248 lines
46 KiB
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248 lines
46 KiB
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<document id="2AB9913C94F83B95EEB21F434FBDF29A" ID-CLB-Dataset="291999" ID-DOI="10.25674/so95iss1id311" ID-GBIF-Dataset="06b03cc4-e211-4f65-be07-742740b0053c" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFCDFFFABF14FE6C08100580" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus anuliflagellatus Frisch 2022, spec. nov." docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="36" updateTime="1711467745701" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="438FBDA132D33BE2B0EABC6A30532383">
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<mods:title id="173473CCCD697945D370991146694B01">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="32CCB7B8252FD5A5864A84BA5206A891" type="personal">
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<mods:roleTerm id="1B7F6338A5B1B5A64DDE053ED57B0A5F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2AFA3870E8681E954F5F068E078E8973">Frisch, Johannes</mods:namePart>
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</mods:name>
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<mods:name id="CE515ECAABC76DF30730EFB6F1F837E6" type="personal">
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</mods:role>
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<mods:namePart id="9214D4392B460A5E597343F85408DCF2">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="BF1E66D5D36A985FAB66E2E15FE371E1">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="F3E557AB6B399E052F7816C337FA6DFC">text</mods:typeOfResource>
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<mods:title id="31E3EB5AF213C3F8C8D84CD40CE14A22">Soil Organisms</mods:title>
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</mods:titleInfo>
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<mods:part id="7404A1642994243BCDBB5A283CC7A11E">
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<mods:date id="B8F324883A20FBE561CD58E1CEE16E36">2022</mods:date>
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<mods:detail id="339BAB0AB4CFDDAB65D4EDC5D2E65506" type="pubDate">
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<mods:number id="50536E41F396E05C14B5DE047B72379C">2022-04-01</mods:number>
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<mods:number id="19890BE00EAA6B59258BD6CFA2512791">95</mods:number>
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<mods:number id="D45F7022AD20B813F7BB9BA809249A8F">1</mods:number>
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<mods:start id="468CAAE431A236B4DEA951AFDC668CBF">23</mods:start>
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<mods:end id="CF0746B35AAC87EACFCDBC5E9D47B493">73</mods:end>
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<mods:url id="6AE2013B0CBDFEDAC2C3CDD597B033C8">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="8B7173309380652E498BFDC4D8FE0517">journal article</mods:classification>
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<mods:identifier id="4050795CFFCE1B321908757CC828573F" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFCDFFFABF14FE6C08100580" ID-DOI="http://doi.org/10.5281/zenodo.10724286" ID-Zenodo-Dep="10724286" LSID="urn:lsid:plazi:treatment:03B7C77AFFCDFFFABF14FE6C08100580" httpUri="http://treatment.plazi.org/id/03B7C77AFFCDFFFABF14FE6C08100580" lastPageId="14" lastPageNumber="37" pageId="13" pageNumber="36">
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<subSubSection id="C30425E7FFCDFFF9BF14FE6C087D05A2" box="[847,1376,404,431]" pageId="13" pageNumber="36" type="nomenclature">
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<paragraph id="8BA1766CFFCDFFF9BF14FE6C087D05A2" blockId="13.[847,1376,404,464]" box="[847,1376,404,431]" pageId="13" pageNumber="36">
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<heading id="D0E9C100FFCDFFF9BF14FE6C087D05A2" box="[847,1376,404,431]" fontSize="11" level="2" pageId="13" pageNumber="36" reason="5">
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<taxonomicName id="4C1E0DEFFFCDFFF9BF14FE6C09C105A3" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[847,1244,404,431]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="anuliflagellatus" status="sp. nov.">
