81721554A15EEE036EFF81A" box="[816,897,1934,1959]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
E) moderately sclerotized, Y-shaped, as long as sternite IX; apical 1/3 (parameres) bifid, bearing minute spines in apical parts. Penis (
<figureCitation id="133EC0EBC81721554D41EEDC31DBF876" box="[1124,1205,1970,1995]" captionStart="FIGURE 18" captionStartId="22.[151,250,1405,1427]" captionTargetBox="[330,1260,612,1375]" captionTargetId="figure@22.[324,1263,609,1383]" captionTargetPageId="22" captionText="FIGURE 18. Ypsiloncyphon bicolor sp. n., holotype, male. A) abdominal sternites III – VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen, F) penis in ventral view." httpUri="https://zenodo.org/record/239917/files/figure.png" pageId="22" pageNumber="473">Fig. 18</figureCitation>
F) short, moderately sclerotized, about 0.75 times as long as tegmen; pala relatively long, expanded in basal part; parameroids long and wide, sparsely punctate in lateral parts, obtuse at apex; trigonium long and slender, as long as parameroids, rounded at apex.
</paragraph>
<paragraph id="8BBADC6EC816215449E2E9B1314AFE6D" blockId="23.[151,1436,151,717]" pageId="23" pageNumber="474">
Female (
<figureCitation id="133EC0EBC8162154480DE9B13417FF45" box="[296,377,223,248]" captionStart="FIGURE 13" captionStartId="17.[151,250,1610,1632]" captionTargetBox="[155,1420,202,1578]" captionTargetId="figure@17.[151,1430,193,1583]" captionTargetPageId="17" captionText="FIGURE 13. Habitus of Ypsiloncyphon spp., holotype, male (except for F). A) Y. guamensis sp. n., B) Y. philippinensis sp. n., C) Y. panayensis sp. n., D) Y. chlorizanoides sp. n., E) Y. bicolor sp. n., F) Y. bicolor sp. n., paratype, female, G) Y. sumatranus sp. n., H) Y. acer sp. n., I) Y. nokrekensis sp. n., J) Y. boukali sp. n., K) Y. javanicus sp. n., L) Y. nepalensis sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239912/files/figure.png" pageId="23" pageNumber="474">Fig. 13</figureCitation>
F). Sexual dimorphism distinct in the presence of U-shaped depressed excitator situated around basal 2/5 of elytral sutures; PW/PL 2.03–2.32 (2.18); EL/EW 1.18–1.36 (1.30); EL/PL 3.78–4.64 (4.24); EW/PW 1.39–1.62 (1.50); TL/EW 1.47–1.68 (1.60). Caudal margin of sternite VII (
<figureCitation id="133EC0EBC81621544ADCE8493125FEFD" box="[1017,1099,295,320]" captionStart="FIGURE 19" captionStartId="23.[151,250,1754,1776]" captionTargetBox="[297,1285,753,1726]" captionTargetId="figure@23.[292,1295,746,1733]" captionTargetPageId="23" captionText="FIGURE 19. Ypsiloncyphon bicolor sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239918/files/figure.png" pageId="23" pageNumber="474">Fig. 19</figureCitation>
A) arcuate. Tergite VIII (
<figureCitation id="133EC0EBC81621544C4BE84935DDFED9" captionStart="FIGURE 19" captionStartId="23.[151,250,1754,1776]" captionTargetBox="[297,1285,753,1726]" captionTargetId="figure@23.[292,1295,746,1733]" captionTargetPageId="23" captionText="FIGURE 19. Ypsiloncyphon bicolor sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239918/files/figure.png" pageId="23" pageNumber="474">Fig. 19</figureCitation>
B) moderately sclerotized, trapezoidal, bearing short spines on caudal margin, with a pair of long and slender apodemes protruding from antero-lateral corners. Sternite VIII (