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<emphasis id="B96AAA7EFFCDFFF9BF14FE6C09C105A3" bold="true" box="[847,1244,404,431]" pageId="13" pageNumber="36">
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<emphasis id="B96AAA7EFFCDFFF9BF14FE6C099805A2" bold="true" box="[847,1157,404,431]" italics="true" pageId="13" pageNumber="36">Scopaeus anuliflagellatus</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFCDFFF9B8B0FE6D087D05A2" box="[1259,1376,405,431]" pageId="13" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFCDFFF9BF14FE4D09EE05DD" box="[847,1267,437,464]" pageId="13" pageNumber="36" type="description">
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<paragraph id="8BA1766CFFCDFFF9BF14FE4D09EE05DD" blockId="13.[847,1376,404,464]" box="[847,1267,437,464]" pageId="13" pageNumber="36">
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(
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<figureCitation id="13256AE9FFCDFFF9BF0CFE4D0EAD05C2" box="[855,944,437,463]" captionStart="Figures 11–13" captionStartId="7.[189,262,1931,1953]" captionTargetBox="[232,1409,674,1880]" captionTargetId="figure-224@7.[189,1436,660,1912]" captionTargetPageId="7" captionText="Figures 11–13. Habitus of Scopaeus anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (11); S. grandis, paratype, Jawa Barat, Sukabumi: Cidahu (12); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (13)." figureDoi="http://doi.org/10.5281/zenodo.10724244" httpUri="https://zenodo.org/record/10724244/files/figure.png" pageId="13" pageNumber="36">Figs 11</figureCitation>
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,
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<figureCitation id="13256AE9FFCDFFF9BFE6FE4D0EC005C2" box="[957,989,437,463]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">36</figureCitation>
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,
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<figureCitation id="13256AE9FFCDFFF9BFB2FE4E092005C2" box="[1001,1085,437,464]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77 – 79</figureCitation>
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,
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<figureCitation id="13256AE9FFCDFFF9B812FE4D096505C2" box="[1097,1144,437,463]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="13" pageNumber="36">129</figureCitation>
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,
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<figureCitation id="13256AE9FFCDFFF9B8D8FE4D09F605DD" box="[1155,1259,437,464]" captionStart="Figures 155–162" captionStartId="41.[189,262,1839,1861]" captionTargetBox="[260,1402,386,1672]" captionTargetId="figure-168@41.[189,1436,316,1820]" captionTargetPageId="41" captionText="Figures 155–162. Sperm pump (155–162) and bursa (155, 159, 161) of ♀ of Scopaeus posoanus, paratype, Sulawesi Tengah: Lake Poso (155, with lateral gonocoxal plates IX; 156); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (157, with bursal duct; 158); S. anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (159, with lateral gonocoxal plates folded laterally; 160); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (161, 162). Long scale bar: 156, 158, 160, 162; short scale bar: 155, 157, 159, 161. Abbreviations: ap – apophysis of chamber segment, b – (sclerotized end of) bursa, bd – bursal duct, cs – chamber segment, dp – distal process of chamber segment, lt – lateral gonocoxal plate IX, sd – spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.10724311" httpUri="https://zenodo.org/record/10724311/files/figure.png" pageId="13" pageNumber="36">159, 160</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFCDFFF9BF14FE010EFB06DB" pageId="13" pageNumber="36" type="materials_examined">
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<paragraph id="8BA1766CFFCDFFF9BF14FE010EFB06DB" blockId="13.[824,1437,504,1975]" pageId="13" pageNumber="36">
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<emphasis id="B96AAA7EFFCDFFF9BF14FE010911061D" bold="true" box="[847,1036,504,529]" pageId="13" pageNumber="36">Type specimens:</emphasis>
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<materialsCitation id="3B767C31FFCDFFF9B84EFE000918067E" collectingDate="2015-09-29" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="1210" latitude="-6.7383337" location="Cirasamala River" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" pageId="13" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
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<collectingCountry id="F30936FCFFCDFFF9B84EFE000962061D" box="[1045,1151,504,528]" name="Indonesia" pageId="13" pageNumber="36">Indonesia</collectingCountry>
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,
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<collectingRegion id="49DAB88EFFCDFFF9B8D6FE010814061D" box="[1165,1289,505,528]" country="Indonesia" name="Jawa Barat" pageId="13" pageNumber="36">Jawa Barat</collectingRegion>
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:
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<typeStatus id="54A5C8CEFFCDFFF9B94EFE000865061D" box="[1301,1400,504,528]" pageId="13" pageNumber="36" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFCDFFF9B9D9FE010885061D" box="[1410,1432,505,528]" count="1" pageId="13" pageNumber="36" type="male">♂</specimenCount>
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,
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<collectingCounty id="62C00EE0FFCDFFF9BF63FDE10EBA063C" box="[824,935,537,561]" pageId="13" pageNumber="36">Sukabumi</collectingCounty>
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, NNW
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<collectingMunicipality id="6BC5EC16FFCDFFF9BFA0FDE10951063C" box="[1019,1100,537,561]" pageId="13" pageNumber="36">Cidahu</collectingMunicipality>
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:
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<location id="8EC120B7FFCDFFF9B803FDE1080E063C" LSID="urn:lsid:plazi:treatment:03B7C77AFFCDFFFABF14FE6C08100580:8EC120B7FFCDFFF9B803FDE1080E063C" box="[1112,1299,537,561]" country="Indonesia" county="Sukabumi" latitude="-6.7383337" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" name="Cirasamala River" pageId="13" pageNumber="36" stateProvince="Jawa Barat">Cirasamala River</location>
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(S-slope
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<location id="8EC120B7FFCDFFF9B926FDE20E6A065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFCDFFFABF14FE6C08100580:8EC120B7FFCDFFF9B926FDE20E6A065F" country="Indonesia" county="Sukabumi" latitude="-6.7383337" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" name="Mt Salak" pageId="13" pageNumber="36" stateProvince="Jawa Barat">Mt Salak</location>
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), (
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<geoCoordinate id="EE2A10ABFFCDFFF9BFC8FDC2090A065F" box="[915,1047,570,594]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="13" pageNumber="36" precision="15" seconds="18" value="-6.7383337">06°44’18’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFCDFFF9B878FDC209AA065F" box="[1059,1207,570,594]" degrees="106" direction="east" minutes="42" orientation="longitude" pageId="13" pageNumber="36" precision="15" seconds="52" value="106.71444">106°42’52’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFCDFFF9B890FDC20802065F" box="[1227,1311,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="13" pageNumber="36" unit="m" value="1210.0">
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<elevation id="0033915FFFCDFFF9B890FDC20802065F" box="[1227,1311,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="13" pageNumber="36" unit="m" value="1210.0">1210 m</elevation>
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</quantity>
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,
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<date id="FFA050ACFFCDFFF9B970FDC20885065F" box="[1323,1432,570,594]" pageId="13" pageNumber="36" value="2015-09-29">
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<collectingDate id="EFE4A944FFCDFFF9B970FDC20885065F" box="[1323,1432,570,594]" pageId="13" pageNumber="36" value="2015-09-29">29.9.2015</collectingDate>
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</date>
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, leg.