<figureCitation id="133EC0EBC81621544A55E80136ACFE35" box="[880,962,367,392]" captionStart="FIGURE 19" captionStartId="23.[151,250,1754,1776]" captionTargetBox="[297,1285,753,1726]" captionTargetId="figure@23.[292,1295,746,1733]" captionTargetPageId="23" captionText="FIGURE 19. Ypsiloncyphon bicolor sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239918/files/figure.png" pageId="23" pageNumber="474">Fig. 19</figureCitation>
C) moderately sclerotized, oblong, with a long median apodeme. Ovipositor (
<figureCitation id="133EC0EBC81621544B0BE8FA37EFFE11" box="[558,641,404,429]" captionStart="FIGURE 19" captionStartId="23.[151,250,1754,1776]" captionTargetBox="[297,1285,753,1726]" captionTargetId="figure@23.[292,1295,746,1733]" captionTargetPageId="23" captionText="FIGURE 19. Ypsiloncyphon bicolor sp. n., paratype, female. A) sternites V – VII, B) tergite VIII, C) sternite VIII, D) ovipositor." httpUri="https://zenodo.org/record/239918/files/figure.png" pageId="23" pageNumber="474">Fig. 19</figureCitation>
D) long, with a branch in apical 8/9 of baculus; approximate ratio of stylus, coxite and baculus (n = 1) as 1.0: 5.0: 31.7; sclerotized structures absent.
</paragraph>
<paragraph id="8BBADC6EC816215449E2E8B231E2FD81" blockId="23.[151,1436,151,717]" pageId="23" pageNumber="474">
<emphasis id="B971007CC816215449E2E8B23413FE48" bold="true" box="[199,381,476,501]" pageId="23" pageNumber="474">Measurements.</emphasis>
Male (n = 4): TL 1.72–1.86 (1.77) mm; PW 0.70–0.75 (0.73) mm; PL 0.33–0.38 (0.35) mm; EW 1.10–1.20 (1.15) mm; EL 1.38–1.48 (1.42) mm. Female (n = 9): TL 1.58–1.88 (1.77) mm; PW 0.65–0.80 (0.74) mm; PL 0.28–0.38 (0.34) mm; EW 0.97–1.15 (1.10) mm; EL 1.25–1.53 (1.43) mm.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C816215449E2EB293751FDDD" box="[199,575,583,608]" pageId="23" pageNumber="474" type="distribution">
<paragraph id="8BBADC6EC816215449E2EB293751FDDD" blockId="23.[151,1436,151,717]" box="[199,575,583,608]" pageId="23" pageNumber="474">
<emphasis id="B971007CC816215449E2EB29340EFDDD" bold="true" box="[199,352,583,608]" pageId="23" pageNumber="474">Distribution.</emphasis>
<collectingCountry id="F3129CFEC81621544842EB2934A3FDDD" box="[359,461,583,608]" name="Thailand" pageId="23" pageNumber="474">Thailand</collectingCountry>
,
<collectingCountry id="F3129CFEC816215448FCEB293752FDDD" box="[473,572,583,608]" name="Vietnam" pageId="23" pageNumber="474">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C816215449E2EB023486FD15" pageId="23" pageNumber="474" type="discussion">
<paragraph id="8BBADC6EC816215449E2EB023486FD15" blockId="23.[151,1436,151,717]" pageId="23" pageNumber="474">
<emphasis id="B971007CC816215449E2EB023455FD38" bold="true" box="[199,315,620,645]" pageId="23" pageNumber="474">Remarks.</emphasis>
This is a remarkable species in having a distinctive penis in which the trigonium is long and slender and the parameroids are wide.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C816215449B2EBDA301CFD70" box="[151,1394,692,717]" pageId="23" pageNumber="474" type="etymology">
<paragraph id="8BBADC6EC816215449B2EBDA301CFD70" blockId="23.[151,1436,151,717]" box="[151,1394,692,717]" pageId="23" pageNumber="474">
<emphasis id="B971007CC816215449B2EBDA3470FD70" bold="true" box="[151,286,692,717]" pageId="23" pageNumber="474">Etymology.</emphasis>
The species name refers to the bicolored body, i.e., yellowish head and pronotum and black elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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