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<collectorName id="26EB13BAFFCDFFF9BF32FDA30EB1067E" box="[873,940,603,627]" pageId="13" pageNumber="36">Frisch</collectorName>
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(
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<collectionCode id="ED0FEEA9FFCDFFF9BFE5FDA30EE0067E" box="[958,1021,603,627]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="13" pageNumber="36" type="Museum">MZB</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B767C31FFCDFFF9B849FDA40EFF06DB" collectingDate="2015-05-10" collectionCode="MFNB" collectorName="Halimun-Salak NP & Frisch" country="Indonesia" county="Sukabumi" elevation="1020" latitude="-6.7461114" location="Cikaniki River" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" pageId="13" pageNumber="36" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
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<typeStatus id="54A5C8CEFFCDFFF9B849FDA409690679" box="[1042,1140,604,628]" pageId="13" pageNumber="36" type="paratype">Paratype</typeStatus>
|
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:
|
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|
|
<specimenCount id="9D18BDE5FFCDFFF9B8DBFDA309B1067E" box="[1152,1196,603,627]" count="1" pageId="13" pageNumber="36" type="female">1 ♀</specimenCount>
|
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,
|
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|
|
<collectingCounty id="62C00EE0FFCDFFF9B8E7FDA30836067E" box="[1212,1323,603,627]" pageId="13" pageNumber="36">Sukabumi</collectingCounty>
|
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,
|
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<collectingMunicipality id="6BC5EC16FFCDFFF9B962FDA30884067E" box="[1337,1433,603,627]" pageId="13" pageNumber="36">Cikaniki</collectingMunicipality>
|
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:
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|
<location id="8EC120B7FFCDFFF9BF63FD840EF80699" LSID="urn:lsid:plazi:treatment:03B7C77AFFCDFFFABF14FE6C08100580:8EC120B7FFCDFFF9BF63FD840EF80699" box="[824,997,636,660]" country="Indonesia" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Cikaniki River" pageId="13" pageNumber="36" stateProvince="Jawa Barat">Cikaniki River</location>
|
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(
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|
<location id="8EC120B7FFCDFFF9B859FD8409840699" LSID="urn:lsid:plazi:treatment:03B7C77AFFCDFFFABF14FE6C08100580:8EC120B7FFCDFFF9B859FD8409840699" box="[1026,1177,636,660]" country="Indonesia" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Mt Halimun" pageId="13" pageNumber="36" stateProvince="Jawa Barat">Mt Halimun</location>
|
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,
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<collectorName id="26EB13BAFFCDFFF9B8E8FD84088C0699" box="[1203,1425,636,660]" pageId="13" pageNumber="36">Halimun-Salak NP</collectorName>
|
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), (
|
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|
|
<geoCoordinate id="EE2A10ABFFCDFFF9BF1BFD650ED806B8" box="[832,965,669,693]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="13" pageNumber="36" precision="15" seconds="46" value="-6.7461114">06°44’46’’S</geoCoordinate>
|
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,
|
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|
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<geoCoordinate id="EE2A10ABFFCDFFF9BF83FD65097106B8" box="[984,1132,669,693]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="13" pageNumber="36" precision="15" seconds="25" value="106.540276">106°32’25’’E</geoCoordinate>
|
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),
|
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|
|
<quantity id="4CE6DB89FFCDFFF9B8DCFD6509FF06B9" box="[1159,1250,669,693]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="13" pageNumber="36" unit="m" value="1020.0">
|
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|
|
<elevation id="0033915FFFCDFFF9B8DCFD6509FF06B9" box="[1159,1250,669,693]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="13" pageNumber="36" unit="m" value="1020.0">1020 m</elevation>
|
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|
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</quantity>
|
|||
|
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,
|
|||
|
|
<date id="FFA050ACFFCDFFF9B8AEFD65087F06B8" box="[1269,1378,669,693]" pageId="13" pageNumber="36" value="2015-05-10">
|
|||
|
|
<collectingDate id="EFE4A944FFCDFFF9B8AEFD65087F06B8" box="[1269,1378,669,693]" pageId="13" pageNumber="36" value="2015-05-10">5.10.2015</collectingDate>
|
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</date>
|
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, leg.
|
|||
|
|
<collectorName id="26EB13BAFFCDFFF9BF63FD460E6606DB" box="[824,891,702,726]" pageId="13" pageNumber="36">Frisch</collectorName>
|
|||
|
|
(
|
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|
|
<collectionCode id="ED0FEEA9FFCDFFF9BFD2FD460EC706DB" box="[905,986,702,726]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="36">MFNB</collectionCode>
|
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|
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)
|
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|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
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|
|
<subSubSection id="C30425E7FFCDFFFABF14FD260FDA0527" lastPageId="14" lastPageNumber="37" pageId="13" pageNumber="36" type="description">
|
|||
|
|
<paragraph id="8BA1766CFFCDFFF9BF14FD260848006D" blockId="13.[824,1437,504,1975]" pageId="13" pageNumber="36">
|
|||
|
|
<emphasis id="B96AAA7EFFCDFFF9BF14FD260EC106FB" bold="true" box="[847,988,734,758]" pageId="13" pageNumber="36">Description:</emphasis>
|
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|
|
Habitus and coloring as in
|
|||
|
|
<figureCitation id="13256AE9FFCDFFF9B91BFD27088506FA" box="[1344,1432,735,759]" captionStart="Figures 11–13" captionStartId="7.[189,262,1931,1953]" captionTargetBox="[232,1409,674,1880]" captionTargetId="figure-224@7.[189,1436,660,1912]" captionTargetPageId="7" captionText="Figures 11–13. Habitus of Scopaeus anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (11); S. grandis, paratype, Jawa Barat, Sukabumi: Cidahu (12); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (13)." figureDoi="http://doi.org/10.5281/zenodo.10724244" httpUri="https://zenodo.org/record/10724244/files/figure.png" pageId="13" pageNumber="36">Fig. 11</figureCitation>
|
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|
|
. Head subquadrate. Antennae with penultimate segment slightly elongate or quadrate. Mesotibia moderately thickened. Forebody surface with dense, rugulose, setose punctation, matt; abdomen somewhat shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body black except for light tip of abdomen, medium brown maxillary palpi and yellow-brown legs with femora dark brown except for distal end; antennae with scapus medium brown, from blackish pedicellus evenly lighter towards terminal segments. Total body length 3.6 – 4.0 mm; forebody length
|
|||
|
|
<quantity id="4CE6DB89FFCDFFF9B89DFBB0084C006D" box="[1222,1361,1096,1120]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="13" pageNumber="36" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1 – 2.2 mm</quantity>
|
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|
|
.
|
|||
|
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</paragraph>
|
|||
|
|
<paragraph id="8BA1766CFFCDFFF9BF14FB91084C0358" blockId="13.[824,1437,504,1975]" pageId="13" pageNumber="36">
|
|||
|
|
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII widely emarginate in almost posterior sixth with median third of emargination somewhat convex (
|
|||
|
|
<figureCitation id="13256AE9FFCDFFF9B8E4FB34080200E9" box="[1215,1311,1228,1252]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="13" pageNumber="36">Fig. 129</figureCitation>
|
|||
|
|
). Aedeagus (
|
|||
|
|
<figureCitation id="13256AE9FFCDFFF9BF1BFB150E8A0108" box="[832,919,1261,1285]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 36</figureCitation>
|
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|
|
,
|
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|
|
<figureCitation id="13256AE9FFCDFFF9BFFDFB150EEF0108" box="[934,1010,1261,1285]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77 – 79</figureCitation>
|
|||
|
|
) about
|
|||
|
|
<quantity id="4CE6DB89FFCDFFF9B810FB1509830109" box="[1099,1182,1261,1285]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="36" unit="mm" value="1.2">1.2 mm</quantity>
|
|||
|
|
long with distal lobes strongly asymmetrical; dextral apical lobe remarkably enlarged apically with broad, convex apex and acute, proximoventral end (
|
|||
|
|
<figureCitation id="13256AE9FFCDFFF9B87BFAB7096B016A" box="[1056,1142,1359,1383]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 36</figureCitation>
|
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,
|
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|
|
<figureCitation id="13256AE9FFCDFFF9B8DFFAA809BF0165" box="[1156,1186,1360,1384]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77</figureCitation>
|
|||
|
|
), in ventral and dorsal view slender, parallel (
|
|||
|
|
<figureCitation id="13256AE9FFCDFFF9B812FA8809C40185" box="[1097,1241,1392,1416]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Figs 78, 79</figureCitation>
|
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|
); sinistral apical lobe, compared to dextral apical lobe, considerably narrower with convex, proximoventral end in lateral view, but much wider in ventral and dorsal view (
|
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|
|
<figureCitation id="13256AE9FFCDFFF9B937FA2B0E9D0201" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Figs 78, 79</figureCitation>
|
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|
|
); apex of dorsal lobe curved ventrad, extended in dextral, hooked end and sinistral, lobiform end (
|
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|
|
<figureCitation id="13256AE9FFCDFFF9B964F9ED08920220" box="[1343,1423,1557,1581]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Fig. 77</figureCitation>
|
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); flagellum comparatively short, not projecting ventrally from wide, distal portion of apical lobes, with annular winding to left of aedeagus (
|
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|
|
<figureCitation id="13256AE9FFCDFFF9B8DFF98F09C00282" box="[1156,1245,1655,1679]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 36</figureCitation>
|
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|
,
|
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|
|
<figureCitation id="13256AE9FFCDFFF9B8B6F98008240282" box="[1261,1337,1655,1679]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77, 78</figureCitation>
|
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|
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); ventral lobe divided in scythe-shaped, distally curved, dextral lobe (
|
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|
|
<figureCitation id="13256AE9FFCDFFF9BF20F9410EC902DC" box="[891,980,1721,1745]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 36</figureCitation>
|
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|
|
,
|
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|
|
<figureCitation id="13256AE9FFCDFFF9BFBEF941091C02DC" box="[997,1025,1721,1745]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77</figureCitation>
|
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|
) and short, sinistral tooth (
|
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<figureCitation id="13256AE9FFCDFFF9B919F9410E4A02FF" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Fig. 77, 78</figureCitation>
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); lateral lobes reduced, marked by longer dextral and shorter sinistral group of setae (
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<figureCitation id="13256AE9FFCDFFF9B8F1F9030829031E" box="[1194,1332,1787,1811]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Figs 78, 79</figureCitation>
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); median foramen small, subtriangular, with evenly, convexly distad curved, transversal ridge (
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<figureCitation id="13256AE9FFCDFFF9B8C7F8C509EC0358" box="[1180,1265,1853,1877]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 36</figureCitation>
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,
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<figureCitation id="13256AE9FFCDFFF9B8A6F8C508580358" box="[1277,1349,1853,1877]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">77, 78</figureCitation>
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).
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</paragraph>
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<paragraph id="8BA1766CFFCDFFF9BF14F8A50EFE03BA" blockId="13.[824,1437,504,1975]" pageId="13" pageNumber="36">
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Female: Bursal duct membranous, about three times as long as sperm pump; bursa with specific sclerotization (
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<figureCitation id="13256AE9FFCDFFF9BF64F8670ECA03BA" box="[831,983,1951,1975]" captionStart="Figures 155–162" captionStartId="41.[189,262,1839,1861]" captionTargetBox="[260,1402,386,1672]" captionTargetId="figure-168@41.[189,1436,316,1820]" captionTargetPageId="41" captionText="Figures 155–162. Sperm pump (155–162) and bursa (155, 159, 161) of ♀ of Scopaeus posoanus, paratype, Sulawesi Tengah: Lake Poso (155, with lateral gonocoxal plates IX; 156); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (157, with bursal duct; 158); S. anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (159, with lateral gonocoxal plates folded laterally; 160); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (161, 162). Long scale bar: 156, 158, 160, 162; short scale bar: 155, 157, 159, 161. Abbreviations: ap – apophysis of chamber segment, b – (sclerotized end of) bursa, bd – bursal duct, cs – chamber segment, dp – distal process of chamber segment, lt – lateral gonocoxal plate IX, sd – spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.10724311" httpUri="https://zenodo.org/record/10724311/files/figure.png" pageId="13" pageNumber="36">Figs 159, 160</figureCitation>
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).
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</paragraph>
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<paragraph id="8BA1766CFFCEFFFABCF5FF0A0FDA0527" blockId="14.[151,764,241,397]" pageId="14" pageNumber="37">
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<emphasis id="B96AAA7EFFCEFFFABCF5FF0A0C300507" bold="true" box="[174,301,242,266]" pageId="14" pageNumber="37">Phylogeny:</emphasis>
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<taxonomicName id="4C1E0DEFFFCEFFFABD6EFF0A0F500504" authorityName="Frisch" authorityYear="2022" box="[309,589,241,265]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="anuliflagellatus">
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<emphasis id="B96AAA7EFFCEFFFABD6EFF0A0F500504" box="[309,589,241,265]" italics="true" pageId="14" pageNumber="37">Scopaeus anuliflagellatus</emphasis>
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</taxonomicName>
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is a member of the
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<taxonomicName id="4C1E0DEFFFCEFFFABCE4FEEB0C300527" authorityName="Oke" authorityYear="1933" box="[191,301,274,298]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="B96AAA7EFFCEFFFABCE4FEEB0C300527" box="[191,301,274,298]" italics="true" pageId="14" pageNumber="37">S. gracilis</emphasis>
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</taxonomicName>
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species group (
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<bibRefCitation id="EF8F0B9DFFCEFFFABD8EFEEA0FA60527" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[469,699,274,298]" pageId="14" pageNumber="37" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFCEFFFABCF5FECB0DC60580" pageId="14" pageNumber="37" type="distribution">
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<paragraph id="8BA1766CFFCEFFFABCF5FECB0DC60580" blockId="14.[151,764,241,397]" pageId="14" pageNumber="37">
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<emphasis id="B96AAA7EFFCEFFFABCF5FECB0C5A0546" bold="true" box="[174,327,307,331]" pageId="14" pageNumber="37">Distribution:</emphasis>
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<taxonomicName id="4C1E0DEFFFCEFFFABD15FECC0F740546" authorityName="Frisch" authorityYear="2022" box="[334,617,307,331]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="anuliflagellatus">
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<emphasis id="B96AAA7EFFCEFFFABD15FECC0F740546" box="[334,617,307,331]" italics="true" pageId="14" pageNumber="37">Scopaeus anuliflagellatus</emphasis>
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</taxonomicName>
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was found in two localities at Mount Salak and Mount Halimun,
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<collectingRegion id="49DAB88EFFCEFFFABE9DFEAD0DCB0580" country="Indonesia" name="Jawa Barat" pageId="14" pageNumber="37">Jawa Barat</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFCEFFFABF70FF0A08100580" pageId="14" pageNumber="37" type="etymology">
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<paragraph id="8BA1766CFFCEFFFABF70FF0A08100580" blockId="14.[788,1399,241,397]" pageId="14" pageNumber="37">
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<emphasis id="B96AAA7EFFCEFFFABF70FF0A0EAC0507" bold="true" box="[811,945,242,266]" pageId="14" pageNumber="37">Etymology:</emphasis>
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The epithet
|
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<taxonomicName id="4C1E0DEFFFCEFFFAB80AFF0909E40504" authorityName="Frisch" authorityYear="2022" box="[1105,1273,241,265]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="37" phylum="Arthropoda" rank="species" species="anuliflagellatus">
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<emphasis id="B96AAA7EFFCEFFFAB80AFF0909E40504" box="[1105,1273,241,265]" italics="true" pageId="14" pageNumber="37">anuliflagellatus</emphasis>
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</taxonomicName>
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[adjective, Latin, composed of the noun
|
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<emphasis id="B96AAA7EFFCEFFFAB820FEEA09DE0527" box="[1147,1219,274,298]" italics="true" pageId="14" pageNumber="37">anulus</emphasis>
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(ring) and the adjective
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<emphasis id="B96AAA7EFFCEFFFABFDBFECB0EED0546" box="[896,1008,307,331]" italics="true" pageId="14" pageNumber="37">flagellatus</emphasis>
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|
(indicating a characteristic/feature of the flagellum)] refers to the annular flagellum of the aedeagus (
|
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<figureCitation id="13256AE9FFCEFFFABFDDFE8D0EC40580" box="[902,985,373,397]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="14" pageNumber="37">Figs 36</figureCitation>
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,
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<figureCitation id="13256AE9FFCEFFFABFB8FE8E09300580" box="[995,1069,373,397]" captionStart="Figures 74–79" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 74–79. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (74–76); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (77–79). Abbreviations: al – apical lobes, al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, dl(d) – dextral end of dorsal lobe, dl(s) – sinistral end of dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="14" pageNumber="37">77 – 79</figureCitation>
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) of this new species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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